Catasetum and Cycnoches — Part 1 - American Orchid Society
Catasetum and Cycnoches — Part 1 - American Orchid Society
Catasetum and Cycnoches — Part 1 - American Orchid Society
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BEGINNER'S SERIES <strong>—</strong> 25<br />
<strong>Catasetum</strong> <strong>and</strong> <strong>Cycnoches</strong> <strong>—</strong> <strong>Part</strong> 1<br />
<strong>—</strong> <strong>Catasetum</strong>s with "Perfect" Flowers<br />
STEPHEN R. BATCHELOR<br />
THUS FAR in this series, the emphasis has been on genera which grow almost<br />
continuously, though slowly. For these genera, beginners are able to adopt<br />
cultural practices which require little variation. And yet, some genera do not<br />
grow constantly, but fluctuate between a state of rapid growth <strong>and</strong> one of nearly<br />
complete inactivity <strong>—</strong> all in the course of a year. Treatment of these genera, in<br />
contrast, must vary considerably for the best growth <strong>and</strong> flowering.<br />
Photo: Greg Allikas<br />
FIGURE 1 <strong>—</strong> Reaching a leafy maturity, the newest<br />
pseudobulb of this plant of <strong>Catasetum</strong> expansum initiates<br />
four inflorescences in mid-summer.<br />
Species <strong>and</strong> hybrids of <strong>Catasetum</strong> <strong>and</strong> <strong>Cycnoches</strong>, two closely related genera,<br />
are excellent examples of orchids which have a changeable growth rate. Their<br />
growth generally occurs in a great flush during the spring <strong>and</strong> summer, only to<br />
come to a st<strong>and</strong>still during the fall <strong>and</strong> winter. This may be disconcerting at first<br />
to the beginner. But with some experience, it soon becomes apparent that the<br />
plants follow a cycle of growth, clearly marked by changes in their behavior <strong>and</strong><br />
appearance. Observing these changes, <strong>and</strong> altering cultural practices<br />
accordingly, the beginner should have much success with catasetums <strong>and</strong><br />
cycnochcs, for they are among the most vigorous <strong>and</strong> floriferous of orchids.<br />
A large part of this series on catasetums <strong>and</strong> cycnoches will be devoted to
heir cycle of growth, <strong>and</strong> to how several talented growers meet the fluctuating<br />
needs of these genera. Prior to that, however, representative species <strong>and</strong><br />
hybrids need to be introduced. <strong>Catasetum</strong>s in particular offer an astounding<br />
variety of flowers, from the beautiful to the bizarre. Not every one will appeal!<br />
Just the same, those which lack conventional good-looks often compensate<br />
with a few surprises.<br />
PLANT AND FLOWERING HABIT OF CATASETUMS<br />
The genus <strong>Catasetum</strong> contains over one hundred epiphytic species native to<br />
the lower elevations of tropical America. Because of their warm origins,<br />
catasetums require warm conditions for proper growth <strong>and</strong> flowering. Though<br />
flower characteristics <strong>and</strong> presentation vary widely in this genus, plant habit is<br />
similar throughout. The plant of <strong>Catasetum</strong> expansum pictured in FIGURE 1,<br />
photographed in mid-summer, is representative of the genus. The growth begun<br />
in the springtime is already reaching maturity, producing a fleshy, cylindrical<br />
pseudobulb 6 inches (15 cm) high. This pseudobulb is fully ensheathed in a fanlike<br />
arrangement of eight thin, plicate (pleated) leaves, the largest of which<br />
measures about 12 inches (30 cm) long <strong>and</strong> 3 inches (8 cm) wide. The entire<br />
plant, excluding pot, is over 1'/2 feet (45 cm) high.<br />
Following closely behind the leafy pseudobulb in FIGURE 1 are those pseudobulbs<br />
produced in previous years. Their leafless state is an indication of the<br />
deciduous nature of catasetums. Leaves produced during one growing season<br />
are usually lost by the beginning of the next. In other words, by the time an eye<br />
swells at the base of this most recently matured pseudobulb, signaling a<br />
resumption of active growth, its leaves will have yellowed <strong>and</strong> fallen, or will be in<br />
the process of doing so. The actual timing of this leaf-fall, <strong>and</strong>, consequently, the<br />
duration of the plant's leafless state, largely depends on the growing conditions<br />
<strong>and</strong> the health of the plant. This will be discussed at length in upcoming articles.<br />
Though a <strong>Catasetum</strong> pseudobulb will lose its leaves within a year, the bases of<br />
these leaves remain, forming a dry, papery sheathing. Older pseudobulbs may<br />
lose this covering, as well as their plumpness, becoming exposed <strong>and</strong> furrowed<br />
