Annotated checklist of the beetles (Coleoptera) of the California Channel Islands
- Published
- Accepted
- Received
- Academic Editor
- Jason Bond
- Subject Areas
- Biodiversity, Biogeography, Entomology, Taxonomy, Zoology
- Keywords
- Island biogeography, Biodiversity informatics, Taxonomy, Nomenclature, Conservation, Endemism, Faunistics
- Copyright
- © 2023 Gimmel et al.
- Licence
- This is an open access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, reproduction and adaptation in any medium and for any purpose provided that it is properly attributed. For attribution, the original author(s), title, publication source (PeerJ) and either DOI or URL of the article must be cited.
- Cite this article
- 2023. Annotated checklist of the beetles (Coleoptera) of the California Channel Islands. PeerJ 11:e14793 https://doi.org/10.7717/peerj.14793
Abstract
The beetle fauna of the California Channel Islands is here enumerated for the first time in over 120 years. We provide an annotated checklist documenting species-by-island diversity from an exhaustive literature review and analysis of a compiled dataset of 26,609 digitized specimen records to which were added over 3,000 individual specimen determinations. We report 825 unique species from 514 genera and 71 families (including 17 new family records) comprising 1,829 species-by-island records. Species totals for each island are as follows: Anacapa (74); San Clemente (197); San Miguel (138); San Nicolas (146); Santa Barbara (64); Santa Catalina (370); Santa Cruz (503); and Santa Rosa (337). This represents the largest list of species published to date for any taxonomic group of animals on the Channel Islands; despite this, we consider the checklist to be preliminary. We present evidence that both inventory and taxonomic efforts on Channel Islands beetles are far from complete. Rarefaction estimates indicate there are at least several hundred more species of beetles yet to be recorded from the islands. Despite the incomplete nature of existing records, we found that species diversity is highly correlated with island area. We report 56 species which are putatively geographically restricted (endemic) to the Channel Islands, with two additional species of questionable endemic status. We also report 52 species from the islands which do not natively occur in the southern California region.
Introduction
The California Channel Islands are an archipelago of eight main islands between 20 and 98 km off the coast of southern California, USA. Often referred to as “North America’s Galápagos”, the biological diversity of the California Channel Islands has long captured the attention of natural historians of western North America. Detailed information about the islands’ geography, geologic history, natural history, and history of human activity may be found in many other publications, especially Schoenherr, Feldmeth & Emerson (1999) and Moody (2000); Miller (1985a) provided an introduction to the history of entomology of the islands.
Attempts to document species diversity of the islands have been scattered and unequal across taxonomic groups, with most organismal groups not having a reliable checklist or taxonomic treatment completed and made available. The largest published list is that of Ratay, Vanderplank & Wilder (2014) which cited 976 vascular plant taxa from the California Channel Islands, including species (922), subspecies, varieties, and forms, of which 278 taxa are nonnative; Carter (2015) provided a list of 157 bryophyte species from the islands. Within animals, vertebrates are best documented: Mammalia—34 species, including 14 native and 20 nonnative (von Bloeker, 1965); Amphibia—eight species (Nafis, 2022); Reptilia—16 species and subspecies (Nafis, 2022); Aves—422 species (Collins & Jones, 2015). Several non-beetle insect groups have published inventories for all California Channel Islands: Dermaptera—two species, both nonnative (Miller, 1984); Orthoptera—54 species (Weissman & Rentz, 1976; Weissman, 1985), list/guide published in Rentz & Weissman (1982); Hemiptera: Sternorrhyncha: Pseudococcidae—43 species (Rust, Menke & Miller, 1985), list published in same; Lepidoptera—purportedly >800 species (Powell, 1994, 2005), but no species list published; Hymenoptera: Apoidea—243 species (Rust, Menke & Miller, 1985), list published in same.
Prior efforts at inventorying the beetles of the California Channel Islands have been significant yet have not resulted in a published list of species for over a century. Fall (1897) made the first attempt at a comprehensive list of the species of Coleoptera of the southern California islands, including the Channel Islands and Guadalupe Island, Mexico. He recorded 212 species from the Channel Islands. Fall & Davis (1934) recorded 51 species of Coleoptera from Santa Cruz Island. Darlington (1943: 54–55) made general observations about the carabid beetle fauna of the “Santa Barbara Islands” but presented no new taxon or island records. Miller (1985a) recorded 36 endemic beetle species from the Channel Islands (not including the 10 endemic species of Trigonoscuta), plus two additional endemic subspecies. Miller & Miller (1985) reported 52 species of Coleoptera from Santa Barbara Island. Elbert L. Sleeper purportedly had an unpublished beetle list for Santa Catalina Island, with 330 species (see Caterino & Chandler, 2010). Scott E. Miller had an unpublished checklist of 225 beetle species prior to the efforts of the California Beetle Project (Caterino, Chatzimanolis & Richmond, 2015: 279). Caterino, Chatzimanolis & Richmond (2015) reported on an unpublished list of “over 640 named species of beetles” from the Channel Islands.
We here report, for the first time since 1897, a comprehensive list of Coleoptera species from the Channel Islands, annotated with full supporting citations, specimen records, nomenclatural authority, and notes.
Materials and Methods
To generate a comprehensive list of the Coleoptera known from the Channel Islands, we utilized two primary data streams: published literature and digitized specimen records. We also physically examined specimens from several institutions to complement and refine the specimen-level data delivered by them. Our methods generally follow those suggested by Johnston, Aalbu & Franz (2018) to create a thoroughly traceable checklist to encourage a verified, reproducible, and readily updatable product. Specific methods for each data type are discussed below with an assessment of their strengths and limitations.
Harvesting published literature records
The taxonomic and faunistic literature for Coleoptera of North America is incredibly expansive and intractable for a single research team to fully scour. Nevertheless, a rigorous review of the literature was attempted to identify Channel Islands distribution records. The comprehensive bibliographies for California Island entomology by Miller & Menke (1981), Miller (1985b), and Miller (1993) were used as a baseline. We then examined literature sources we were familiar with, particularly those published after the last (1993) supplement, and finally broadened the literature search by searching for keywords of “Channel Islands” and “Coleoptera” as well as by examining modern works that treated species and genera of beetles known to occur in California. Gray literature was generally not consulted for additional species or island records, although specimens resulting from such works made available as voucher specimens in institutional collections were frequently encountered. The result, we believe, is a thorough baseline literature review across all relevant publications. Every publication was vetted by us and a full citation with page number was generated for every island record of a taxon the publication presented.
Literature records, especially those from authoritative taxonomic revisions, can provide some of the best information available for Channel Island Coleoptera. In particular, revisionary studies are often based upon borrowed material from many institutions and likely report on specimens that have not yet been digitized and made available by the owning institutions. Conversely, historical literature records may often be doubtful as taxonomic names and concepts have shifted through time. Additionally, some publications do not cite particular specimens so a proper vetting of the island records may never be fully possible.
Future revisionary works and taxon-specific studies may overturn some of the records reported in the literature. However, we have here reported all such taxa and literature citations in order to make them transparent for future researchers. In the event of a publication explicitly discounting earlier published records or ascribing them to new taxa, the original citations are included in the notes under each taxon, while the island records presented have been adjusted according to more recent authorities. Notes under each taxon detail any perceived ambiguities, irregularities, or importance for each literature citation.
Harvesting digitized specimen records
Natural history collections house the primary distributional data for insects. Each collection has idiosyncratic strengths which are often a result of the activities of its workers through time. For the Channel Islands, material is scattered throughout the world’s collections and even collections with limited holdings from the region may contain valuable species records that correspond to taxonomic expertise of its staff. Visiting all collections and examining every island beetle within them is impractical and inefficient for building a checklist. Therefore, we have focused on publicly available digitized specimen records for Channel Islands Coleoptera.
Data sources. Our dataset is built upon three primary groups of specimen records: (1) the beetle holdings of the Natural History Museum of Los Angeles County (LACM), which contains vast amounts of historical island survey material; (2) the beetle holdings of the Santa Barbara Museum of Natural History (SBMNH), which has a focus on both Coleoptera and the Channel Islands; and (3) digitized records available from the Symbiota Collections of Arthropods Network (SCAN, https://scan-bugs.org). The holdings from LACM and SBMNH are fully (or very nearly) digitized to the specimen level. The records from SCAN were compiled by performing two searches of the portal (in November 2020): (1) taxon “Coleoptera” within a polygon drawn around all eight Channel Islands, and (2) taxon “Coleoptera” with state “California” and “island” contained in the locality. Specimens from the following institutional or personal collections appear in our checklist:
- ASUHIC
-
Arizona State University Hasbrouck Insect Collection, Tempe, AZ, USA
- AUMNH
-
Auburn University Museum of Natural History, Auburn, AL, USA
- BYUC
-
Brigham Young University Arthropod Collection, Provo, UT, USA
- CASC
-
California Academy of Sciences, San Francisco, CA, USA
- CSCA
-
California State Collection of Arthropods, Sacramento, CA, USA
- CSUC
-
Colorado State University Insect Collection, Fort Collins, CO, USA
- DMNS
-
Denver Museum of Nature and Science, Denver, CO, USA
- EMEC
-
Essig Museum, University of California Berkeley, Berkeley, CA, USA
- JNRC
-
Jacques N. Rifkind Collection, Sacramento, CA, USA
- LACM
-
Natural History Museum of Los Angeles County, Los Angeles, CA, USA
- MAJC
-
M. Andrew Johnston Research Collection, Tempe, AZ, USA
- SBMNH
-
Santa Barbara Museum of Natural History, Santa Barbara, CA, USA
- SDNHM
-
San Diego Natural History Museum, San Diego, CA, USA
- SEMC
-
Snow Entomological Museum, University of Kansas, Lawrence, KS, USA
- TAMU
-
Texas A&M University Insect Collection, College Station, TX, USA
- UCMC
-
University of Colorado Museum of Natural History, Boulder, CO, USA
- UCRC
-
University of California Riverside Insect Collection, Riverside, CA, USA
- UCSB
-
University of California Santa Barbara, Santa Barbara, CA, USA
- USNM
-
United States National Museum of Natural History (Smithsonian Institution), Washington, DC, USA
- UTCI
-
University of Tennessee Chattanooga, Chattanooga, TN, USA
- UASM
-
University of Alberta Strickland Entomology Museum, Edmonton, AB, Canada
- YPMC
-
Yale Peabody Museum of Natural History, New Haven, CT, USA
- iNat
-
iNaturalist Research Grade Observations (https://inaturalist.org)
Specimen determinations. Specimen records came with determination information from the original data providers. In addition, 3,309 taxonomic redeterminations/annotations were made by us. Most of these (2,352) were performed on specimens examined in person at SBMNH, LACM, and the UCSB collections. Additional nomenclatural adjustments (957) were made for records that had obvious misspellings or old combinations.
Data cleaning. All data were imported into the Symbiota portal Ecdysis (https://serv.biokic.asu.edu/ecdysis/) utilizing best practices according to the Darwin Core data standard and FAIR (Findable, Accessible, Interoperable, Reusable) data principles. All records were examined and georeferenced (where not previously done) and added to island-specific datasets. California “island” records not located in the Channel Islands (e.g., “Farallon Islands”) were excluded. Throughout this process, the owner institution and metadata were preserved with each record. Records that were deemed untrustworthy, typically due to a mismatch in locality data and provided GPS coordinates or records lacking any data, were pruned from the dataset. The final set of records numbered 26,609; these are fully available in their final, cleaned form (Johnston & Gimmel, 2022).
Excluded specimen data. Digitized data for insects is not yet as complete or mature as for other groups of organisms (e.g., vertebrates and plants). Many collections have no specimens digitized and most are only partially digitized. This issue is compounded by the fact that not all museums share their data publicly or do not frequently refresh their data to online aggregators. In addition, online taxonomic resources are woefully incomplete for insects, particularly beetles, so many publicly available records are not appropriately indexed to family or order level. With each of these hurdles, otherwise valuable records are in effect made unavailable to research projects like this one. Other websites (e.g., California Beetle Project) occasionally provided additional species or specimen information, but these were ignored as unverifiable since they do not have a unique identifier to relocate the presumed specimen(s) anchoring the record. We hope to see increased focus on the Channel Islands by coleopterists in the future where taxonomic experts can continue to add to and refine the knowledge aggregated and summarized in this work.
Checklist validation
Each taxon listed from the islands was critically examined as part of our literature review. In addition, we examined any records of taxa not known from southern California and, where possible, confirmed the identification of the physical specimen. The most modern and reliable treatments for all taxa were used for determining taxon validity, and were cited in full.
Non-unique order, family, or genus records in the literature were generally ignored for purposes of this checklist. For example, if a record was identified only to the genus level where a species from that genus was already known from the Channel Islands, that was not considered a new taxon for the tally. In cases where only genus-level or higher records are known for a given taxon, then that taxon was included in the species count as an undetermined species of the genus (or “undetermined genus and species” for a family in one case). All digitized and literature records are included for the genus level; many of these represent unique island records for that group. Subspecies were not counted as separate taxa in our checklist; instead, subspecies are discussed under each species where relevant.
Numbers quoted in the family accounts for California beetle diversity are mostly derived from an unpublished checklist of author MLG with other sources being cited when used. We do not include general biological information except in special cases and except as it relates specifically to island-collected or island-observed specimens; such biological information can be found in more general guides and references.
We cite the most relevant work where we derived our taxon name, combination, and authority from as the nomenclatural authority. This is often the most recent catalog, revision, or book chapter known to us. Many of the groups represented across the California Channel Islands are in desperate need of revision and have a long and complicated taxonomic history. We anticipate that taxonomic experts will come to conclusions different than those of historical workers who have published on the Channel Islands and identified material in collections. It is with this in mind that we strive to explicitly document all name usages such that they can be tracked, validated, and updated by future generations of coleopterists (see also Johnston, Aalbu & Franz, 2018).
Biogeographic and diversity analyses
Our final species list and dataset of digitized specimen records were analyzed to explore trends and correlations using R (R Core Team, 2022). Species diversity from each island was plotted against geographic and rainfall data for each island (taken from Miller, 1985a) using linear models in the R ggplot2 software package (Wickham, 2016). The final digitized record dataset (i.e., not counting literature records but including identifications to higher taxon ranks as a single unique taxon) was used to generate specimen totals for each unique taxon on each island and totals for each taxon pooled across all of the islands. Totals for each species by island were further pooled into collecting events where all records with the same collector (recordedBy field) and collection date (eventDate field) were considered to belong to the same collecting event to examine possible limitations of the dataset. Total specimen counts were analyzed in a rarefaction and extrapolation species diversity analysis using the R iNEXT software package (Chao et al., 2014) to generate both rarefaction curves and an estimate of actual species diversity given the observed data. An annotated R script with raw data is available via Zenodo (Johnston, 2022).
Results and Discussion
We here provide an annotated checklist of 825 unique species (= taxa) comprising 1,829 species-by-island records (see checklist below). Individual island species counts and geographic data are given in Table 1. Curated digitized specimen records are archived and available on Zenodo (Johnston & Gimmel, 2022).
| Islands | Species | DigRecords | Area (km2) | Distance (km) |
|---|---|---|---|---|
| Anacapa | 74 | 814 | 2.9 | 61 |
| San Clemente | 197 | 5,092 | 145 | 79 |
| San Miguel | 138 | 2,795 | 37 | 42 |
| San Nicolas | 146 | 2,946 | 58 | 98 |
| Santa Barbara | 64 | 751 | 2.6 | 61 |
| Santa Catalina | 370 | 3,068 | 194 | 32 |
| Santa Cruz | 503 | 7,341 | 249 | 30 |
| Santa Rosa | 337 | 3,802 | 217 | 44 |
Note:
Species and DigRecords (digitized records) are counts from the checklist and dataset reported in this study. Area and distance from mainland for each island are taken from Miller (1985a).
We report 56 species putatively restricted to the Channel Islands along with two more that are questionably so. This represents a 22% increase in the number of known endemic species since Miller (1985a), which is mostly accounted for by recognition of newly described and still-undescribed endemic species. An additional 52 species in the fauna are not native to the southern California region.
Beetle diversity on individual islands was plotted against island and dataset characteristics to elucidate potential driving factors and biases in our results. Beetle diversity is extremely strongly correlated with island land area with a linear relationship (Fig. 1A) but not with island distance to mainland (Fig. 1B). Interestingly, species richness was linearly, and not logarithmically, correlated with island area, which is counter to Darlington’s rule hypothesizing a doubling of species for each ten-fold increase in island area (Darlington, 1957). Our finding is similar to that of Moody (2000) for native vascular plants on the Channel Islands; however, Powell (1994) found only a weak species-area relationship in Channel Islands Lepidoptera. While observed species richness could be the result of bias in sampling effort, the number of species on an island did not strongly correlate with the number of digitized records (Fig. 2A). The number of records showed a similar relationship with the size of the island (Fig. 2B), perhaps indicating a somewhat even sampling per island area.
Figure 1: Species by island size and position.
(A) Plotted against total area (km2). (B) Plotted against distance to mainland (km).
Figure 2: Correlation of digitized specimen records to number of species and island size.