with age. The thin, upward-growing secondary roots emanating from the potting<br />
medium in FIGURE 1 are a curious feature of many <strong>Catasetum</strong> species.<br />
The plant of <strong>Catasetum</strong> expansum in FIGURE 1 is also characteristic of the genus<br />
in that it is producing four inflorescences from the bases of its newest<br />
pseudobulbs. Though all basal in origin, the inflorescences of catasetums can<br />
differ in the position they ultimately take. Many, such as those of <strong>Catasetum</strong><br />
expansum pictured in FIGURE 1, are upright, becoming more arching as the<br />
flowers develop <strong>and</strong> add weight to the inflorescence. A few are strictly<br />
pendulous. The initiation of inflorescences from new growth soon after it<br />
matures results in most catasetums being summer- <strong>and</strong> fall-flowering. Yet there<br />
are notable exceptions to this generalization, which should come to light upon<br />
closer examination of the species <strong>and</strong> hybrids.<br />
CATASETUMS WITH PERFECT FLOWERS<br />
By the title of this first article, I do not mean to suggest that the catasetums<br />
about to be discussed arc flawless <strong>and</strong> therefore free from criticism! Rather, this<br />
title refers to the fact that the flowers of these catasetums have functioning<br />
male <strong>and</strong> female parts; they are, in botanical jargon, perfect, or bisexual,<br />
flowers. This is not a distinguishing feature within the orchid family, yet it is<br />
within the genus <strong>Catasetum</strong>. The great majority of <strong>Catasetum</strong> species have<br />
unisexual flowers, male flowers <strong>and</strong> female flowers, a highly unique trait within<br />
the orchid family. The ten or so <strong>Catasetum</strong> species with perfect flowers are<br />
different in this, as well as other floral features. So distinct are they that Dr.<br />
Calaway H. Dodson, a prominent taxonomist, has proposed separating them into
two genera of their own: Clowesia, to include <strong>Catasetum</strong> roseum, Ctsm.<br />
warscewiczii, Ctsm. thylaciochilum, Ctsm. russellianum <strong>and</strong> Ctstn. glaucoglossa; <strong>and</strong><br />
Dressleria, to contain <strong>Catasetum</strong> dilectum, Ctsm. eburneum, Ctsm. suavis <strong>and</strong> one new<br />
species (Dodson, 1975).<br />
Beginners will find, however, that for the most part these species with perfect<br />
flowers are still occasionally called catasetums by orchid growers. Hybrids made<br />
with these species are registered as <strong>Catasetum</strong> hybrids. Nevertheless, from a<br />
horticultural point of view as well, these species need to be treated separately, as<br />
they offer growers quite a different look from that of most <strong>Catasetum</strong> species.<br />
For these reasons, I will maintain here the two separate groups set forth by<br />
Dodson, while continuing to call these species <strong>and</strong> their hybrids catasetums.<br />
CLOWESIA<br />
The five species classified by Dodson into the genus Clowesia had been set apart<br />
previously, though not to that degree, as belonging to the Clowesia section of the<br />
Photo: Greg Allikas<br />
FIGURE 2 <strong>—</strong> <strong>Catasetum</strong> russellianum (syn. Clowesia russelliana)<br />
genus <strong>Catasetum</strong> (Hawkes, 1965). Species of the section, or genus Clowesia, differ<br />
from the majority of catasetums in a number of ways. In plant habit, they are<br />
considerably smaller. Their pseudobulbs are rarely over 4 inches (10 cm) high,<br />
often a good deal less, <strong>and</strong> are more conical than cylindrical in shape. Their<br />
inflorescences are not upright, or arching, but strongly pendulous, <strong>and</strong> tend to<br />
be shorter than most <strong>Catasetum</strong> species. The flowers of this group, though<br />
generally smaller, are perhaps the loveliest <strong>and</strong> most fragrant of all catasetums.<br />
They are, regrettably, quite typical of the genus in being short-lived, lasting little<br />
more than a week in most cases.<br />
Of the group, <strong>Catasetum</strong> russellianum has the most in common with the genus<br />
at large. Native to southern Mexico <strong>and</strong> Central America, this species has<br />
fragrant, light green flowers striped in a darker green (FIGURE 2). Though the<br />
sepals are narrower than the petals, both segments are long <strong>and</strong> well-spaced,<br />
creating a large <strong>and</strong> fairly open flower. Measuring 2'/2-3 inches (6.5-8 cm)<br />
across, these flowers are comparable in size to some of the largest-flowered<br />
catasetums. Their thin, almost transparent substance, however, sets them apart<br />