(A) Species by digitized records. (B) Digitized records by island area.
The distribution of the number of islands a single species inhabits (Fig. 3A) was strongly left-skewed with over 50% being recorded from just a single island. However, numbers of specimens and collecting events per species across all islands were also highly left-skewed (Figs. 3B and 3C); in fact, 154 species (19% of the fauna) in our list are represented by a single digitized specimen (Fig. 3B) while 248 species (30% of the fauna) are represented by a single digitized collecting event (i.e., a series of specimens which had identical values for collector and date) (Fig. 3C). The observed distribution patterns may therefore not be a true reflection of biological diversity but are likely subject to bias from insufficient sampling.
Figure 3: Species distribution and collection frequency.
(A) Number of islands each species inhabits. (B) Raw number of digitized records per species. (C) Unique collecting events per species.
Species richness on each island is likely still far from fully documented (Table 2; Fig. 4). The entire island beetle fauna seems to have at least several hundred species still undocumented (Fig. 4C). Interestingly, the smallest islands, Anacapa (Fig. 4A) and Santa Barbara (Fig. 4B), appear to be the least completely inventoried (62.1% and 63.1% estimated complete, respectively; Table 2). Santa Cruz, at 87.7% estimated complete, appears to be the most completely inventoried island (Fig. 4A; Table 2).
| Islands | Observed taxa | Estimated taxa | Estimated % complete | s.e. | 95% LCL | 95% UCL |
|---|---|---|---|---|---|---|
| San Clemente | 164 | 197.992 | 82.8% | 15.424 | 178.557 | 243.374 |
| San Nicolas | 147 | 186.371 | 78.9% | 18.771 | 163.215 | 242.59 |
| Santa Barbara | 46 | 72.955 | 63.1% | 17.593 | 54.386 | 132.636 |
| Santa Catalina | 300 | 418.221 | 71.7% | 34.606 | 367.396 | 507.375 |
| Anacapa | 79 | 127.221 | 62.1% | 27.169 | 96.231 | 213.95 |
| San Miguel | 136 | 164.254 | 82.8% | 13.35 | 147.721 | 204.106 |
| Santa Cruz | 569 | 648.879 | 87.7% | 20.09 | 618.162 | 698.79 |
| Santa Rosa | 326 | 406.031 | 80.3% | 24.022 | 371.004 | 468.318 |
| All combined | 832 | 966.744 | 86.1% | 28.507 | 921.412 | 1,035.061 |
Note:
Observed and estimated taxa are from rarefaction analyses based upon digitized records (excludes taxa only known from literature records). Estimated percent complete is the proportion of the estimated taxa already observed. s.e. = standard error; LCL and UCL are lower and upper 95% confidence levels, respectively.
Figure 4: Species diversity rarefaction and estimation curves.
(A) North islands. (B) South islands. (C) All islands combined.
The ten most species-rich families on the islands are: Staphylinidae (105 species), Carabidae (87 species), Curculionidae (65 species), Tenebrionidae (61 species), Coccinellidae (43 species), Scarabaeidae (41 species), Chrysomelidae (38 species), Cerambycidae (34 species), Hydrophilidae (32 species), and Melyridae (27 species). The three most species-rich genera on the islands are, with 11 species each: Eleodes (Tenebrionidae), Scymnus (Coccinellidae), and Trigonoscuta (Curculionidae); Bembidion (Carabidae) follows closely behind with 10 species. Table 3 lists all species reported in the annotated checklist below and summarizes their status (native vs adventive) and known island-level distribution.
| Scientific name | Duplicate genus record | Endemic | Adventive | Anacapa | San Clemente | San Miguel | San Nicolas | Santa Barbara | Santa Catalina | Santa Cruz | Santa Rosa |
|---|---|---|---|---|---|---|---|---|---|---|---|
| ADEPHAGA | |||||||||||
| Carabidae | |||||||||||
| Agonum decorum | D,L | ||||||||||
| Agonum limbatum | D | D,L | D | D | |||||||
| Agonum piceolum | D | ||||||||||
| Agonum punctiforme | D | D | D | ||||||||
| Akephorus marinus | D | D | D | D,L | |||||||
| Amara aurata | D,L | D | |||||||||
| Amara californica | D | D,L | D | D | D,L | D,L | |||||
| Amara conflata | D | ||||||||||
| Amara insignis | D,L | D | L | ||||||||
| Amara insularis | end | D | D,L | D | D,L | D,L | D | D | D,L | ||
| Amara pomona | L | L | |||||||||
| Amara scitula | D | ||||||||||
| Anchomenus funebris | L | L | D,L | ||||||||
| Anisodactylus californicus | D | D,L | D | D,L | D,L | D,L | |||||
| Anisodactylus consobrinus | L | D,L | D,L | ||||||||
| Anisodactylus similis | D,L | D | |||||||||
| Apristus | dup | D | |||||||||
| Apristus pugetanus | D | ||||||||||
| Axinopalpus biplagiatus | D | D | |||||||||
| Bembidion corgenoma | D | D,L | D,L | ||||||||
| Bembidion ephippigerum | D | L | |||||||||
| Bembidion indistinctum | L | L | |||||||||
| Bembidion insulatum | L | ||||||||||
| Bembidion iridescens | L | ||||||||||
| Bembidion laticeps | D | ||||||||||
| Bembidion palosverdes | D,L | ||||||||||
| Bembidion platynoides | L | ||||||||||
| Bembidion striola | L | L | |||||||||
| Bembidion versicolor | L | ||||||||||
| Brachinus costipennis | L | D,L | D | ||||||||
| Brachinus gebhardis | D | D,L | D | ||||||||
| Brachinus mexicanus | D,L | ||||||||||
| Brachinus quadripennis | L | ||||||||||
| Bradycellus californicus | D | D | D | D | D | D,L | |||||
| Bradycellus nitidus | D | D | D,L | D,L | D,L | ||||||
| Bradycellus rupestris | D | D | L | D | D | ||||||
| Bradycellus sejunctus | D | ||||||||||
| Calathus ruficollis | D,L | L | D,L | D,L | D,L | D,L | |||||
| Calosoma eremicola | D,L | D,L | |||||||||
| Calosoma parvicolle | L | ||||||||||
| Calosoma semilaeve | L | L | D,L | D | L | ||||||
| Chlaenius cumatilis | D,L | D | |||||||||
| Chlaenius obsoletus | L | D | |||||||||
| Chlaenius tricolor | D | ||||||||||
| Chlaenius variabilipes | D | D,L | D | ||||||||
| Dicheirus dilatatus | D,L | D | D,L | D,L | D,L | ||||||
| Dicheirus piceus | D | D,L | D,L | D | D,L | ||||||
| Dromius piceus | D | D | |||||||||
| Dyschirius aratus | D | ||||||||||
| Dyschirius consobrinus | D | ||||||||||
| Dyschirius gibbipennis | D | D | D,L | ||||||||
| Dyschirius varidens | D | ||||||||||
| Elaphropus undet. sp. | D | ||||||||||
| Harpalus | dup | D | |||||||||
| Harpalus caliginosus | D | D | |||||||||
| Harpalus pensylvanicus | D | ||||||||||
| Lachnophorus elegantulus | D | ||||||||||
| Laemostenus complanatus | adv | D | D | ||||||||
| Lebia cyanipennis | D | D | |||||||||
| Lebia perita | D | D | |||||||||
| Microlestes undet. sp. | D | ||||||||||
| Notiophilus semiopacus | L | D | |||||||||
| Omophron dentatum | D | D,L | D,L | ||||||||
| Phrypeus rickseckeri | D | ||||||||||
| Platynus brunneomarginatus | D | D | D,L | D,L | D,L | ||||||
| Poecilus laetulus | D | D,L | L | L | |||||||
| Pterostichus gliscans | end | L | D | ||||||||
| Pterostichus illustris | D | ||||||||||
| Pterostichus inermis | D | ||||||||||
| Pterostichus isabellae | L | L | |||||||||
| Pterostichus jacobinus | D | ||||||||||
| Pterostichus lustrans | D,L | ||||||||||
| Pterostichus menetriesii | D | D | L | ||||||||
| Scaphinotus crenatus | D | D | D | D | |||||||
| Scaphinotus punctatus | D,L | ||||||||||
| Scaphinotus ventricosus | D,L | ||||||||||
| Schizogenius depressus | D | L | |||||||||
| Stenolophus anceps | D | D | D | ||||||||
| Stenolophus flavipes | D | D | D | ||||||||
| Stenolophus limbalis | L | ||||||||||
| Stenolophus lineola | D | D | D,L | ||||||||
| Stenolophus ochropezus | D | ||||||||||
| Stenolophus rugicollis | D | ||||||||||
| Tachys corax | L | D | |||||||||
| Tachys vittiger | L | ||||||||||
| Tachys vorax | D | ||||||||||
| Tanystoma cuyama | D | ||||||||||
| Tanystoma maculicolle | L | D,L | D,L | D,L | D,L | D,L | D,L | ||||
| Thalassotrechus barbarae | D | D | |||||||||
| Cicindelidae | |||||||||||
| Cicindela hirticollis | L | L | |||||||||
| Cicindela oregona | D,L | D,L | D,L | L | D,L | D,L | |||||
| Cicindela senilis | D,L | ||||||||||
| Cicindelidia hemorrhagica | D,L | D,L | |||||||||
| Cicindelidia trifasciata | D,L | ||||||||||
| Dytiscidae | |||||||||||
| Agabinus glabrellus | D,L | D,L | D | ||||||||
| Agabinus sculpturellus | L | ||||||||||
| Agabus obsoletus | D | ||||||||||
| Dytiscus marginicollis | D | D,L | |||||||||
| Eretes sticticus | D | ||||||||||
| Hydrovatus brevipes | L | ||||||||||
| Hygrotus lutescens | D | D | D,L | D | |||||||
| Ilybiosoma lugens | D | D,L | D | D,L | |||||||
| Ilybiosoma regulare | D | D | |||||||||
| Ilybiosoma seriatum | D | D | |||||||||
| Ilybius discors | L | ||||||||||
| Ilybius lineellus | D | ||||||||||
| Ilybius walsinghami | L | D | |||||||||
| Laccophilus fasciatus | D | D,L | |||||||||
| Laccophilus maculosus | D,L | ||||||||||
| Leconectes striatellus | D | D | D,L | D,L | D,L | ||||||
| Liodessus obscurellus | D | D | D | D | |||||||
| Neoclypeodytes pictodes | D | ||||||||||
| Rhantus gutticollis | D,L | D | D,L | D,L | D | ||||||
| Sanfilippodytes barbarensis | D | D | D | ||||||||
| Sanfilippodytes latebrosus | D | D | D | ||||||||
| Sanfilippodytes vilis | D | D | L | L | |||||||
| Sanfilippodytes williami | L | D | L | ||||||||
| Uvarus subtilis | D | D | |||||||||
| Gyrinidae | |||||||||||
| Gyrinus plicifer | D | D,L | D | ||||||||
| Haliplidae | |||||||||||
| Haliplus undet. sp. | D | ||||||||||
| Peltodytes simplex | D,L | D,L | D | ||||||||
| MYXOPHAGA | |||||||||||
| Hydroscaphidae | |||||||||||
| Hydroscapha natans | D,L | D | |||||||||
| Sphaeriusidae | |||||||||||
| Sphaerius politus | D | D | |||||||||
|
SCIRTOIDEA Scirtidae |
|||||||||||
| Undet. genus, undet. sp. | L | ||||||||||
|
CLAMBOIDEA Clambidae |
|||||||||||
| Clambus undet. sp. | D | ||||||||||
| Loricaster rotundus | D | D | |||||||||
|
DASCILLOIDEA Dascillidae |
|||||||||||
| Anorus piceus | D | D | D | D | |||||||
| Rhipiceridae | |||||||||||
| Sandalus cribricollis | D | ||||||||||
|
BUPRESTOIDEA Buprestidae |
|||||||||||
| Acmaeodera hepburnii | D,L | D,L | D,L | ||||||||
| Acmaeodera mariposa | D | ||||||||||
| Acmaeodera prorsa | D | D,L | |||||||||
| Agrilus quadriguttatus | D | ||||||||||
| Anthaxia aeneogaster | D | ||||||||||
| Buprestis aurulenta | D,L | ||||||||||
| Chrysobothris mali | D | D | |||||||||
| Melanophila consputa | D | ||||||||||
|
DRYOPOIDEA Dryopidae |
|||||||||||
| Postelichus productus | L | ||||||||||
| Elmidae | |||||||||||
| Ordobrevia nubifera | L | ||||||||||
| Heteroceridae | |||||||||||
| Heterocerus | dup | D | D | D | D | ||||||
| Heterocerus mexicanus | D,L | ||||||||||
| Limnichidae | |||||||||||
| Limnichites nebulosus | D | ||||||||||
|
ELATEROIDEA Cantharidae |
|||||||||||
| Cultellunguis americanus | L | ||||||||||
| Cultellunguis hatchi | L | D,L | |||||||||
| Frostia laticollis | D,L | ||||||||||
| Pacificanthia consors | D | D,L | D | ||||||||
| Podabrus pruinosus | D | ||||||||||
| Podabrus undet. sp. | D | ||||||||||
| Silis | dup | D | |||||||||
| Silis carmelita | D | ||||||||||
| Elateridae | |||||||||||
| Ampedus longicornis | D,L | ||||||||||
| Ampedus rhodopus | D | ||||||||||
| Anchastus cinereipennis | D | D,L | L | D | D | ||||||
| Athous axillaris | D | D | |||||||||
| Athous nigropilis | D | ||||||||||
| Athous rufiventris | D | D | |||||||||
| Cardiophorus | dup | D | |||||||||
| Cardiophorus tenebrosus | D | ||||||||||
| Dalopius | dup | D | |||||||||
| Dalopius luteolus | D | ||||||||||
| Dalopius undet. sp. | D | D | |||||||||
| Elater lecontei | D | ||||||||||
| Euthysanius lautus | D | D | |||||||||
| Hemicrepidius californicus | D | D | |||||||||
| Hemicrepidius tumescens | L | ||||||||||
| Heteroderes amplicollis | adv | D | |||||||||
| Horistonotus inanus | D | D | |||||||||
| Limonius canus | D | D | |||||||||
| Melanactes densus | D | ||||||||||
| Melanotus longulus | D,L | D | D | ||||||||
| Octinodes frater | D | ||||||||||
| Paradonus inops | D | ||||||||||
| Eucnemidae | |||||||||||
| Asiocnemis hospitalis | D,L | ||||||||||
| Lampyridae | |||||||||||
| Pterotus obscuripennis | D | ||||||||||
| Pyropyga nigricans | D | D | |||||||||
| Phengodidae | |||||||||||
| Zarhipis integripennis | D | ||||||||||
| Throscidae | |||||||||||
| Trixagus sericeus | D | ||||||||||
|
HISTEROIDEA Histeridae |
|||||||||||
| Aphelosternus interstitialis | L | ||||||||||
| Bacanius undet. sp. | D,L | ||||||||||
| Carcinops opuntiae | D | ||||||||||
| Euspilotus scissus | D | D | D | D | |||||||
| Euspilotus sp. near laridus | L | ||||||||||
| Geomysaprinus undet. sp. | D | D | |||||||||
| Halacritus maritimus | D | D | |||||||||
| Hololepta vicina | L | ||||||||||
| Hypocaccus bigemmeus | D | D | D | D | D | ||||||
| Hypocaccus gaudens | D | D | D | D | D | ||||||
| Hypocaccus lucidulus | D,L | D,L | D,L | D,L | D,L | ||||||
| Hypocaccus serrulatus | D | ||||||||||
| Iliotona cacti | D | ||||||||||
| Margarinotus sexstriatus | D,L | L | |||||||||
| Neopachylopus sulcifrons | D | D | D,L | D | D | D | |||||
| Plegaderus undet. sp. | D | ||||||||||
| Saprinus lugens | L | D | D,L | D,L | D | D,L | D,L | ||||
| Saprinus oregonensis | L | ||||||||||
| Xerosaprinus fimbriatus | L | ||||||||||
| Xerosaprinus lubricus | L | D,L | D,L | D,L | |||||||
| Xerosaprinus vitiosus | L | ||||||||||
|
HYDROPHILOIDEA Helophoridae |
|||||||||||
| Helophorus linearis | D | ||||||||||
| Hydrophilidae | |||||||||||
| Agna capillata | D | ||||||||||
| Anacaena signaticollis | D | D,L | D | ||||||||
| Berosus fraternus | L | ||||||||||
| Berosus hatchi | L | ||||||||||
| Berosus infuscatus | D | ||||||||||
| Berosus punctatissimus | D | D,L | D | ||||||||
| Cercyon fimbriatus | D | D,L | D | D,L | D | D | |||||
| Cercyon haemorrhoidalis | adv | L | D | ||||||||
| Cercyon luniger | D | L | L | D,L | |||||||
| Cercyon quisquilius | adv | D | D | ||||||||
| Chaetarthria hespera | D,L | D | |||||||||
| Chaetarthria nigrella | D | D | |||||||||
| Chaetarthria punctulata | D | ||||||||||
| Chaetarthria pusilla | D | ||||||||||
| Cymbiodyta columbiana | D | ||||||||||
| Cymbiodyta dorsalis | D,L | D | D,L | D,L | D,L | ||||||
| Cymbiodyta punctatostriata | D,L | ||||||||||
| Enochrus carinatus | L | D | |||||||||
| Enochrus cristatus | D | ||||||||||
| Enochrus hamiltoni | D | ||||||||||
| Enochrus piceus | D | D | D | D | |||||||
| Enochrus pygmaeus | D | ||||||||||
| Helochares normatus | D,L | D | |||||||||
| Hydrobius fuscipes | L | ||||||||||
| Hydrochara lineata | D,L | L | |||||||||
| Hydrophilus triangularis | D | D,L | |||||||||
| Laccobius californicus | D,L | ||||||||||
| Laccobius ellipticus | D,L | D,L | D,L | ||||||||
| Laccobius insolitus | D | D | |||||||||
| Sphaeridium scarabaeoides | adv | D | D | D | |||||||
| Tropisternus affinis | D,L | D | D | ||||||||
| Tropisternus californicus | L | L | |||||||||
|
SCARABAEOIDEA Geotrupidae |
|||||||||||
| Bolbocerastes regalis | L | ||||||||||
| Odonteus obesus | D | ||||||||||
| Scarabaeidae | |||||||||||
| Aegialia convexa | L | ||||||||||
| Aegialia crassa | L | ||||||||||
| Aegialia nigrella | L | ||||||||||
| Aegialia punctata | L | ||||||||||
| Amblonoxia palpalis | D | L | D,L | L | |||||||
| Aphodius fimetarius | adv | D | D | D | |||||||
| Calamosternus granarius | adv | D | D | D | D,L | ||||||
| Canthon simplex | L | ||||||||||
| Cinacanthus militaris | L | ||||||||||
| Coenonycha clementina | end | D,L | |||||||||
| Coenonycha clypeata | end | D,L | |||||||||
| Coenonycha fulva | end | D | D,L | ||||||||
| Coenonycha santacruzae | end | D,L | |||||||||
| Cotinis mutabilis | D | ||||||||||
| Cremastocheilus schaumii | D,L | ||||||||||
| Cyclocephala borealis | L | L | |||||||||
| Cyclocephala hirta | D | ||||||||||
| Cyclocephala longula | L | L | L | L | |||||||
| Cyclocephala melanocephala | L | L | L | ||||||||
| Cyclocephala pasadenae | L | L | L | ||||||||
| Dichelonyx backii | D | ||||||||||
| Dichelonyx fulgida | L | ||||||||||
| Dichelonyx pusilla | D,L | D,L | D,L | ||||||||
| Diplotaxis fimbriata | L | L | |||||||||
| Diplotaxis subangulata | D,L | D,L | D,L | ||||||||
| Hoplia callipyge | D,L | ||||||||||
| Labarrus pseudolividus | D | D,L | D,L | D | L | D,L | D,L | ||||
| Ligyrus gibbosus | D,L | D,L | D,L | D | D,L | ||||||
| Otophorus haemorrhoidalis | adv | D | |||||||||
| Phobetus | dup | D,L | D | L | D | ||||||
| Phobetus ciliatus | end | D,L | |||||||||
| Phobetus testaceus | end | D,L | |||||||||
| Phyllophaga mucorea | L | ||||||||||
| Planolinellus vittatus | D,L | D,L | |||||||||
| Polyphylla | dup | L | |||||||||
| Polyphylla crinita | D | D,L | L | ||||||||
| Polyphylla nigra | D | D,L | L | ||||||||
| Rugaphodius rugatus | D,L | D,L | |||||||||
| Serica alternata | L | L | L | ||||||||
| Serica catalina | end | L | |||||||||
| Serica cruzi | end | D,L | |||||||||
| Serica mixta | L | L | L | L | L | ||||||
| Tesarius mcclayi | D | D | |||||||||
| Trogidae | |||||||||||
| Trox atrox | D,L | ||||||||||
| Trox gemmulatus | D,L | ||||||||||
|
STAPHYLINOIDEA Colonidae |
|||||||||||
| Colon forceps | D | ||||||||||
| Hydraenidae | |||||||||||
| Hydraena | dup | D | D | ||||||||
| Hydraena arenicola | L | ||||||||||
| Hydraena circulata | L | ||||||||||
| Hydraena vandykei | L | ||||||||||
| Ochthebius | dup | D | D | D | D | ||||||
| Ochthebius discretus | L | ||||||||||
| Ochthebius interruptus | L | ||||||||||
| Ochthebius puncticollis | D,L | ||||||||||
| Leiodidae | |||||||||||
| Agathidium pulchrum | D | ||||||||||
| Agathidium virile | D | D | |||||||||
| Leiodes antennata | D | ||||||||||
| Leiodes paludicola | D | ||||||||||
| Pinodytes gibbosus | D,L | D,L | D,L | ||||||||
| Ptiliidae | |||||||||||
| Acrotrichis undet. sp. | D | D | |||||||||
| Actidium undet. sp. | D | ||||||||||
| Motschulskium sinuatocolle | D | D | D | ||||||||
| Ptenidium undet. sp. | D | ||||||||||
| Pteryx undet. sp. | D | D | |||||||||
| Ptiliolum undet. sp. | D | D | D | ||||||||
| Staphylinidae | |||||||||||
| Acrotona | dup | D | D | ||||||||
| Acrotona recondita | L | ||||||||||
| Acrotona sonomana | L | ||||||||||
| Actium californicum | L | ||||||||||
| Actium vestigialis | end | D,L | |||||||||
| Adota maritima | L | ||||||||||
| Aleochara bimaculata | L | L | |||||||||
| Aleochara curtidens | L | ||||||||||
| Aleochara densissima | L | ||||||||||
| Aleochara fumata | adv | L | |||||||||
| Aleochara lanuginosa | adv | D,L | |||||||||
| Aleochara littoralis | L | L | |||||||||
| Aleochara sulcicollis | D | D,L | D,L | D | D,L | D,L | |||||
| Aleochara valida | D | D | L | D | |||||||
| Aploderus | dup | D | D | ||||||||
| Aploderus flavipennis | L | ||||||||||
| Apocellus analis | L | ||||||||||
| Astenus undet. sp. | D | D | |||||||||
| Atheta hampshirensis | D | ||||||||||
| Belonuchus ephippiatus | D | D | |||||||||
| Bisnius albionicus | D | D | |||||||||
| Bisnius sordidus | adv | D | D | ||||||||
| Bledius albonotatus | D,L | D,L | D | D | D | ||||||
| Bledius fenyesi | D | D,L | D,L | D | D,L | D | |||||
| Bledius opacifrons | D | D | |||||||||
| Bledius ruficornis | D,L | D | D | ||||||||
| Blepharhymenus undet. sp. | D | D | D | ||||||||
| Brachycepsis undet. sp. | L | D,L | D | ||||||||
| Bryoporus rufescens | D | D,L | D | ||||||||
| Bryothinusa catalinae | D,L | ||||||||||
| Cafius canescens | D | D,L | L | D | D | ||||||
| Cafius lithocharinus | D | D | D,L | D | D | D,L | |||||
| Cafius luteipennis | D | D | D | D,L | D | D,L | |||||
| Cafius opacus | L | ||||||||||
| Cafius seminitens | D | D,L | D,L | D | D | ||||||
| Cafius sulcicollis | D | L | D,L | ||||||||
| Carpelimus undet. sp. | D | D | D | D | D | ||||||
| Cephennium urbanum | D,L | ||||||||||
| Creophilus maxillosus | D,L | L | D | D | |||||||
| Diaulota fulviventris | D | D | |||||||||
| Diestota undet. sp. | D | D | D | D | D | D | |||||
| Erichsonius puncticeps | D | L | D | D | |||||||
| Euconnus undet. sp. | D | D | D | ||||||||
| Falagriota occidua | D,L | D | |||||||||
| Gabrius | dup | D | |||||||||
| Gabrius nigritulus | adv | D | L | ||||||||
| Gnypeta undet. sp. | D | D | |||||||||
| Habrocerus capillaricornis | adv | D | |||||||||
| Hadrotes crassus | D | D,L | D | D,L | D,L | D,L | D,L | ||||
| Hesperotychus undet. sp. | D,L | ||||||||||
| Heterosilpha ramosa | D | L | D,L | D,L | |||||||
| Heterothops conformis | D | D,L | D | ||||||||
| Heterothops fusculus | L | ||||||||||
| Holobus undet. sp. | D | ||||||||||
| Hydrosmecta undet. sp. | D | ||||||||||
| Leptotyphlinae undet. sp. | D | ||||||||||
| Linohesperus | dup | D | D | ||||||||
| Linohesperus borealis | L | D | |||||||||
| Linohesperus cuspifer | L | ||||||||||
| Lobrathium | dup | D | |||||||||
| Lobrathium jacobinum | L | ||||||||||
| Lordithon thoracicus | D | D | |||||||||
| Medon undet. sp. | D | D | D | D,L | D | ||||||
| Meoticina undet. sp. | D | ||||||||||
| Mycetoporus neotomae | D | D | |||||||||
| Myllaena undet. sp. | D | D | D | D | |||||||
| Neobisnius occidentoides | D,L | D | D | D | |||||||
| Neobisnius sobrinus | D | ||||||||||
| Neobisnius terminalis | D | ||||||||||
| Nicrophorus guttula | D,L | D | D | ||||||||
| Nicrophorus marginatus | D | ||||||||||
| Nicrophorus nigrita | L | L | L | D,L | D,L | D,L | |||||
| Nitidotachinus agilis | D | D | |||||||||
| Nudobius pugetanus | D | ||||||||||
| Oligota undet. sp. | D | D | D | ||||||||
| Omalium algarum | D | ||||||||||
| Oropus undet. sp. | L | D,L | |||||||||
| Orus undet. sp. | D | ||||||||||
| Oxypoda undet. sp. | D | D | D | ||||||||
| Palporus nitidulus | D | D | |||||||||
| Philonthus cruentatus | adv | D | D,L | D | D | ||||||
| Philonthus davus | D | D | |||||||||
| Philonthus flavolimbatus | D | ||||||||||
| Philonthus hepaticus | D | ||||||||||
| Philonthus lecontei | L | ||||||||||
| Philonthus longicornis | adv | L | |||||||||
| Philonthus quadrulus | D | ||||||||||
| Philonthus triangulum | L | ||||||||||
| Phloeopora undet. sp. | D | ||||||||||
| Platystethus americanus | D | ||||||||||
| Pontomalota opaca | D,L | D | D | ||||||||
| Pseudopsis | dup | L | |||||||||
| Pseudopsis minuta | D,L | ||||||||||
| Quedius limbifer | D,L | D | |||||||||
| Sepedophilus castaneus | D | D | |||||||||
| Sonoma | dup | D,L | |||||||||
| Sonoma isabellae | D,L | D,L | |||||||||
| Sonomota undet. sp. | D | D | |||||||||
| Stenichnus undet. sp. | D,L | ||||||||||
| Stictalia undet. sp. | D | D | D | ||||||||
| Sunius | dup | L | |||||||||
| Sunius mobilis | L | ||||||||||
| Sunius reductus | L | ||||||||||
| Tachinus debilis | D | ||||||||||
| Tachyporus | dup | L | |||||||||
| Tachyporus californicus | D | D | L | D | D,L | ||||||
| Tarphiota fucicola | D | D | D | D | |||||||
| Tarphiota geniculata | D | D | D | D | D | D | |||||
| Tasgius ater | adv | D,L | D | L | D | ||||||
| Thanatophilus lapponicus | L | ||||||||||
| Thinobius undet. sp. | D | D | D | ||||||||
| Thinopinus pictus | D,L | D,L | D,L | D,L | D,L | ||||||
| Thinusa fletcheri | D | D | D | D | |||||||
| Thinusa maritima | D,L | ||||||||||
|
BOSTRICHOIDEA Bostrichidae |
|||||||||||
| Amphicerus cornutus | D | D | |||||||||
| Lyctus cavicollis | D | ||||||||||
| Lyctus linearis | adv | D | |||||||||
| Lyctus planicollis | D | ||||||||||
| Melalgus confertus | D | ||||||||||
| Polycaon stoutii | D | L | |||||||||
| Psoa maculata | D | ||||||||||
| Psoa quadrisignata | L | ||||||||||
| Scobicia declivis | L | D | |||||||||
| Scobicia suturalis | D | D | D | ||||||||
| Stephanopachys substriatus | D | ||||||||||
| Dermestidae | |||||||||||
| Anthrenus lepidus | D | D | |||||||||
| Anthrenus verbasci | adv | D,L | |||||||||
| Cryptorhopalum apicale | D,L | ||||||||||
| Cryptorhopalum triste | D | ||||||||||
| Dermestes | dup | D | |||||||||
| Dermestes caninus | L | L | D,L | L | |||||||
| Dermestes frischi | adv | D,L | D,L | D | D,L | D,L | |||||
| Dermestes marmoratus | D,L | L | L | L | |||||||
| Dermestes rattus | D | L | |||||||||
| Dermestes talpinus | L | D | |||||||||
| Megatoma variegata | D | D | |||||||||
| Trogoderma sternale | D,L | D,L | D | D | |||||||
| Ptinidae | |||||||||||
| Actenobius pleuralis | D | ||||||||||
| Byrrhodes ulkei | D | ||||||||||
| Colposternus tenuilineatus | L | D | |||||||||
| Ernobius debilis | D,L | ||||||||||
| Ernobius punctulatus | L | ||||||||||
| Euceratocerus hornii | L | ||||||||||
| Euvrilletta catalinae | end | L | |||||||||
| Euvrilletta occidentalis | D | ||||||||||
| Hemicoelus nelsoni | D | D | |||||||||
| Lasioderma serricorne | adv | D | |||||||||
| Oligomerus delicatulus | D | ||||||||||
| Ozognathus cornutus | D | D | D | ||||||||
| Petalium californicum | D | ||||||||||
| Priobium punctatum | D | D,L | |||||||||
| Ptilinus basalis | D | ||||||||||
| Ptinomorphus granosus | D | ||||||||||
| Ptinus agnatus | D | D | |||||||||
| Ptinus fallax | D | ||||||||||
| Stegobium paniceum | adv | D | |||||||||
| Tricorynus | dup | D | D | D | D | D,L | |||||
| Tricorynus nubilus | L | ||||||||||
| Tricorynus obsoletus | L | ||||||||||
| Vrilletta blaisdelli | L | D | D | ||||||||
| Xarifa insularis | end | L | D,L | D | D | ||||||
| Xestobium marginicolle | L | D | |||||||||
| Xyletinus undet. sp. | D | ||||||||||
|
CLEROIDEA Byturidae |
|||||||||||
| Xerasia grisescens | D | D,L | D,L | D | |||||||
| Cleridae | |||||||||||
| Cymatodera angustata | L | ||||||||||
| Cymatodera caterinoi | end | L | D,L | D,L | |||||||
| Cymatodera insularis | end | L | D | D,L | |||||||
| Cymatodera ovipennis | L | ||||||||||
| Loedelia maculicollis | D | ||||||||||
| Necrobia ruficollis | adv | D | L | D | |||||||
| Necrobia rufipes | adv | D | D,L | D | D | D,L | D | D,L | |||
| Phyllobaenus | dup | D | |||||||||
| Phyllobaenus funebris | D | D | |||||||||
| Phyllobaenus scaber | D | D | |||||||||
| Trichodes ornatus | D | ||||||||||
| Melyridae | |||||||||||
| Attalus transmarinus | end | L | |||||||||
| Attalus undesc. sp. | end | D | |||||||||
| Charopus undesc. sp. | L | D | |||||||||
| Collops cribrosus | D | D | D,L | ||||||||
| Collops crusoe | end | D | D,L | D,L | D,L | ||||||
| Collops vittatus | D | ||||||||||
| Dasytastes | dup | D | D | D | D | D | |||||
| Dasytastes catalinae | end | D,L | |||||||||
| Dasytastes insularis | end | L | |||||||||
| Dasytes | dup | D | |||||||||
| Dasytes clementae | end | L | |||||||||
| Endeodes basalis | D | D | D | L | D | D | |||||
| Endeodes collaris | L | D,L | |||||||||
| Endeodes insularis | L | D | L | D | |||||||
| Eschatocrepis constrictus | D | D | L | D,L | D | ||||||
| Listrus | dup | D,L | D | L | D | D,L | |||||
| Listrus anacapaensis | end | L | |||||||||
| Listrus interruptus | D | ||||||||||
| Malachius undet. sp. | L | ||||||||||
| Microasydates punctipennis | end | D,L | |||||||||
| Microasydates sanclemente | end | D,L | |||||||||
| Microasydates santabarbara | D,L | D,L | D,L | ||||||||
| Microlipus laticeps | D | D | D | D | |||||||
| Pseudasydates explanatus | D | ||||||||||
| Trichochrous brevicornis | D | D | |||||||||
| Trichochrous calcaratus | end | D,L | D,L | D,L | D,L | ||||||
| Trichochrous pedalis | D,L | ||||||||||
| Trichochrous undesc. sp. 1 near brevicornis | end | D | D | D | D | ||||||
| Trichochrous undesc. sp. 2 near brevicornis | end | D | |||||||||
| Trichochrous undesc. sp. near pedalis | end | D,L | D | D,L | |||||||
| Trogossitidae | |||||||||||
| Temnoscheila chlorodia | D,L | ||||||||||
| Tenebroides crassicornis | L | ||||||||||
| Tenebroides occidentalis | D | ||||||||||
|
TENEBRIONOIDEA Anthicidae |
|||||||||||
| Amblyderus obesus | D | D,L | |||||||||
| Amblyderus parviceps | D | D | |||||||||
| Anthicus cribratus | D,L | ||||||||||
| Anthicus maritimus | D | ||||||||||
| Anthicus nanus | D | D | |||||||||
| Anthicus punctulatus | D | D | D | ||||||||
| Anthicus rufulus | D | ||||||||||
| Cyclodinus annectens | L | D,L | |||||||||
| Ischyropalpus nitidulus | D | L | D,L | D,L | |||||||
| Notoxus desertus | D,L | D | D | ||||||||
| Notoxus sparsus | L | ||||||||||
| Omonadus floralis | adv | L | |||||||||
| Ciidae | |||||||||||
| Ceracis californicus | D | ||||||||||
| Cis undet. sp. | L | ||||||||||
| Hadreule blaisdelli | D | ||||||||||
| Orthocis punctatus | D | D | |||||||||
| Sulcacis curtulus | D | ||||||||||
| Meloidae | |||||||||||
| Cordylospasta opaca | L | ||||||||||
| Epicauta puncticollis | D,L | ||||||||||
| Lytta stygica | D | ||||||||||
| Meloe barbarus | D,L | D,L | L | D,L | D,L | D | |||||
| Meloe strigulosus | D,L | ||||||||||
| Mordellidae | |||||||||||
| Mordella albosuturalis | D | ||||||||||
| Mordella hubbsi | D,L | ||||||||||