from the waxy flowers characteristic of most <strong>Catasetum</strong> species. The pouchshaped<br />
(saccate) lip in the lowermost position of the flower is the most
distinguishing feature of Calasetum russellianum. <strong>Catasetum</strong> thylaciochilum is very<br />
similar to Calasetum russellianum in flower form <strong>and</strong> coloration, but its lip is not<br />
nearly so pouched (Teuscher, 1977).<br />
Though the inflorescences of <strong>Catasetum</strong> russellianum, being pendent, are<br />
typical of Clowesia, they are exceptional in length <strong>and</strong> floriferousness, again<br />
matching the best of the genus. Less robust plants of <strong>Catasetum</strong> russellianum will<br />
naturally produce fewer flowers, but 15 to 20 flowers on inflorescences<br />
approximately 15 inches (38 cm) long can be expected of mature, healthy plants<br />
(Hawkes, 1965). The flowering season of <strong>Catasetum</strong> russellianum is like that of<br />
most catasetums, running from late summer into fall.<br />
FIGURE 3 <strong>—</strong> <strong>Catasetum</strong> warscewiczii 'Severini' HCC/AOS<br />
<strong>Catasetum</strong> warscewiczii, native to portions of Central <strong>and</strong> South America, is<br />
another species of this group which has fragrant, greenish flowers with darker<br />
green striping (FIGURE 3), but its similarity with <strong>Catasetum</strong> russellianum ends<br />
there. The sepals <strong>and</strong> petals of <strong>Catasetum</strong> warscewiczii are equally broad <strong>and</strong><br />
cupped, forming a full, round but small flower 1-1 Vi inches (2.5-4 cm) across.<br />
The marvelous, trilobed labellum has two distinctly striped side-lobes which<br />
arc upright, cradling the nose-like column between them. Its wonderfully<br />
fringed mid-lobe juts out like a bearded chin. The cavity formed at the<br />
convergence of the three lobes of the lip, with its darker coloration, adds depth<br />
to the flower.<br />
<strong>Catasetum</strong> warscewiczii produces pendent inflorescences, like Calasetum<br />
russellianum, but they are generally much shorter, less than 6 inches (15 cm) in<br />
length. Each inflorescence also tends to carry fewer flowers, no more than 10 or<br />
12, even for those specimen plants which have received cultural awards. Even so,<br />
the dense <strong>and</strong> uniform arrangement of flowers on the inflorescence results in a<br />
concentration of flowers near the plant. This pleasing, cascade effect is enhanced<br />
when a pseudobulb produces more than one inflorescence, as is often the case with<br />
<strong>Catasetum</strong> warscewiczii (FIGURE 3). Unlike <strong>Catasetum</strong> russellianum <strong>and</strong> most<br />
catasetums, <strong>Catasetum</strong> warscewic-zii tends to bloom during the winter. By this<br />
time, not only have the newest pseudobulbs matured, they have lost their leaves<br />
as well. In this way, too, the flowers are shown to their full advantage, without<br />
obstruction from view by the large leaves typical of this group <strong>and</strong> the entire<br />
genus.<br />
<strong>Catasetum</strong> roseum has much in common with <strong>Catasetum</strong> warscewiczii. This Mexican
species also flowers during the winter months from recently matured pseudobulbs<br />
Grower: Lincoln Adderley Photo: Charles M. Fitch<br />
FIGURE 4<strong>—</strong> <strong>Catasetum</strong> roseum ‘Chico’ CCM/AOS<br />
which have since lost their leaves. Robust pseudobulbs of <strong>Catasetum</strong> roseum, like<br />
Ctsm. warscewiczii, can produce multiple inflorescences, each measuring up to 6<br />
inches (15 cm) long <strong>and</strong> carrying a dozen flowers. The flowers of <strong>Catasetum</strong><br />
roseum are also of similar size, usually 1 inch (2.5 cm) across. But unlike Ctsm.<br />
warscewiczii, <strong>Catasetum</strong> roseum, as its name suggests, has flowers which are an<br />
attractive pink overall, with the exception of a green column (FIGURE 4). These<br />
flowers, in addition, have sepals <strong>and</strong> petals which are more pointed <strong>and</strong> less<br />
cupped. The lip of <strong>Catasetum</strong> roseum also has a highly fringed mid-lobe, but its<br />
side-lobes are not as well developed as Ctsm. warscewiczii.<br />
The specimen plant of <strong>Catasetum</strong> roseum in FIGURE 4 well illustrates the<br />
desirable traits of this species. In its leafless state at least, <strong>Catasetum</strong> roseum<br />
could certainly be classified as a miniature. The short, densely clustered<br />
pseudobulbs of this plant are contained in a 6-inch pot. At the same time, the<br />
plant is flowering profusely on equally compact, 5-inch (13-cm) inflorescences,<br />
nearly surrounding the plant <strong>and</strong> its pot in a wreath of 140 flowers. A small<br />