| Mordellina undet. sp. | D | D | D | D | D | D | D | ||||
| Mordellistena undet. sp. | D | D | D | D,L | D,L | D,L | D | ||||
| Mycetophagidae | |||||||||||
| Litargus balteatus | D | D | |||||||||
| Mycetophagus pluriguttatus | D | D | |||||||||
| Typhaea stercorea | adv | D | |||||||||
| Mycteridae | |||||||||||
| Lacconotus pinicola | D,L | D,L | |||||||||
| Oedemeridae | |||||||||||
| Copidita quadrimaculata | D | D | D | D | D | D | |||||
| Nacerdes melanura | adv | D | |||||||||
| Xanthochroa marina | D | ||||||||||
| Pyrochroidae | |||||||||||
| Pedilus bardii | D | ||||||||||
| Salpingidae | |||||||||||
| Rhinosimus undet. sp. | D | ||||||||||
| Scraptiidae | |||||||||||
| Anaspis atrata | D | ||||||||||
| Anaspis collaris | D,L | ||||||||||
| Pentaria trifasciata | D | D,L | D | ||||||||
| Tenebrionidae | |||||||||||
| Alaudes singularis | D,L | L | |||||||||
| Apocrypha anthicoides | D | D | |||||||||
| Apsena barbarae | L | L | |||||||||
| Apsena grossa | end | D,L | D,L | D,L | D,L | D,L | D,L | ||||
| Apsena pubescens | D | D,L | D | D | |||||||
| Apsena rufipes | D | D | |||||||||
| Batuliodes rotundicollis | D | ||||||||||
| Blapstinus angustus | D | ||||||||||
| Blapstinus brevicollis | D | D,L | D,L | ||||||||
| Blapstinus discolor | D | L | D | D | |||||||
| Cibdelis bachei | end | L | L | D,L | D,L | D | |||||
| Coelocnemis magna | D,L | ||||||||||
| Coelus ciliatus | L | D | D | D | |||||||
| Coelus globosus | D,L | D,L | D,L | L | L | D,L | D,L | ||||
| Coelus pacificus | end | L | D,L | D,L | D,L | D,L | D,L | D,L | D,L | ||
| Conibius seriatus | D | D | D | ||||||||
| Coniontis elliptica | L | L | |||||||||
| Coniontis lamentabilis | L | ||||||||||
| Coniontis lata | end | D,L | D,L | D,L | D,L | D,L | D,L | L | |||
| Coniontis microsticta | D | ||||||||||
| Coniontis nemoralis | D | ||||||||||
| Coniontis santarosae | end | D,L | D,L | D,L | |||||||
| Coniontis subpubescens | L | L | |||||||||
| Coniontis viatica | L | ||||||||||
| Corticeus opaculus | D,L | ||||||||||
| Cryptadius inflatus | D,L | D | |||||||||
| Eleodes acuticauda | D,L | D,L | D,L | D,L | D,L | D | D,L | D | |||
| Eleodes carbonaria | D | D,L | D | ||||||||
| Eleodes clavicornis | D | ||||||||||
| Eleodes dentipes | L | L | L | L | L | ||||||
| Eleodes gigantea | D | D | |||||||||
| Eleodes inculta | end | D,L | D,L | L | D | D,L | D,L | ||||
| Eleodes littoralis | D | D | D | D,L | D,L | D,L | |||||
| Eleodes nigropilosa | D,L | D | D | ||||||||
| Eleodes osculans | D | D | D,L | D,L | |||||||
| Eleodes scabripennis | L | L | |||||||||
| Eleodes subvestita | end | D,L | |||||||||
| Epantius obscurus | D,L | D | D | D,L | D,L | D,L | D,L | ||||
| Eusattus difficilis | L | ||||||||||
| Eusattus politus | end | D,L | L | D,L | |||||||
| Eusattus robustus | end | D,L | D | D,L | D | D | |||||
| Helops bachei | ?end | D | D,L | D | D | D,L | D | D | D | ||
| Helops blaisdelli | L | ||||||||||
| Helops rugicollis | D | ||||||||||
| Hylocrinus longulus | D | ||||||||||
| Hymenorus | dup | L | |||||||||
| Hymenorus infuscatus | L | ||||||||||
| Isomira | dup | D | |||||||||
| Isomira comstocki | D | D,L | |||||||||
| Isomira damnata | D | ||||||||||
| Isomira luscitiosa | D | D | |||||||||
| Isomira variabilis | L | ||||||||||
| Lepidocnemeplatia sericea | D | ||||||||||
| Metoponium | dup | D | |||||||||
| Metoponium convexicolle | L | ||||||||||
| Metoponium insulare | end | D,L | |||||||||
| Mycetochara | dup | D | |||||||||
| Mycetochara pubipennis | L | ||||||||||
| Nyctoporis carinata | D | D,L | D,L | D,L | D,L | ||||||
| Phaleria rotundata | D,L | D | D,L | D,L | D,L | D | |||||
| Platydema oregonensis | D | ||||||||||
| Telabis serratus | D | ||||||||||
| Tonibius sulcatus | D,L | ||||||||||
| Tribolium castaneum | adv | D | |||||||||
| Ulus crassus | L | ||||||||||
| Zopheridae | |||||||||||
| Lasconotus linearis | D | D | |||||||||
| Megataphrus tenuicornis | D | ||||||||||
| Phloeodes diabolicus | L | ||||||||||
| Phloeodes plicatus | D,L | D,L | |||||||||
| Rhagodera costaefragmenta | end | L | |||||||||
| Rhagodera interrupta | D,L | ||||||||||
| Rhagodera tuberculata | L | L | D,L | ||||||||
| Synchita lecontei | D | ||||||||||
|
COCCINELLOIDEA Akalyptoischiidae |
|||||||||||
| Akalyptoischion heterotrichos | D | ||||||||||
| Akalyptoischion hormathos | D | L | D | D | D | ||||||
| Cerylonidae | |||||||||||
| Cerylon unicolor | D | ||||||||||
| Coccinellidae | |||||||||||
| Axion plagiatum | D | ||||||||||
| Carinodulinka undesc. sp. near baja | D | ||||||||||
| Cephaloscymnus occidentalis | L | ||||||||||
| Chilocorus undet. sp. | L | D | |||||||||
| Coccidophilus atronitens | D | D | |||||||||
| Coccinella californica | D,L | D,L | D | D,L | D,L | D,L | D,L | D,L | |||
| Coccinella johnsoni | D,L | D,L | D,L | ||||||||
| Coccinella novemnotata | D | ||||||||||
| Coccinella septempunctata | adv | D | D | D | D | D | D | D | D | ||
| Cycloneda polita | L | D,L | D | ||||||||
| Cycloneda sanguinea | D | D,L | D | ||||||||
| Delphastus catalinae | L | D,L | D,L | ||||||||
| Diomus debilis | D | D | |||||||||
| Hippodamia convergens | D,L | D | D,L | D | D,L | D,L | D,L | D,L | |||
| Hippodamia quinquesignata | D | D,L | D | D,L | D,L | D,L | D,L | ||||
| Hyperaspidius | dup | D | |||||||||
| Hyperaspidius comparatus | L | ||||||||||
| Hyperaspis | dup | D | D | ||||||||
| Hyperaspis lateralis | D,L | D,L | |||||||||
| Hyperaspis sp. near annexa | D,L | ||||||||||
| Hyperaspis taeniata | D,L | ||||||||||
| Microweisea undet. sp. | D | ||||||||||
| Nephus binaevatus | adv | D,L | |||||||||
| Nephus guttulatus | D | D | L | D | |||||||
| Nephus sordidus | D | D | D | D | |||||||
| Nipus niger | D | ||||||||||
| Olla v-nigrum | D | D | |||||||||
| Paranaemia vittigera | L | ||||||||||
| Psyllobora renifer | D | ||||||||||
| Psyllobora vigintimaculata | D | D | D,L | D,L | D | ||||||
| Rhyzobius forestieri | adv | L | D,L | ||||||||
| Rhyzobius lophanthae | adv | L | D | D | D | ||||||
| Scymnus ardelio | L | L | |||||||||
| Scymnus cervicalis | L | D | D | ||||||||
| Scymnus coniferarum | D | ||||||||||
| Scymnus difficilis | D | D | |||||||||
| Scymnus falli | end | D | L | D,L | D,L | ||||||
| Scymnus fenderi | D | ||||||||||
| Scymnus jacobianus | D | D | D | ||||||||
| Scymnus loewii | D | D,L | |||||||||
| Scymnus marginicollis | D | D | D,L | D,L | D | ||||||
| Scymnus nebulosus | L | D,L | D | D | |||||||
| Scymnus pallens | L | D,L | D | ||||||||
| Stethorus punctum | D | D | |||||||||
| Zagloba ornata | D | L | D | ||||||||
| Corylophidae | |||||||||||
| Aenigmaticum californicum | D,L | D | D | D,L | |||||||
| Orthoperus undet. sp. | D | ||||||||||
| Sericoderus undet. sp. | D | D | D | ||||||||
| Endomychidae | |||||||||||
| Aphorista morosa | D | D | D,L | ||||||||
| Latridiidae | |||||||||||
| Cartodere australica | adv | D | |||||||||
| Corticaria undet. sp. | L | D | |||||||||
| Corticarina | dup | D | D | ||||||||
| Corticarina cavicollis | D | ||||||||||
| Corticarina herbivagans | L | D,L | |||||||||
| Corticarina milleri | end | D | L | L | L | D | L | ||||
| Corticarina minuta | D | D | |||||||||
| Dienerella undet. sp. | D,L | ||||||||||
| Enicmus aterrimus | D | D | |||||||||
| Fuchsina undesc. sp. | end | D | D | D,L | D | ||||||
| Melanophthalma | dup | D | D | ||||||||
| Melanophthalma americana | L | L | L | D | |||||||
| Melanophthalma casta | L | D,L | |||||||||
| Melanophthalma insularis | end | L | |||||||||
| Metophthalmus haigi | D | D | D,L | ||||||||
| Metophthalmus rudis | D | D | D,L | D | |||||||
| Metophthalmus trux | D | D | D,L | D | |||||||
| Revelieria californica | D | D | |||||||||
| Stephostethus armatulus | L | ||||||||||
| Stephostethus costicollis | D | D,L | |||||||||
| Stephostethus liratus | D | ||||||||||
|
EROTYLOIDEA Erotylidae |
|||||||||||
| Cryptophilus angustus | adv | D | |||||||||
| Dacne californica | D | D,L | D,L | D | |||||||
|
NITIDULOIDEA Kateretidae |
|||||||||||
| Amartus tinctus | D,L | L | D,L | ||||||||
| Heterhelus sericans | L | ||||||||||
| Monotomidae | |||||||||||
| Hesperobaenus abbreviatus | D | D,L | D | ||||||||
| Macreurops longicollis | D | ||||||||||
| Phyconomus marinus | D | D | |||||||||
| Nitidulidae | |||||||||||
| Brassicogethes aeneus | D | ||||||||||
| Carpophilus | dup | D | |||||||||
| Carpophilus discoideus | D | ||||||||||
| Carpophilus ligneus | D | D | D | ||||||||
| Cryptarcha gila | D | L | D | ||||||||
| Glischrochilus quadrisignatus | adv | D,L | |||||||||
| Glischrochilus sanguinolentus | adv | D,L | |||||||||
| Nitidula flavomaculata | adv | D | |||||||||
| Nitops pallipennis | D,L | D | D | D | D,L | D,L | D | ||||
| Thalycra undet. sp. | D | ||||||||||
|
CUCUJOIDEA Cryptophagidae |
|||||||||||
| Atomaria | dup | L | |||||||||
| Atomaria lewisi | adv | D | |||||||||
| Atomaria nubipennis | D | ||||||||||
| Atomaria puella | D | ||||||||||
| Cryptophagus tuberculosus | D,L | L | D | ||||||||
| Laemophloeidae | |||||||||||
| Narthecius striaticeps | D | ||||||||||
| Phalacridae | |||||||||||
| Phalacrus undet. sp. 1 | D | ||||||||||
| Phalacrus undet. sp. 2 | D,L | ||||||||||
| Silvanidae | |||||||||||
| Silvanoprus angusticollis | adv | D | |||||||||
|
CHRYSOMELOIDEA Cerambycidae |
|||||||||||
| Anastrangalia laetifica | D | ||||||||||
| Arhopalus asperatus | L | ||||||||||
| Arhopalus productus | L | ||||||||||
| Asemum nitidum | D | ||||||||||
| Brachysomida californica | D | ||||||||||
| Brothylus gemmulatus | D | ||||||||||
| Callidiellum rufipenne | adv | L | |||||||||
| Callimus ruficollis | D | D | |||||||||
| Centrodera autumnata | D | ||||||||||
| Centrodera spurca | D | ||||||||||
| Desmocerus californicus | D | D | |||||||||
| Enaphalodes hispicornis | D,L | ||||||||||
| Holopleura marginata | D | ||||||||||
| Ipochus fasciatus | D,L | L | D,L | D | D,L | D,L | D,L | D,L | |||
| Lophopogonius crinitus | D | D | |||||||||
| Megobrium edwardsi | D | L | |||||||||
| Nathrius brevipennis | adv | D | |||||||||
| Necydalis laevicollis | D | ||||||||||
| Oberea quadricallosa | D | ||||||||||
| Paranoplium gracile | D,L | ||||||||||
| Phoracantha recurva | adv | D | D | ||||||||
| Phoracantha semipunctata | adv | D | D | ||||||||
| Phymatodes decussatus | D | D,L | |||||||||
| Phymatodes grandis | D | D | |||||||||
| Prionus californicus | D | ||||||||||
| Saperda horni | D | ||||||||||
| Stenocorus vestitus | D | ||||||||||
| Sternidocinus barbarus | D,L | ||||||||||
| Strophiona tigrina | D | D | |||||||||
| Styloxus fulleri | D | ||||||||||
| Trichocnemis spiculatus | D | ||||||||||
| Xestoleptura crassipes | D | ||||||||||
| Xylotrechus insignis | D,L | ||||||||||
| Xylotrechus nauticus | D | D,L | |||||||||
| Chrysomelidae | |||||||||||
| Acanthoscelides margaretae | D | D | D | D | |||||||
| Acanthoscelides napensis | D | L | D | D | |||||||
| Acanthoscelides pauperculus | L | L | |||||||||
| Acanthoscelides pullus | D | D | D | D | L | D | D | ||||
| Altica undet. sp. | D | D | |||||||||
| Aulacothorax recticollis | D | D | |||||||||
| Calligrapha sigmoidea | D | ||||||||||
| Charidotella sexpunctata | D | D | |||||||||
| Colaspidea smaragdula | D,L | D,L | D | ||||||||
| Cryptocephalus sanguinicollis | D | ||||||||||
| Diabrotica undecimpunctata | D,L | L | D,L | D | |||||||
| Diachus auratus | D | D,L | D,L | D | D,L | D,L | D,L | ||||
| Dibolia californica | D | ||||||||||
| Disonycha latiovittata | D | D | |||||||||
| Epitrix similaris | L | ||||||||||
| Epitrix subcrinita | D,L | ||||||||||
| Erynephala morosa | L | ||||||||||
| Gastrophysa cyanea | D | D | D | ||||||||
| Lema daturaphila | D,L | D | |||||||||
| Longitarsus undet. sp. 1 | D | ||||||||||
| Longitarsus undet. sp. 2 | D | ||||||||||
| Megacerus impiger | D | D,L | |||||||||
| Monoxia undet. sp. | D | ||||||||||
| Pachybrachis melanostictus | D | ||||||||||
| Pachybrachis mobilis | D | ||||||||||
| Pachybrachis pluripunctatus | D | ||||||||||
| Pachybrachis punctatus | L | D | |||||||||
| Pachybrachis quadratus | L | ||||||||||
| Phaedon prasinellus | D,L | ||||||||||
| Phyllotreta | dup | D | |||||||||
| Phyllotreta pusilla | L | ||||||||||
| Plagiodera californica | D,L | ||||||||||
| Scelolyperus torquatus | L | ||||||||||
| Spintherophyta punctum | end | L | |||||||||
| Stator limbatus | D,L | ||||||||||
| Trachymela sloanei | adv | D | |||||||||
| Trirhabda confusa | D | ||||||||||
| Trirhabda sericotrachyla | D | D | |||||||||
| Yingabruxia sordida | L | D | D,L | ||||||||
|
CURCULIONOIDEA Attelabidae |
|||||||||||
| Deporaus glastinus | D | D,L | |||||||||
| Temnocerus aeratoides | D | ||||||||||
| Temnocerus aureus | L | D | |||||||||
| Temnocerus insularis | L | L | |||||||||
| Temnocerus naso | D | ||||||||||
| Brentidae | |||||||||||
| Apioninae | dup | D | D | D | |||||||
| Coelocephalapion antennatum | L | L | |||||||||
| Coelocephalapion californicum | L | ||||||||||
| Coelocephalapion oedorhynchum | L | ||||||||||
| Curculionidae | |||||||||||
| Anthonomus | dup | D | |||||||||
| Anthonomus inermis | L | D,L | L | ||||||||
| Anthonomus pauperculus | L | ||||||||||
| Anthonomus undesc. sp. | ?end | D | |||||||||
| Apleurus jacobinus | D,L | ||||||||||
| Carphobius undesc. sp. | end | D | |||||||||
| Carphoborus declivis | D | ||||||||||
| Ceutorhynchus assimilis | adv | D | |||||||||
| Coccotrypes dactyliperda | adv | L | |||||||||
| Curculio aurivestis | D,L | ||||||||||
| Curculio uniformis | L | D,L | |||||||||
| Dendrocranulus cucurbitae | D | D | D | D,L | D | ||||||
| Dendroctonus valens | D | ||||||||||
| Elassoptes marinus | D | D | D | D | D | ||||||
| Emphyastes fucicola | D | D | D | D | |||||||
| Geodercodes latipennis | D | D,L | D,L | ||||||||
| Gilbertiola undet. sp. | D,L | ||||||||||
| Gnathotrichus pilosus | D | ||||||||||
| Hypera postica | adv | D | D | D | D | ||||||
| Hypothenemus eruditus | D | D | D | ||||||||
| Ips paraconfusus | D,L | ||||||||||
| Listroderes | dup | D | |||||||||
| Listroderes costirostris | adv | D | D | D | D | ||||||
| Listronotus | dup | D | |||||||||
| Listronotus sordidus | L | ||||||||||
| Micromastus gracilis | D | ||||||||||
| Monarthrum scutellare | D | ||||||||||
| Naupactus cervinus | adv | D,L | D | ||||||||
| Nemocestes undet. sp. | D | D,L | D | ||||||||
| Notiodes aeratus | D | ||||||||||
| Otiorhynchus cribricollis | adv | D | |||||||||
| Peritelinus undet. sp. | D | ||||||||||
| Pityophthorus carmeli | D | D | |||||||||
| Procryphalus utahensis | D | ||||||||||
| Pselactus spadix | adv | D | |||||||||
| Pseudips mexicanus | L | ||||||||||
| Pseudopityophthorus agrifoliae | D | ||||||||||
| Pseudopityophthorus pubipennis | D | D | |||||||||
| Rhinocyllus conicus | adv | D,L | L | ||||||||
| Rhyncolus | dup | D | L | ||||||||
| Rhyncolus cylindricollis | D | ||||||||||
| Scaphomorphus americanus | L | ||||||||||
| Sciopithes insularis | end | L | |||||||||
| Sciopithes setosus | L | L | |||||||||
| Scyphophorus yuccae | D | ||||||||||
| Sibinia maculata | L | L | D,L | ||||||||
| Sitona californius | D | D,L | D | D,L | D | ||||||
| Smicronyx | dup | D | |||||||||
| Smicronyx cinereus | L | ||||||||||
| Sphenophorus graminis | D | ||||||||||
| Sphenophorus phoeniciensis | D | ||||||||||
| Sphenophorus simplex | D | D | |||||||||
| Sphenophorus vomerinus | L | ||||||||||
| Stenoclyptus sulcatus | D | ||||||||||
| Stenoptochus undet. sp. | D | ||||||||||
| Thalasselephas testaceus | D | D | D | ||||||||
| Trichobaris compacta | D | ||||||||||
| Trigonoscuta anacapensis | end | D,L | |||||||||
| Trigonoscuta catalina | end | D,L | |||||||||
| Trigonoscuta clemente | end | D,L | D | ||||||||
| Trigonoscuta curviscroba | end | L | |||||||||
| Trigonoscuta miguelensis | end | D,L | |||||||||
| Trigonoscuta nesiotis | end | D,L | |||||||||
| Trigonoscuta nicolana | end | D,L | D | ||||||||
| Trigonoscuta pilosa | L | L | |||||||||
| Trigonoscuta sanctabarbarae | end | D,L | |||||||||
| Trigonoscuta sanctarosae | end | D,L | |||||||||
| Trigonoscuta stantoni | end | D,L | |||||||||
| Tychius | dup | L | |||||||||
| Tychius lineellus | D,L | D,L | |||||||||
| Xyleborinus saxesenii | adv | D | D |
Note:
The sequence of suborders, superfamilies, and families corresponds to the sequence in the Annotated Checklist; the sequence of genera and species is alphabetical within families. Taxa are marked as “undet. sp.” for undetermined species that simply have not been identified to lower rank and “undesc. sp.” for undescribed species where the specimens have been specifically identified to an unnamed new taxon. “Duplicate genus record” (“dup”) means that the line is not counted as a unique taxon but contains additional island records for the given genus. “Endemic” (“end”) = nominal species that are purportedly restricted to the Channel Islands. “Adventive” (“adv”) marks species whose native ranges do not include southern California. In the columns representing each of the eight Channel Islands, “D” = digitized records anchoring the species-by-island presence, “L” = literature records.
Notable taxa
In this section we highlight some island-specific findings notable for their taxonomic or biogeographic implications, and highlight taxa in need of further investigation. For specific island records and other details, please refer to the taxon entries in the main checklist.
Among Carabidae, the subgenus Pterostichus (Hypherpes), a California-centric subgenus, is in serious need of investigation; currently there are six “known” species from the islands, including one putative endemic, but their taxonomy needs revision. Amara insularis is currently considered endemic and occurring on all islands, but based on investigation of hundreds of specimens housed in SBMNH (M. L. Gimmel, 2021, personal observation) this species is questionably distinct from mainland (and island) A. insignis.
Among non-endemic taxa, a few are notable for having flightless females, yet have clearly dispersed from source populations on the mainland: Anorus piceus (Dascillidae), Pterotus obscuripennis (Lampyridae), and Zarhipis integripennis (Phengodidae).
Apparently undescribed, and possibly endemic, island species exist in the genera Dalopius (Elateridae), Mordellistena (Mordellidae), Carinodulinka (Coccinellidae), Fuchsina (Latridiidae), Dacne (Erotylidae), Longitarsus (Chrysomelidae; at least two species), Anthonomus (Curculionidae), Gilbertiola (Curculionidae) (all M. L. Gimmel, 2021, personal observation), Phobetus (Scarabaeidae; at least two species; A. Evans, 2021, personal communication), and Carphobius (A. Cognato & S. Smith, 2022, personal communication), as well as the subfamily Leptotyphlinae (Staphylinidae). Interestingly, the Fuchsina, Gilbertiola, and Leptotyphlinae are eyeless and flightless (as is Pinodytes gibbosus; Leiodidae), while the two Longitarsus taxa are both flightless with abbreviated elytra; one undescribed Mordellistena has vestigial hind wings.
Taxa needing taxonomic investigation that will almost certainly reveal additional species, possibly including endemics, are as follows: Bacanius species (Histeridae), Plegaderus species (Histeridae), Anthaxia species (Buprestidae), Hyperaspis species (Coccinellidae), and Dienerella species (Latridiidae).
At least 63 known, unique taxa (not including known, undescribed species) still lack species determinations, predominantly in groups that lack modern taxonomic treatments, including all or most genera in the Scirtidae, Ptiliidae, Staphylinidae (Aleocharinae, Paederinae, Scydmaeninae), Salpingidae, Latridiidae, and Brentidae (Apioninae).
Island endemic taxa are distributed across many families of Coleoptera; the family with the highest proportion of endemics was found to be Melyridae, with 13 out of 28 species, or 46% of the island fauna. This family is the current research focus for MLG, and several of these island endemics are undescribed and will be receiving treatment in the near future; based on much recent fieldwork and museum work, the island endemism in this family is believed to be genuine rather than artefactual. Other families with significant proportions of endemics, either real or artefactual, include Scarabaeidae (eight out of 41 species, or 20%), Curculionidae (13 out of 65 species, or 20%), Cleridae (two out of 10 species, or 20%), Tenebrionidae (11 out of 61 species, or 18%), Latridiidae (three out of 19 species, or 16%), Zopheridae (one out of eight species, or 13%), and Ptinidae (two out of 25 species, or 8%). Notable genera with endemics include Coenonycha (Scarabaeidae), in which all four species from the islands are endemic; Serica (Scarabaeidae), which contains two endemic species; Xarifa (Ptinidae), a genus with one island endemic species and one rarely collected mainland species, neither of which have been studied since their original description, or illustrated; and Trigonoscuta (Curculionidae), with 10 putative endemic species that desperately need a taxonomic reassessment.
Missing taxa
The balance of available evidence suggests that the Channel Islands have never been connected to the mainland via a land bridge (Miller, 1985a). Consequently, the entire beetle fauna of the islands was acquired via over-water dispersal events or human-aided transport. As a result, the fauna is notably depauperate compared to that of the mainland (Miller, 1985a). While we do not provide a comprehensive faunal comparison to the mainland here, we hope that the following family-level assessment of missing taxa will aid in emphasizing this conclusion, or perhaps serve to encourage further sampling and scouring of collections to discover the existence on the islands of these “missing taxa”.
Not all families occurring in California are considered to be candidates for missing taxa from the Channel Islands. These include: Amphizoidae, Archeocrypticidae, Biphyllidae, Bothrideridae, Brachypsectridae, Cerophytidae, Derodontidae, Diphyllostomatidae, Eulichadidae, Hybosoridae, Ischaliidae, Mauroniscidae, Megalopodidae, Nosodendridae, Noteridae, Prostomidae, Ptilodactylidae, Smicripidae, Sphaeritidae, Stenotrachelidae, Teredidae, Thanerocleridae, and Trachypachidae. These all occur in distant regions of the state and/or are restricted to elevations or habitat types not present on the Channel Islands, and are not likely to occur there.
Families absent from the Channel Islands but present on nearby mainland
Cupedidae (Archostemata). This family of two genera and two species in California (M. L. Gimmel, 2022, unpublished data) contains a species, Prolixocupes lobiceps (LeConte, 1874), widely distributed across dry areas of southern California that may eventually be found on the Channel Islands.
Eucinetidae (Clamboidea). This family of two genera and three species in California (M. L. Gimmel, 2022, unpublished data) contains at least one widespread coastal species, Nycteus infumatus (LeConte, 1853), that may yet be discovered on the Channel Islands.
Schizopodidae (Buprestoidea). This family of three genera and seven species in California (Nelson et al., 2008) contains species of Dystaxia LeConte, 1866 and Glyptoscelimorpha Horn, 1893 present on the nearby mainland.
Byrrhidae (Byrrhoidea). This family of three subfamilies, eight genera, and 10 species in California (M. L. Gimmel, 2022, unpublished data), although primarily boreal and montane in distribution, contains a few taxa occurring along California’s Central Coast south into Santa Barbara and Ventura counties (SBMNH specimen data).
Psephenidae (Dryopoidea). This family of three subfamilies and as many genera and species in California (Shepard, 1993) contains a widely distributed and abundant coastal species, Eubrianax edwardsii (LeConte, 1874), that is conspicuously absent from the Channel Islands. Lack of suitable microhabitat may explain this absence.
Artematopodidae (Elateroidea). This family of two subfamilies, four genera, and six species in California (M. L. Gimmel, 2022, unpublished data) contains a species, Brevipogon confusus (Fall, 1901), widely distributed in the southern half of California (Lawrence, 2005).
Lycidae (Elateroidea). This family of two subfamilies, five genera, and seven species in California (M. L. Gimmel, 2022, unpublished data), contains at least a couple of species occurring in the Coast Ranges.
Omethidae (Elateroidea). This family of two subfamilies, five genera, and seven species in California (M. L. Gimmel, 2022, unpublished data), including at least one, Ginglymocladus luteicollis Van Dyke, 1918, occurring in coastal Santa Barbara County (SBMNH specimen data).
Georissidae (Hydrophiloidea). This family of one species in California, Georissus californicus LeConte, 1874 (Hansen, 1999), occurs at lower elevations within the Transverse Ranges (SBMNH specimen data).
Hydrochidae (Hydrophiloidea). This family of one genus, Hydrochus Leach, 1817, and four species in California (Hansen, 1999), has species that occur in the Coast Ranges (SBMNH specimen data).
Glaphyridae (Scarabaeoidea). This family of one genus, Lichnanthe Burmeister, 1844, and six species in California, contains species occurring in coastal portions of the state, including Santa Barbara, Ventura, and Los Angeles counties (Carlson, 1980).
Glaresidae (Scarabaeoidea). This family of one genus, Glaresis Erichson, 1848, and 13 species in California (Gordon & Hanley, 2014), has species that occur within the Transverse Ranges of California (SBMNH specimen data).
Lucanidae (Scarabaeoidea). This family consists of two subfamilies, four genera, and 18 species in California (M. L. Gimmel, 2022, unpublished data). Species of Platycerus Geoffroy, 1792 and Sinodendron Hellwig, 1894 occur in southern coastal California (SBMNH specimen data).
Ochodaeidae (Scarabaeoidea). This family of two subfamilies, four genera, and five species in California (Paulsen, 2007) has at least one species, Parochodaeus californicus (Horn, 1895), occurring in coastal southern California (SBMNH specimen data).
Pleocomidae (Scarabaeoidea). This family contains one genus, Pleocoma LeConte, 1856, and about 23 species in California (M. L. Gimmel, 2022, unpublished data), many of them occurring in coastal California.
Agyrtidae (Staphylinoidea). This family of three subfamilies, four genera, and seven species in California (Newton, 1997) contains species, notably Necrophilus hydrophiloides Guérin-Méneville, 1835, occurring in nearby coastal California (SBMNH specimen data).
Lophocateridae (Cleroidea). This family of three genera and five species in California (M. L. Gimmel, 2022, unpublished data) contains species of the genus Eronyxa Reitter, 1876 occurring in nearby coastal California (Barron, 1971; SBMNH specimen data).
Peltidae (Cleroidea). This family of one genus, Peltis Müller, 1764, and three species in California (Barron, 1971, as Ostoma Laicharting, 1781) contains species occurring at lower elevations within the Transverse Ranges (SBMNH specimen data).
Rhadalidae (Cleroidea). This family of two genera and three species in California (M. L. Gimmel, 2022, unpublished data) contains species of both Rhadalus LeConte, 1852 and Semijlulistus Schilsky, 1894 occurring at lower elevations within the Transverse Ranges (SBMNH specimen data).
Aderidae (Tenebrionoidea). This family of four genera and five species in California (M. L. Gimmel, 2022, unpublished data) contains species occurring in nearby coastal California (SBMNH specimen data).
Melandryidae (Tenebrionoidea). This family of two subfamilies, 12 genera, and 15 species in California (M. L. Gimmel, 2022, unpublished data) contains several genera and species in coastal southern regions of California (SBMNH specimen data). Osphya lutea (Horn, 1879) is a particularly abundant and well-collected species we expect might occur on the islands.
Pythidae (Tenebrionoidea). This family of three genera and three species in California (M. L. Gimmel, 2022, unpublished data) contains at least one species, Sphalma quadricollis Horn, 1888, occurring at lower elevations in the Transverse Ranges (SBMNH specimen data).
Ripiphoridae (Tenebrionoidea). This family of two genera and 20 species in California (Linsley & MacSwain, 1951; M. L. Gimmel, 2022, unpublished data) contains species occurring, but rarely collected, in coastal California (Linsley & MacSwain, 1951; SBMNH specimen data).