number of hybrids have been made using species from the Clowesia section of<br />
<strong>Catasetum</strong>. <strong>Catasetum</strong> Rebecca Northen (Grace Dunn [warscewiczii X roseum] X<br />
roseum), registered by W.W.G. Moir in 1971, is an attractive hybrid involving<br />
two of the group's most desirable species. The awarded clone of <strong>Catasetum</strong><br />
Rebecca Northen illustrated in FIGURE 5 combines the more rounded, cupped<br />
shape of <strong>Catasetum</strong> warscewiczii with the brighter color of <strong>Catasetum</strong> roseum,<br />
while maintaining floriferousness <strong>and</strong> a compact flowering <strong>and</strong> plant habit.<br />
When awarded in April, 1978, this clone displayed 47 flowers on three<br />
inflorescences, all produced by the same plump, leafless pseudobulb. The<br />
flowers averaged not quite one inch (2.2 cm) across.<br />
DRESSLERIA<br />
The four Central <strong>and</strong> South <strong>American</strong> <strong>Catasetum</strong> species with perfect flowers<br />
which Dodson classified in a new genus called Dressleria provide a vivid contrast<br />
to the ones just discussed. In plant habit, they tend to be much larger, looking
FIGURE 5--<br />
<strong>Catasetum</strong> Rebecca Northen ‘Marta’ HCC/AOS (Grace Dunn<br />
[warceswiczii x roseum] x roseum)<br />
more like typical catasetums. The inflorescences of these species are likewise<br />
considerably longer, <strong>and</strong> are upright, or nearly upright, not pendent. Plants of<br />
<strong>Catasetum</strong> dilectum (Dressleria dilecta), for example, often have pseudobulbs 8<br />
inches (20 cm) long, producing inflorescences up to 14 inches (35 cm) in length<br />
(Bechtel etal., 1981). It is the flowers of Dressleria, however, which are most<br />
distinct, bearing little resemblance to those of Clowesia.<br />
The flowers of the four species classified in Dressleria by Dodson are very similar, a<br />
fact which has caused some confusion in their identification. The flowers<br />
illustrated in FIGURE 6, for instance, while identified as <strong>Catasetum</strong> eburneum, bear<br />
closer resemblance to those of <strong>Catasetum</strong> dilectum (Dressleria dilecta), as<br />
described by Dodson (1975) <strong>and</strong> pictured in Bechtel et al. (1981). In any event,<br />
these flowers, photographed in summer, are representative of the group. Borne<br />
approximately a dozen in number on an upright inflorescence bending under<br />
their weight, these fleshy flowers have conspicuous, helmet-shaped lips. Their<br />
uppermost position indicates that the flowers are non-resupinate, as opposed to<br />
the resupinate flowers of Clowesia, with their lips lowermost. The lips of<br />
Dressleria flowers also lack the highly fringed nature of most Clowesia species,<br />
<strong>and</strong> are attached (adnate) to the column, not separate. The columns of the<br />
flowers pictured in FIGURE 6 are short <strong>and</strong> wide, <strong>and</strong> like the lip, have an opening<br />
or cavity. Strongly reflexed away from the lip, the greenish sepals <strong>and</strong> petals lack<br />
the prominence these segments have in Clowesia. Sepals <strong>and</strong> petals of other<br />
members of Dressleria, however, are not as reflexed as these. The widest portion<br />
of the flowers pictured measures less than one inch (2.1 cm) across. species also<br />
flowers during the winter months from recently matured pseudobulbs.
Grower: Mr. & Mrs. Grenville Seibels, II Photography: Edwin S. Boyett, Jr.<br />
FIGURE 6-- <strong>Catasetum</strong> eburneum ‘Memoria Vinton Overholt’, CBM/AOS<br />
Many of the remaining 90 or so <strong>Catasetum</strong> species have non-resupinate<br />
flowers with helmet-shaped lips like thosejust described. Other <strong>Catasetum</strong><br />
species have both resupinate <strong>and</strong> non-resupinate flowers, depending on whether<br />
those flowers are male or female! The unisexual catasetums <strong>—</strong> considered<br />
catasetums by everyone <strong>—</strong> will be the subject of the next article for this series<br />
REFERENCES<br />
Bechtel, Helmut, Cribb, Phillip <strong>and</strong> Edmund Launert. 1981. The Manual of Cultivated<br />
<strong>Orchid</strong> Species. The MIT Press, Cambridge, Massachusetts. Dodson, C.H. 1975.<br />
Dressleria <strong>and</strong> Clowesia: A New Genus <strong>and</strong> an Old One Revived in the Catasetinae<br />
(<strong>Orchid</strong>aceae). Selbyana 1(2): 130.<br />
Hawkes, Alex D. 1965. Encyclopaedia of Cultivated <strong>Orchid</strong>s. Faber <strong>and</strong> Faber,<br />
Limited, London. Teuscher, Henry. 1977.<br />
<strong>Catasetum</strong> thylaciochilum <strong>—</strong> Collector's Item. Amer. <strong>Orchid</strong>Soc. Bull. 46(11):