Tetratomidae (Tenebrionoidea). This family of four subfamilies, seven genera, and nine species in California (M. L. Gimmel, 2022, unpublished data) contains species occurring at lower elevations in the Transverse Ranges (SBMNH specimen data).
Anamorphidae (Coccinelloidea). This family contains a single, introduced species, Symbiotes gibberosus (Lucas, 1846), occurring in California (Shockley, Tomaszewska & McHugh, 2009), including at lower elevations across the southern coastal portion of the state (SBMNH specimen data).
Murmidiidae (Coccinelloidea). This family contains a single, introduced species in California, Murmidius ovalis (Beck, 1817), which is a cosmopolitan stored product associate (Lawrence & Stephan, 1975).
Mycetaeidae (Coccinelloidea). This family contains a single, introduced species in California, Mycetaea subterranea (Fabricius, 1801), which is a cosmopolitan species (Shockley, Tomaszewska & McHugh, 2009).
Sphindidae (Nitiduloidea). This family contains two subfamilies, two genera, and three species in California (M. L. Gimmel, 2022, unpublished data), at least one of which, Sphindus crassulus Casey, 1898, occurs in nearby coastal California (SBMNH specimen data).
Cucujidae (Cucujoidea). This family contains two genera and six species in California (M. L. Gimmel, 2022, unpublished data) and is widely distributed in forested areas of California (SBMNH specimen data).
Orsodacnidae (Chrysomeloidea). This family contains a single species in California, Orsodacne atra (Ahrens, 1810), which occurs on the nearby coastal mainland (SBMNH specimen data).
Anthribidae (Curculionoidea). This family of two subfamilies, five genera, and 11 species in California (M. L. Gimmel, 2022, unpublished data) contains at least two species occurring on the nearby coastal mainland (SBMNH specimen data).
Cimberididae (Curculionoidea). This family of four genera and 10 species in California (Kuschel, 1989, as Nemonychidae) contains species occurring at lower elevations in the Transverse Ranges (SBMNH specimen data). Their life histories are closely tied with Pinus species (Pinaceae); consequently, they should be searched for during spring in the pine groves occurring on the Channel Islands.
Annotated checklist
The format of this annotated checklist is structured to provide a foundation for future research on the taxa included. Higher classification is arranged phylogenetically by suborder and superfamily, adopting the higher groupings of Cai et al. (2022). Taxa of family rank and lower are arranged alphabetically within higher taxa. Notes on the taxonomy and diversity within California are given for all higher taxa. The standardized sections for each taxon are briefly defined below. We did not attempt to provide an overview of the biology of the taxa in the checklist; this information can be gleaned from general works on beetle biology and as well as taxon-specific references cited herein.
In order to keep close accounting of the actual number of unique taxa known from the islands, the following system of presentation is employed: (1) Most family-level digital and literature records are excluded, except for select groups presenting taxonomic challenges (e.g., Apioninae), and for certain notable higher-taxon literature records, which are included in the respective Notes field; (2) Island records of supraspecific taxa that have no records at lower levels (e.g., a genus record with no identified species reported from the islands) are treated with a unique header as “(Taxon) undetermined species” to identify it as a unique taxon within the checklist; (3) When a genus-level taxon does have lower-level representation in our list, then any records determined only to the higher level are included merely under that higher-level heading and not listed as an additional taxon.
Nomenclatural Authority: This field contains a reference, or set of references, from which the valid name and taxon authorship were derived for this list. This is typically a recent revision or catalog and is intended to validate the use of the name here and serve as an anchor and starting point for future use of this list as taxonomic names and concepts continue to change (see Johnston, Aalbu & Franz, 2018).
Literature Records: In this field, every published island record for the taxon is cited with a page number. In the Notes field under the taxon we provide additional comments on such records, including previous nomenclatural combinations used in cited works, discounted or ambiguous references, etc.
Digitized Records: In this field, all digitized specimen records included in our final dataset are tallied by island for each taxon, and are listed by collection. For detailed information on each specimen record see the recordset discussed and referenced above.
Range: This field denotes whether the taxon is known only from the Channel Islands or also from the mainland. References are given to support the claim of endemicity (only known from the Channel Islands) or presence on the mainland. Full ranges of species are not included or covered in this checklist.
Notes: This field contains a wide variety of information about the taxon. For entries within supraspecific taxa, we include information on diversity in California, and discuss recent taxonomic revisions or catalogs. For species-level entries, we include any information about subspecies classification of island specimens. Any additional information deemed relevant, including island-specific natural history notes, as well as decisions on taxon validity and discrepancies or issues relating to literature or digitized specimens, are included under this section.
ADEPHAGA
Carabidae
Notes. There are 103 genera and 647 species of this family known from California, placed in 38 tribes (Bousquet, 2012; M. L. Gimmel, 2022, unpublished data). A subfamily classification is not widely agreed upon in the literature; we use only tribes below, following the arrangement of Bousquet (2012), with the exclusion of Cicindelidae, which has been recently recognized at the family level (e.g., Cai et al., 2022). Bousquet (2012) provided an extensively annotated distributional catalog and bibliography for the North American taxa.
Bembidiini
Notes. Thirteen genera and 148 species of Bembidiini are known to occur in California (Bousquet, 2012; M. L. Gimmel, 2022, unpublished data).
Bembidion Latreille, 1802
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): San Clemente (1 EMEC; 2 LACM; 8 SBMNH), San Miguel (8 LACM; 14 SBMNH), San Nicolas (4 LACM; 28 SBMNH), Santa Catalina (1 LACM; 23 SBMNH), Santa Cruz (18 LACM; 105 SBMNH), Santa Rosa (9 LACM; 18 SBMNH).
Notes. This genus was widely known in the earlier literature as Bembidium. Fully 115 species have been reported to occur in California (M. L. Gimmel, 2022, unpublished data). No complete keys exist for North American or Californian species, but Lindroth’s (1963) key included about 75% of the North American fauna (Bousquet, 2012). A record of Bembidion (Hirmoplataphus) recticolle LeConte, 1863 from Santa Cruz Island provided to the California Beetle Database was deemed unverifiable and needs to be substantiated.
Bembidion (Furcacampa) versicolor (LeConte, 1847)
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Cockerell, 1940: 284)
Digitized Records: none
Range: Also known from mainland (Bousquet, 2012).
Bembidion (Lymneops) laticeps (LeConte, 1858)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: San Clemente (1 SBMNH)
Range: Also known from mainland (Bousquet, 2012). This species was transferred from the subgenus Lymnaeum Stephens, 1828 to the subgenus Lymneops Casey, 1918 by Maddison & Maruyama (2019).
Bembidion (Lymneops) palosverdes Kavanaugh & Erwin, 1992
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Caterino, Caterino & Maddison, 2015: 410; Maddison & Maruyama, 2019: 46)
Digitized Records: Santa Catalina (2 SBMNH)
Range: Also known from mainland, but possibly extinct there (Caterino, Caterino & Maddison, 2015).
Notes. This species was thought to be extinct since its original description from the Palos Verdes Peninsula in greater Los Angeles, but was rediscovered by M. and K. Caterino on Santa Catalina in 2010 (Caterino, Caterino & Maddison, 2015). This species was transferred from the subgenus Cillenus Samouelle, 1819 to the subgenus Lymneops Casey, 1918 by Maddison & Maruyama (2019).
Bembidion (Notaphus) indistinctum Dejean, 1831
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Nicolas (Cockerell, 1940: 285), Santa Rosa (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 613).
Bembidion (Notaphus) insulatum (LeConte, 1852)
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Cockerell, 1940: 284)
Digitized Records: none
Range: Also known from mainland (Bousquet, 2012).
Bembidion (Peryphanes) platynoides Hayward, 1897
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Rosa (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 579).
Bembidion (Peryphodes) ephippigerum (LeConte, 1852)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: San Nicolas (9 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 629).
Bembidion (Peryphus) corgenoma Maddison, 2020
Nomenclatural Authority: Maddison (2020)
Literature Records: Santa Cruz (LeConte, 1876: 298; Fall, 1897: 236; Fall & Davis, 1934: 143), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Miguel (1 SBMNH), Santa Cruz (19 SBMNH), Santa Rosa (10 SBMNH)
Range: Also known from mainland (Maddison, 2020).
Notes. LeConte (1876) recorded this species as Bembidium mannerheimii (LeConte, 1852), subsequently considered a junior synonym of Bembidion transversale Dejean, 1831 (see Bousquet, 2012). Fall (1897) and Fall & Davis (1934) reported this species as B. transversale. Maddison (2020), however, concluded based on morphology that the Pacific coast species in the transversale species group represented a new species and provided a revised version of the relevant portion of the key in Lindroth (1963).
Bembidion (Peryphus) striola (LeConte, 1852)
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Cockerell, 1940: 285), Santa Catalina (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 564).
Bembidion (Trechonepha) iridescens (LeConte, 1852)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 654).
Elaphropus Motschulsky, 1839
Nomenclatural Authority: Bousquet (2012)
Notes. Seven species of this genus have been reported from California (Bousquet, 2012). The native North American members of Elaphropus are in need of revision (Bousquet, 2012).
Elaphropus undetermined species
Literature Records: none
Digitized Records: Santa Cruz (10 SBMNH)
Phrypeus Casey, 1924
Nomenclatural Authority: Bousquet (2012)
Notes. Only one species of Phrypeus occurs in North America (Bousquet, 2012).
Phrypeus rickseckeri (Hayward, 1897)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (6 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Tachys Dejean, 1821
Nomenclatural Authority: Bousquet (2012)
Notes. Six species of Tachys have been reported from California (Bousquet, 2012). The subgenus Paratachys Casey, 1918, with two species in California, has frequently been treated as a separate genus (Bousquet, 2012). Both this and the subgenus Tachys need revision (Bousquet, 2012).
Tachys (Paratachys) vorax LeConte, 1852
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Tachys (Tachys) corax LeConte, 1852
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Fall, 1897: 236)
Digitized Records: San Nicolas (5 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Notes. Fall (1897: 239) indicated that the “two specimens from San Clemente are closely allied to corax, Lec., but seem distinct by the obviously less transverse thorax.” This is likely identical to what we have identified as T. corax, and we have included these records together above.
Tachys (Tachys) vittiger LeConte, 1852
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236; Fall, 1901: 43; Bousquet, 2012: 689)
Digitized Records: none
Range: Also known from mainland (Fall, 1901; Bousquet, 2012).
Brachinini
Notes. One genus and 12 species of Brachinini occur in California (Bousquet, 2012).
Brachinus Weber, 1801
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Catalina (1 CASC), Santa Cruz (50 CASC; 31 EMEC; 50 SBMNH; 11 UCSB; 35 UASM), Santa Rosa (5 SBMNH)
Notes. There are 12 species of Brachinus reported from California (Bousquet, 2012), which were treated by Erwin (1965, 1970). This genus appeared in some early literature as Brachynus.
Brachinus costipennis Motschulsky, 1859
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236), Santa Cruz (Erwin, 1965: 6; Erwin, 1970: 88)
Digitized Records: Santa Cruz (1 BYUC; 7 CASC; 4 LACM; 4 SBMNH; 1 iNat), Santa Rosa (15 SBMNH)
Range: Also known from mainland (Erwin, 1965, 1970).
Notes. This species was reported as Brachinus carinulatus Motschulsky, 1859 by Fall (1897), which was synonymized with B. costipennis by Erwin (1965).
Brachinus gebhardis Erwin, 1965
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Cruz (Erwin, 1965: 7; Erwin, 1970: 132)
Digitized Records: Santa Catalina (6 LACM), Santa Cruz (1 CASC; 23 LACM; 9 SBMNH), Santa Rosa (10 SBMNH)
Range: Also known from mainland (Erwin, 1965, 1970).
Brachinus mexicanus Dejean, 1831
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Cruz (Erwin, 1965: 11; Erwin, 1970: 107)
Digitized Records: Santa Cruz (40 CASC)
Range: Also known from mainland (Erwin, 1965, 1970).
Notes. This species was recorded as Brachinus fidelis LeConte, 1863 by Erwin (1965), which was synonymized with B. mexicanus by Erwin (1970).
Brachinus quadripennis Dejean, 1825
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Cruz (Fall & Davis, 1934: 144)
Digitized Records: none
Range: Also known from mainland (Erwin, 1970).
Notes. Fall & Davis (1934) recorded this species as Brachynus tschernikhii Mannerheim, 1843, which was synonymized with B. quadripennis by Erwin (1970: 99).
Carabini
Notes. Two genera and 28 species of Carabini are known to occur in California (Bousquet, 2012).
Calosoma Weber, 1801
Nomenclatural Authority: Bousquet (2012)
Notes. The North American species of Calosoma were revised by Gidaspow (1959). Twenty-six species have been reported as occurring in California (Bousquet, 2012).
Calosoma (Camegonia) parvicolle Fall, 1910
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Gidaspow, 1959: 256)
Digitized Records: none
Range: Also known from mainland (Gidaspow, 1959; Bousquet, 2012).
Calosoma (Carabosoma) eremicola Fall, 1910
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Fall, 1910: 91; Cockerell, 1940: 284; Jeannel, 1940: 206; Gidaspow, 1959: 259; Bousquet, 2012: 235), Santa Catalina (Cockerell, 1940: 284; Gidaspow, 1959: 259)
Digitized Records: San Clemente (1 CASC; 18 LACM; 5 SBMNH), Santa Catalina (2 CASC; 1 SBMNH)
Range: Also known from mainland (Cockerell, 1940; Jeannel, 1940; Gidaspow, 1959; Bousquet, 2012).
Notes. This species was synonymized with Calosoma glabratum sponsum Casey, 1897 by Breuning (1928: 103), then re-validated by Jeannel (1940: 206). Calosoma eremicola was originally described as endemic to San Clemente Island by Fall (1910).
Calosoma (Chrysostigma) semilaeve LeConte, 1852
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Nicolas (Miller & Miller, 1985: 123), Santa Barbara (Miller & Miller, 1985: 123), Santa Catalina (Miller & Miller, 1985: 123), Santa Rosa (Fall, 1897: 236; Miller & Miller, 1985: 123)
Digitized Records: Santa Catalina (5 LACM), Santa Cruz (2 SBMNH; 1 UCSB)
Range: Also known from mainland (Gidaspow, 1959; Bousquet, 2012).
Notes. Recorded from the “Channel Islands” by Bousquet (2012: 243).
Chlaeniini
Notes. One genus and 13 species of Chlaeniini are known from California (Bousquet, 2012).
Chlaenius Brullé, 1834
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Cruz (4 UCSB; 13 UASM)
Notes. Thirteen species of Chlaenius have been reported to occur in California (Bousquet, 2012). The genus was revised for North America by Bell (1960).
Chlaenius (Chlaeniellus) obsoletus LeConte, 1851
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236; Bell, 1960: 150)
Digitized Records: Santa Cruz (3 CASC; 1 SBMNH)
Range: Also known from mainland (Bell, 1960; Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 984).
Chlaenius (Chlaeniellus) tricolor Dejean, 1826
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Bell, 1960; Bousquet, 2012).
Notes. All C. tricolor from California belong to the subspecies C. t. vigilans Say, 1830 (Bell, 1960; Bousquet, 2012).
Chlaenius (Chlaeniellus) variabilipes Eschscholtz, 1833
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Cruz (Bell, 1960: 150)
Digitized Records: Santa Catalina (1 LACM), Santa Cruz (4 LACM; 2 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Bell, 1960; Bousquet, 2012).
Chlaenius (Chlaenius) cumatilis LeConte, 1851
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Cruz (Fall & Davis, 1934: 144)
Digitized Records: Santa Cruz (52 CASC; 11 LACM; 19 SBMNH; 7 TAMU), Santa Rosa (6 SBMNH)
Range: Also known from mainland (Bell, 1960; Bousquet, 2012).
Clivinini
Notes. Three genera and 11 species of Clivinini are known to occur in California (Bousquet, 2012).
Schizogenius Putzeys, 1846
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Cruz (5 CASC; 14 SBMNH)
Notes. Seven species of Schizogenius have been recorded from California (Bousquet, 2012), belonging to two subgenera, Genioschizus Whitehead, 1972 and Schizogenius (s.str.). Whitehead (1972) revised the North American species.
Schizogenius (Schizogenius) depressus LeConte, 1852
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Rosa (Fall, 1897: 236; Whitehead, 1972: 294)
Digitized Records: Santa Cruz (2 CASC)
Range: Also known from mainland (Whitehead, 1972; Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 419).
Cychrini
Notes. One genus and 18 species of Cychrini are known to occur in California (Bousquet, 2012).
Scaphinotus Dejean, 1826
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Catalina (3 EMEC), Santa Cruz (2 EMEC; 1 UASM), Santa Rosa (1 EMEC)
Notes. Eighteen species of Scaphinotus are known from California (Bousquet, 2012). Most of these species belong to the subgenus Brennus Motschulsky, 1866, which was revised by Gidaspow (1968).
Scaphinotus (Brennus) crenatus (Motschulsky, 1859)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: San Miguel (1 CASC; 1 SBMNH), Santa Catalina (1 CASC), Santa Cruz (8 CASC; 1 LACM; 4 SBMNH), Santa Rosa (1 CASC; 9 SBMNH)
Range: Also known from mainland (Gidaspow, 1968).
Scaphinotus (Brennus) punctatus (LeConte, 1859)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236; Baker, 1905: 59; Gidaspow, 1968: 167; Bousquet, 2012: 223)
Digitized Records: Santa Catalina (5 CASC)
Range: Also known from mainland (Gidaspow, 1968).
Notes. This species was previously recorded as Cychrus mimus Horn, 1874 by Fall (1897) and Baker (1905: “Cuchrus mimus”), which was later synonymized with S. punctatus (see Gidaspow, 1968).
Scaphinotus (Brennus) ventricosus (Dejean, 1831)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Gidaspow, 1968: 171)
Digitized Records: Santa Catalina (1 CASC)
Range: Also known from mainland (Gidaspow, 1968).
Dyschiriini
Notes. Two genera and 21 species of Dyschiriini are known to occur in California (Bousquet, 2012).
Akephorus LeConte, 1852
Nomenclatural Authority: Bousquet (2012)
Notes. Two species of this genus occur in California (Bousquet, 2012). They were keyed (as species of Dyschirius) by Bousquet (1988).
Akephorus marinus LeConte, 1852
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Rosa (Fall, 1897: 236)
Digitized Records: San Miguel (33 LACM; 5 SBMNH; 18 CASC), San Nicolas (1 LACM; 5 SBMNH), Santa Cruz (34 LACM; 18 SBMNH; 8 CASC), Santa Rosa (7 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 431).
Dyschirius Bonelli, 1810
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Cruz (12 CASC)
Notes. Nineteen species of Dyschirius have been recorded from California (Bousquet, 2012). A key to most North American species of this genus was provided by Bousquet (1988).
Dyschirius aratus LeConte, 1852
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Rosa (7 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Dyschirius consobrinus LeConte, 1852
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Rosa (4 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Dyschirius gibbipennis LeConte, 1857
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Rosa (Fall, 1897: 236)
Digitized Records: San Miguel (1 CASC), Santa Cruz (2 SBMNH), Santa Rosa (6 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 439).
Dyschirius varidens Fall, 1910
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Harpalini
Notes. Thirteen genera and 78 species of Harpalini are known to occur in California (Bousquet, 2012; M. L. Gimmel, 2022, unpublished data).
Anisodactylus Dejean, 1829
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): San Miguel (1 LACM), Santa Cruz (19 UASM)
Notes. Thirteen species of Anisodactylus are known to occur in California (Bousquet, 2012). Noonan (1996) revised the subgenus Anisodactylus (Anisodactylus), the only subgenus known to occur in the Channel Islands.
Anisodactylus (Anisodactylus) californicus Dejean, 1829
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Miguel (Noonan, 1996: 126), Santa Catalina (Fall, 1897: 236), Santa Cruz (Noonan, 1996: 126), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Clemente (7 SBMNH), San Miguel (1 CASC; 2 SBMNH), San Nicolas (8 SBMNH), Santa Catalina (1 SBMNH), Santa Cruz (3 CASC), Santa Rosa (5 SBMNH)
Range: Also known from mainland (Noonan, 1996; Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 1022).
Anisodactylus (Anisodactylus) consobrinus LeConte, 1851
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236), Santa Cruz (LeConte, 1876: 298; Fall, 1897: 236; Fall & Davis, 1934: 144; Noonan, 1996: 107), Santa Rosa (Fall, 1897: 236)
Digitized Records: Santa Cruz (3 SBMNH), Santa Rosa (8 SBMNH)
Range: Also known from mainland (Noonan, 1996; Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 1021).
Anisodactylus (Anisodactylus) similis LeConte, 1851
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Cruz (Fall & Davis, 1934: 144)
Digitized Records: Santa Cruz (2 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Noonan, 1996; Bousquet, 2012).
Notes. This species was recorded as Anisodactylus semipunctatus LeConte, 1859, a current junior synonym of A. similis, by Fall & Davis (1934). Reported from the “Channel Islands” by Bousquet (2012: 1023).
Bradycellus Erichson, 1837
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): San Clemente (4 SBMNH), San Miguel (3 CASC), San Nicolas (3 LACM), Santa Cruz (5 CASC; 2 EMEC; 1 SBMNH; 3 UASM)
Notes. Four subgenera and 25 species of Bradycellus are known to occur in California (Bousquet, 2012). The two most species-rich subgenera in California, Liocellus Motschulsky, 1864 and Stenocellus Casey, 1914, both need revision (Bousquet, 2012).
Bradycellus (Liocellus) nitidus (Dejean, 1829)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236), Santa Cruz (Fall & Davis, 1934: 144), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Miguel (2 CASC; 2 SBMNH), San Nicolas (15 LACM; 11 SBMNH), Santa Catalina (1 SBMNH), Santa Cruz (4 CASC; 10 SBMNH; 1 UCSB), Santa Rosa (1 LACM; 20 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Notes. Fall (1897) recorded this species as Tachycellus nitidus; Fall & Davis (1934) recorded this species as Glycerius nitidus. Reported from the “Channel Islands” by Bousquet (2012: 1061).
Bradycellus (Stenocellus) californicus (LeConte, 1857)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Rosa (Fall, 1897: 236)
Digitized Records: San Clemente (16 SBMNH), San Miguel (1 SBMNH), San Nicolas (17 SBMNH), Santa Catalina (4 SBMNH), Santa Cruz (8 SBMNH), Santa Rosa (12 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 1068).
Bradycellus (Stenocellus) rupestris (Say, 1823)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: San Clemente (6 SBMNH), San Nicolas (12 SBMNH), Santa Cruz (3 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Bradycellus (Stenocellus) sejunctus (Casey, 1914)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: San Clemente (14 EMEC)
Range: Also known from mainland (Bousquet, 2012).
Dicheirus Mannerheim, 1843
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Cruz (3 CASC)
Notes. Five species of Dicheirus are known to occur in California (Bousquet, 2012). They were revised and keyed by Noonan (1968).
Dicheirus dilatatus (Dejean, 1829)
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Casey, 1914: 201; Noonan, 1968: 298; Noonan, 1975: 7 [map]), Santa Catalina (Noonan, 1968: 298), Santa Cruz (Fall & Davis, 1934: 144), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Clemente (29 LACM; 7 SBMNH), San Miguel (2 SBMNH), Santa Catalina (10 LACM; 5 SBMNH), Santa Cruz (2 SBMNH), Santa Rosa (10 SBMNH)
Range: Also known from mainland (Noonan, 1975; Bousquet, 2012).
Notes. This species was recorded as Anisodactylus dilatatus by Fall (1897), and as Dicheirus australinus Casey, 1914 by Casey (1914). The latter was synonymized with D. dilatatus by Noonan (1968). All island records of D. dilatatus refer to the nominate subspecies, D. d. dilatatus (Dejean, 1829) (Noonan, 1968). Reported from the “Channel Islands” by Bousquet (2012: 1042).
Dicheirus piceus (Ménétriés, 1843)
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Noonan, 1968: 290; Noonan, 1975: 7 [map]), Santa Catalina (Noonan, 1968: 290; Noonan, 1975: 7 [map]), Santa Rosa (Fall, 1897: 236)
Digitized Records: Anacapa (3 SBMNH), San Clemente (11 CASC; 5 LACM; 3 SBMNH), Santa Catalina (26 LACM; 6 SBMNH), Santa Cruz (4 CASC; 18 SBMNH), Santa Rosa (9 SBMNH)
Range: Also known from mainland (Noonan, 1975; Bousquet, 2012).
Notes. This species was recorded as Anisodactylus piceus by Fall (1897). Reported from the “Channel Islands” by Bousquet (2012: 1043).
Harpalus Latreille, 1802
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Anacapa (1 ASUHIC)
Notes. Thirteen species of Harpalus have been reported from California (Bousquet, 2012; M. L. Gimmel, 2022, unpublished data). Most species of the genus were treated for North America by Noonan (1991), with treatments of additional subgenera by Ball & Anderson (1962) and Will (1997).
Harpalus (Megapangus) caliginosus (Fabricius, 1775)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Will, 1997).
Notes. Will (1997) provided a taxonomic review of the subgenus Harpalus (Megapangus), including a shaded range map showing H. caliginosus ranging partially into the Channel Islands (Will, 1997: 46); however, no Channel Island records were given.
Harpalus (Pseudoophonus) pensylvanicus (DeGeer, 1774)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (1 UCSB)
Range: Also known from mainland (Ball & Anderson, 1962; Bousquet, 2012).
Notes. Ball & Anderson (1962) provided a key to the species of the subgenus Pseudoophonus Motschulsky, 1844 (as Pseudophonus). Only one species of the subgenus has been recorded from California (Bousquet, 2012).
Stenolophus Dejean, 1821
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Cruz (2 UASM)
Notes. Eleven species of Stenolophus occur in California, four in subgenus Agonoderus Dejean, 1829 and seven in subgenus Stenolophus (Bousquet, 2012). Lindroth’s (1968) key treated all members of the latter, but a revision of the former is needed (Bousquet, 2012).
Stenolophus (Agonoderus) lineola (Fabricius, 1775)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Rosa (Fall, 1897: 236)
Digitized Records: Anacapa (2 LACM), Santa Cruz (2 LACM), Santa Rosa (1 LACM)
Range: Also known from mainland (Bousquet, 2012).
Notes. Fall (1897) recorded this species as Agonoderus lineola.
Stenolophus (Agonoderus) rugicollis (LeConte, 1859)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: San Miguel (1 LACM)
Range: Also known from mainland (Bousquet, 2012).
Stenolophus (Stenolophus) anceps LeConte, 1857
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: San Miguel (2 SBMNH), Santa Cruz (7 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Stenolophus (Stenolophus) flavipes LeConte, 1858
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (1 LACM; 5 SBMNH), Santa Rosa (14 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Stenolophus (Stenolophus) limbalis LeConte, 1857
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 1050).
Stenolophus (Stenolophus) ochropezus (Say, 1823)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (11 EMEC)
Range: Also known from mainland (Bousquet, 2012).
Lachnophorini
Notes. Three genera and three species of Lachnophorini are known to occur in California (Bousquet, 2012).
Lachnophorus Dejean, 1831
Nomenclatural Authority: Bousquet (2012)
Notes. Only one species of this genus occurs in North America (Bousquet, 2012).
Lachnophorus elegantulus Mannerheim, 1843
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (2 CASC; 1 LACM; 1 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Lebiini
Notes. Fourteen genera and 39 species of Lebiini are known to occur in California (Bousquet, 2012; M. L. Gimmel, 2022, unpublished data).
Apristus Chaudoir, 1846
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Cruz (1 SBMNH), Santa Rosa (4 LACM)
Notes. Seven species of Apristus are known from California (Bousquet, 2012). The genus needs revision (Bousquet, 2012).
Apristus pugetanus Casey, 1920
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (11 CASC)
Range: Also known from mainland (Bousquet, 2012).
Axinopalpus LeConte, 1846
Nomenclatural Authority: Bousquet (2012)
Notes. Three species of Axinopalpus are known from California (Bousquet, 2012). The genus needs revision (Bousquet, 2012).
Axinopalpus biplagiatus (Dejean, 1825)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (1 CASC; 1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Dromius Bonelli, 1810
Nomenclatural Authority: Bousquet (2012)
Notes. One species of Dromius occurs in California (Bousquet, 2012).
Dromius piceus Dejean, 1831
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Anacapa (1 SBMNH), Santa Cruz (3 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Lebia Latreille, 1802
Nomenclatural Authority: Bousquet (2012)
Notes. Ten species of Lebia have been recorded from California (Bousquet, 2012). The North American species were revised by Madge (1967).
Lebia (Lebia) cyanipennis Dejean, 1831
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Madge, 1967).
Lebia (Lebia) perita Casey, 1920
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Madge, 1967).
Microlestes Schmidt-Göbel, 1846
Nomenclatural Authority: Bousquet (2012)
Notes. Five species of Microlestes have been recorded from California (Bousquet, 2012). Lindroth (1969) reviewed the North American species known at the time.
Microlestes undetermined species
Literature Records: none
Digitized Records: Santa Rosa (1 LACM)
Notiophilini
Notes. One genus and four species of Notiophilini have been recorded from California (Bousquet, 2012).
Notiophilus Duméril, 1805
Nomenclatural Authority: Bousquet (2012)
Notes. Four species of Notiophilus have been recorded from California (Bousquet, 2012). These were reviewed and keyed by Lindroth (1961).
Notiophilus semiopacus Eschscholtz, 1833
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1906: 91)
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Fall, 1906; Bousquet, 2012).
Notes. Based on examination of the voucher specimen, the Santa Cruz Island record of “Notiophilus sp.” in Naughton et al. (2014: 303) refers to this species (M. L. Gimmel, 2021, personal observation).
Omophronini
Notes. One genus and six species of Omophronini have been recorded from California (Bousquet, 2012).
Omophron Latreille, 1802
Nomenclatural Authority: Bousquet (2012)
Notes. Six species of Omophron have been recorded from California (Bousquet, 2012). The genus was revised and keyed for North America by Benschoter & Cook (1956).
Omophron dentatum LeConte, 1852
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Cruz (LeConte, 1876: 298; Fall, 1897: 236; Fall & Davis, 1934: 144), Santa Rosa (Fall, 1897: 236; Benschoter & Cook, 1956: 422)
Digitized Records: Santa Catalina (4 CASC; 67 LACM; 1 SBMNH), Santa Cruz (22 CASC; 1 LACM; 9 SBMNH; 8 UASM), Santa Rosa (3 CASC; 12 SBMNH)
Range: Also known from mainland (Benschoter & Cook, 1956; Bousquet, 2012).
Notes. This species was reported from the “Channel Islands” by Bousquet (2012: 389).
Platynini
Notes. Six genera and 33 species of Platynini are known to occur in California (Bousquet, 2012).
Agonum Bonelli, 1810
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Catalina (14 LACM), Santa Cruz (5 LACM; 9 UASM)
Notes. Twenty-one species of Agonum have been recorded from California (Bousquet, 2012). Liebherr (1994) provided a key to the North American species.
Agonum (Agonum) piceolum (LeConte, 1879)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (1 EMEC)
Range: Also known from mainland (Bousquet, 2012).
Agonum (Europhilus) limbatum Motschulsky, 1845
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: San Miguel (1 LACM), Santa Catalina (2 SBMNH), Santa Cruz (1 LACM; 7 SBMNH), Santa Rosa (5 LACM)
Range: Also known from mainland (Bousquet, 2012).
Notes. This species was recorded by Fall (1897) as Platynus variolatus LeConte, 1851. It has been recorded in the recent literature as Agonum variolatum (LeConte, 1851), which is now considered a junior synonym of A. limbatum. Reported from the “Channel Islands” by Bousquet (2012: 1202).
Agonum (Olisares) decorum (Say, 1823)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Cruz (Liebherr, 1983: 350 [map]; Liebherr, 1986: 130 [map])
Digitized Records: Santa Cruz (1 CASC)
Range: Also known from mainland (Liebherr, 1983, 1986).
Agonum (Olisares) punctiforme (Say, 1823)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: San Clemente (9 SBMNH), San Miguel (1 SBMNH), San Nicolas (4 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Anchomenus Bonelli, 1810
Nomenclatural Authority: Bousquet (2012)
Notes. Two species of Anchomenus have been recorded from California (Bousquet, 2012). The genus was revised by Liebherr (1991).
Anchomenus funebris (LeConte, 1854)
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Liebherr, 1991: 55), Santa Catalina (Fall, 1897: 236), Santa Cruz (Fall & Davis, 1934: 143; Liebherr, 1991: 55)
Digitized Records: Santa Cruz (33 CASC; 10 SBMNH; 16 TAMU; 1 UCSB)
Range: Also known from mainland (Liebherr, 1991; Bousquet, 2012).
Notes. This species was recorded as Platynus funebris by Fall (1897) and Fall & Davis (1934). Reported from the “Channel Islands” by Bousquet (2012: 1183).
Platynus Bonelli, 1810
Nomenclatural Authority: Bousquet (2012)
Notes. Three species of Platynus have been recorded from California (Bousquet, 2012). Liebherr & Will (1996) provided a key to the North American species of Platynus known at the time, including all Californian species.
Platynus brunneomarginatus (Mannerheim, 1843)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Seavey, 1892: 262; Fall, 1897: 236), Santa Cruz (LeConte, 1876: 298; Fall, 1897: 236; Fall & Davis, 1934: 143; Liebherr & Will, 1996: 317), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Clemente (3 CASC; 27 LACM; 6 SBMNH), San Miguel (10 SBMNH), Santa Catalina (3 CASC; 4 LACM; 2 SBMNH), Santa Cruz (19 CASC; 10 LACM; 19 SBMNH; 6 TAMU; 1 UCSB; 11 UASM), Santa Rosa (7 SBMNH)
Range: Also known from mainland (Liebherr & Will, 1996; Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 1244).
Tanystoma Motschulsky, 1845
Nomenclatural Authority: Bousquet (2012)
Notes. Four species of Tanystoma have been recorded from California (Bousquet, 2012). The species were revised by Liebherr (1985).
Tanystoma cuyama Liebherr, 1985
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Rosa (9 SBMNH)
Range: Also known from mainland (Liebherr, 1985).
Tanystoma maculicolle (Dejean, 1828)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Anacapa (Liebherr, 1985: 1193 [map]; Liebherr & Hajek, 1986: 22), San Clemente (Liebherr, 1984: 538; Liebherr, 1985: 1193 [map]; Liebherr & Hajek, 1986: 22), San Miguel (Liebherr, 1985: 1193 [map]; Liebherr & Hajek, 1986: 22), San Nicolas (Liebherr, 1985: 1193 [map]; Liebherr & Hajek, 1986: 22), Santa Catalina (Baker, 1905: 59; Liebherr, 1984: 538; Liebherr, 1985: 1193 [map]; Liebherr & Hajek, 1986: 22), Santa Cruz (Fall & Davis, 1934: 143; Liebherr, 1984: 538; Liebherr, 1985: 1193 [map]; Liebherr & Hajek, 1986: 22), Santa Rosa (Fall, 1897: 236; Liebherr, 1985: 1193 [map]; Liebherr & Hajek, 1986: 22)
Digitized Records: San Clemente (8 CASC; 2 LACM; 4 SBMNH), San Miguel (10 CASC; 12 SBMNH), San Nicolas (14 CASC; 1 SBMNH), Santa Catalina (18 CASC; 3 LACM; 13 SBMNH), Santa Cruz (10 CASC; 14 SBMNH; 1 iNat), Santa Rosa (22 CASC; 29 SBMNH; 1 iNat)
Range: Also known from mainland (Liebherr, 1984, 1985; Bousquet, 2012).
Notes. This species was recorded as Platynus maculicollis by Baker (1905), Fall (1897), and Fall & Davis (1934). Populations of this species from the Channel Islands are predominantly brachypterous (Liebherr & Hajek, 1986). Reported from the “Channel Islands” by Bousquet (2012: 1197).
Pogonini
Notes. Two genera and three species of Pogonini have been recorded from California (Bousquet, 2012). The tribe was revised for the Western Hemisphere by Bousquet & Laplante (1997).
Thalassotrechus Van Dyke, 1918
Nomenclatural Authority: Bousquet (2012)
Notes. One species of Thalassotrechus occurs in North America (Bousquet, 2012).
Thalassotrechus barbarae (Horn, 1892)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: San Clemente (2 SBMNH), Santa Catalina (14 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Notes. Bousquet & Laplante (1997) provided a map of known records but did not include any Channel Islands specimens.
Pterostichini
Notes. Two genera and 78 species of Pterostichini are known to occur in California (Bousquet, 2012).
Poecilus Bonelli, 1810
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Cruz (1 iNat)
Notes. Five species of Poecilus have been recorded from California (Bousquet, 2012). The genus needs revision (Bousquet, 2012), but Lindroth’s (1966) key includes all but one of the California species. The iNaturalist observation cited above appears to represent P. laetulus.
Poecilus (Poecilus) laetulus (LeConte, 1863)
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Cockerell, 1940: 285), Santa Cruz (LeConte, 1876: 298; Fall, 1897: 236; Fall & Davis, 1934: 143), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Clemente (1 SBMNH), Santa Catalina (1 SBMNH)
Range: Also known from mainland (Fall, 1901; Bousquet, 2012).
Notes. Fall (1901: 45) collectively reported this and other species from “the islands off the coast”. This species was reported as Pterostichus laetulus LeConte by LeConte (1876), Fall (1897, 1901), and Fall & Davis (1934). Reported from the “Channel Islands” by Bousquet (2012: 775).
Pterostichus Bonelli, 1810
Nomenclatural Authority: Bousquet (2012)
Literature Records (genus-only): Santa Cruz (Naughton et al., 2014: 303), Santa Rosa (Fall, 1897: 236)
Digitized Records (genus-only): San Clemente (24 LACM; 11 SBMNH), San Miguel (1 LACM; 5 SBMNH), Santa Catalina (25 LACM; 5 SBMNH), Santa Cruz (11 LACM; 26 SBMNH; 1 UASM), Santa Rosa (5 LACM; 18 SBMNH)
Notes. Five subgenera and 73 species of Pterostichus have been recorded from California (Bousquet, 2012). Fall (1901: 45) collectively reported an undetermined species and other species of Pterostichus from “the islands off the coast”. Hypherpes Chaudoir, 1838 is by far the largest subgenus of Pterostichus in California, with 48 species recorded (Bousquet, 2012). It is badly in need of revision (Bousquet, 2012). Most or all genus-only digitized records cited above belong to this subgenus. A record of the northern Californian Pterostichus (Hypherpes) congestus (Ménétriés, 1843) from Santa Catalina Island provided to the California Beetle Database probably originated from a specimen determined by H.C. Fall in the Museum of Comparative Zoology, Harvard University (S. Miller, 2022, personal communication); this species record is the result of an erroneous synonymy (see Bousquet, 2012: 846) with P. illustris (K. Will, 2022, personal communication). A record of Pterostichus (Hypherpes) lama (Ménétriés, 1843) from Santa Rosa Island exists in the SBMNH database, but no specimen was found. This species record is geographically suspect (K. Will, 2021, personal communication).
Pterostichus (Bothriopterus) lustrans LeConte, 1851
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Cruz (Fall & Davis, 1934: 143)
Digitized Records: Santa Cruz (10 SBMNH)
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 795). Some of the digital records were erroneously identified as Pterostichus adstrictus Eschscholtz, 1823 previously.
Pterostichus (Hypherpes) gliscans Casey, 1913
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Casey, 1913a: 119; Miller, 1985a: 19; Bousquet, 2012: 847)
Digitized Records: San Miguel (28 LACM)
Range: Endemic (Casey, 1913a; Miller, 1985a; Bousquet, 2012).
Pterostichus (Hypherpes) illustris LeConte, 1851
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Catalina (2 CASC; 1 EMEC)
Range: Also known from mainland (Bousquet, 2012).
Pterostichus (Hypherpes) inermis Fall, 1901
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (28 EMEC)
Range: Also known from mainland (Bousquet, 2012).
Pterostichus (Hypherpes) isabellae LeConte, 1851
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Fall, 1897: 236), Santa Catalina (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Fall, 1901; Bousquet, 2012).
Notes. Fall (1901: 45) collectively reported this and other species from “the islands off the coast”. Reported from the “Channel Islands” by Bousquet (2012: 847).
Pterostichus (Hypherpes) jacobinus Casey, 1913
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Catalina (1 EMEC)
Range: Also known from mainland (Bousquet, 2012).
Pterostichus (Hypherpes) menetriesii LeConte, 1873
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Rosa (Fall, 1897: 236; Fall, 1901: 44; Bousquet, 2012: 849)
Digitized Records: San Miguel (84 CASC), Santa Cruz (4 CASC)
Range: Also known from mainland (Bousquet, 2012).
Notes. This species was thought to be endemic to the Channel Islands by Fall (1901).
Sphodrini
Notes. Two genera and four species of Sphodrini are known to occur in California (Bousquet, 2012).
Calathus Bonelli, 1810
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Santa Cruz (18 UASM)
Notes. Three species of Calathus have been recorded from California (Bousquet, 2012; M. L. Gimmel, 2022, unpublished data). The Western Hemisphere species were revised by Ball & Negre (1972).
Calathus (Neocalathus) ruficollis Dejean, 1828
Nomenclatural Authority: Bousquet (2012)
Literature Records: Anacapa (Cockerell, 1940: 285; Ball & Negre, 1972: 481 [map]), San Clemente (Ball & Negre, 1972: 481 [map]), San Miguel (Cockerell, 1940: 285; Ball & Negre, 1972: 481 [map]), Santa Catalina (Fall, 1897: 236; Baker, 1905: 59; Cockerell, 1940: 285; Ball & Negre, 1972: 481 [map]), Santa Cruz (LeConte, 1876: 298; Fall, 1897: 236; Fall & Davis, 1934: 143; Cockerell, 1940: 285; Ball & Negre, 1972: 481 [map]; Naughton et al., 2014: 303), Santa Rosa (Fall, 1897: 236; Ball & Negre, 1972: 481 [map])
Digitized Records: Anacapa (6 LACM; 1 SBMNH), San Miguel (183 LACM; 16 SBMNH), Santa Catalina (1 CSUC; 9 LACM; 21 SBMNH), Santa Cruz (18 LACM; 31 SBMNH; 4 UCSB), Santa Rosa (55 LACM; 18 SBMNH)
Range: Also known from mainland (Ball & Negre, 1972; Bousquet, 2012).
Notes. This species was cited as C. ruficollis, without subspecies, by authors prior to Ball & Negre (1972), who determined that the only subspecies occurring in southern California is the nominate subspecies, C. r. ruficollis Dejean, 1828. Prior records of Calathus obscurus LeConte (Baker, 1905: 59; Fall, 1897: 236; Cockerell, 1940: 285) also refer to this species, which was later synonymized with C. ruficollis (see Ball & Negre, 1972). Cockerell’s (1940: 285) records from Anacapa, San Miguel, and Santa Cruz were cited as Calathus insularis Casey, which was also later synonymized with C. ruficollis (see Ball & Negre, 1972). Reported from the “Channel Islands” by Bousquet (2012: 1169).
Laemostenus Bonelli, 1810
Nomenclatural Authority: Bousquet (2012)
Notes. One adventive species of Laemostenus occurs in California (Bousquet, 2012).
Laemostenus (Laemostenus) complanatus (Dejean, 1828)
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: San Nicolas (9 SBMNH), Santa Catalina (1 CASC; 2 LACM)
Range: Also known from mainland (Bousquet, 2012).
Notes. This species was introduced from the Palearctic region to North America (Bousquet, 2012: 1173).
Zabrini
Notes. One genus and 36 species of Zabrini are known to occur in California (Bousquet, 2012).
Amara Bonelli, 1810
Nomenclatural Authority: Bousquet (2012)
Digitized Records (genus-only): Anacapa (6 ASUHIC), San Clemente (7 LACM), San Nicolas (3 LACM; 1 SBMNH), Santa Barbara (17 LACM; 4 SBMNH), Santa Cruz (7 UCSB; 1 iNat), Santa Rosa (1 LACM; 9 SBMNH)
Notes. Nine subgenera and 36 species of Amara have been recorded from California (Bousquet, 2012). Miller & Miller (1985) reported many specimens from Santa Barbara Island not determined to species, noting the need for revision of the genus.
Amara (Amara) aurata Dejean, 1828
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Casey, 1918: 274; Miller, 1985a: 19; Hieke, 1993: 121; Bousquet, 2012: 929)
Digitized Records: San Clemente (3 CASC), Santa Cruz (2 CASC)
Range: Also known from mainland (Hieke, 1993; Bousquet, 2012).
Notes. This species was recorded as the purported endemic Celia clementina Casey, 1918 by Casey (1918), and by Miller (1985a) as Amara clementina. It was subsequently synonymized with A. aurata by Hieke (1993).
Amara (Amara) conflata LeConte, 1855
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: Santa Cruz (4 CASC)
Range: Also known from mainland (Bousquet, 2012).
Notes. Reported from the “Channel Islands” by Bousquet (2012: 939).
Amara (Amara) pomona Casey, 1918
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Cruz (Hieke, 1993: 114), Santa Rosa (Hieke, 1993: 114)
Digitized Records: none
Range: Also known from mainland (Hieke, 1993).
Notes. This species was recorded as Amara brunnipes Motschulsky, 1859 by Hieke (1993), which is a junior primary homonym with A. pomona as the next available name.
Amara (Bradytus) insignis Dejean, 1831
Nomenclatural Authority: Bousquet (2012)
Literature Records: Santa Catalina (Fall, 1897: 236; Baker, 1905: 59), Santa Rosa (Fall, 1897: 236)
Digitized Records: Santa Catalina (5 CASC), Santa Cruz (4 CASC)
Range: Also known from mainland (Fall, 1901; Bousquet, 2012).
Notes. Fall (1901: 45) reported this species from “islands”. Reported from the “Channel Islands” by Bousquet (2012: 905). SBMNH specimens of the “Amara insignis group”, represented by the two species A. insignis and A. insularis, were determined based on the key in Lindroth (1968: 659). The characters cited by Lindroth in both the key and species accounts (Lindroth, 1968: 659) involve the punctation of the pronotum and development of the hind wings. However, while mainland specimens of A. insignis appear to be consistently punctate basally, this character appears to break down, at least among northern island populations (M. L. Gimmel, 2021, personal observation). The shape of the aedeagal apices do not seem to covary with this character and, in fact, impunctate “A. insularis” forms do not appear to possess a distinctive aedeagus when compared to mainland A. insignis. Additionally, hind wings appear to be developed in all specimens where this character is visible (M. L. Gimmel, 2021, personal observation). In the end, all island members of the group housed in SBMNH were determined as A. insularis.
Amara (Bradytus) insularis Horn, 1875
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Horn, 1875: 128; Fall, 1897: 236; Casey, 1918: 295; Cockerell, 1940: 285; Lindroth, 1968: 692; Miller, 1985a: 19; Miller & Miller, 1985: 123), San Nicolas (Fall, 1897: 236; Hayward, 1908: 51; Cockerell, 1940: 285), Santa Barbara (Fall, 1897: 236; Hayward, 1908: 51; Cockerell, 1940: 285), Santa Rosa (Hayward, 1908: 51)
Digitized Records: Anacapa (4 CASC), San Clemente (31 CASC; 1 LACM; 10 SBMNH), San Miguel (2 CASC; 7 LACM; 2 SBMNH), San Nicolas (18 CASC; 201 LACM; 2 SBMNH), Santa Barbara (3 CASC; 89 LACM; 36 SBMNH), Santa Catalina (6 CASC), Santa Cruz (3 CASC; 8 SBMNH), Santa Rosa (3 CASC; 2 LACM; 18 SBMNH)
Range: Endemic (Horn, 1875; Fall, 1897; Hayward, 1908; Casey, 1918; Cockerell, 1940; Lindroth, 1968; Miller, 1985a; Bousquet, 2012).
Notes. Fall (1901) reported this species from “all the islands”. Lindroth (1968: 692) incorrectly stated that this species was only known from the type locality (San Clemente Island), though Miller (1985a) noted that “published records from other islands (than San Clemente) cannot be trusted.” This species, however, is questionably distinct from A. insignis (see Notes in that species account above).
Amara (Celia) californica Dejean, 1828
Nomenclatural Authority: Bousquet (2012)
Literature Records: San Clemente (Casey, 1918: 294; Lindroth, 1968: 693; Lindroth, 1975: 131; Bousquet, 2012: 915), Santa Cruz (LeConte, 1876: 298; Fall, 1897: 236; Fall & Davis, 1934: 143), Santa Rosa (Fall, 1897: 236)
Digitized Records: Anacapa (40 CASC), San Clemente (5 CASC), San Miguel (21 CASC), San Nicolas (17 CASC), Santa Cruz (3 CASC), Santa Rosa (1 CASC)
Range: Also known from mainland (Lindroth, 1968; Bousquet, 2012).
Notes. Recorded by Fall & Davis (1934) as Celia californica. Amara perspecta Casey, 1918, described as endemic from San Clemente Island (Casey, 1918: 294), was synonymized with A. californica by Lindroth (1968: 693). The only subspecies of A. californica occurring in California is the nominate subspecies, A. c. californica Dejean, 1828 (Bousquet, 2012).
Amara (Zezea) scitula Zimmermann, 1832
Nomenclatural Authority: Bousquet (2012)
Literature Records: none
Digitized Records: San Miguel (1 CASC)
Range: Also known from mainland (Bousquet, 2012).
Cicindelidae
Notes. The North American fauna of tiger beetles (often treated as a subfamily of Carabidae) was the subject of a thorough field guide by Pearson et al. (2015). Nagano (1982, 1985) assessed the Channel Islands fauna in detail. Three tribes, eight genera, and 34 species have been recorded from California (Bousquet, 2012; Pearson et al., 2015).
Cicindelini
Notes. The California fauna of Cicindelini consists of five genera and 29 species (Bousquet, 2012; Pearson et al., 2015). We follow the genus-level arrangement of Pearson et al. (2015) below.
Cicindela Linnaeus, 1758
Nomenclatural Authority: Bousquet (2012)
Digitized Records: Santa Cruz (6 LACM)
Notes. Seventeen species of Cicindela are known to occur in California (Bousquet, 2012; Pearson et al., 2015).
Cicindela hirticollis Say, 1817
Nomenclatural Authority: Bousquet (2012), Pearson et al. (2015)
Literature Records: Santa Catalina (Nagano, 1982: 36; Nagano, 1985: 106; Graves, Krejci & Graves, 1988: 653 [map]), Santa Rosa (Nagano, 1982: 36; Nagano, 1985: 106; Graves, Krejci & Graves, 1988: 653 [map])
Digitized Records: none
Range: Also known from mainland (Nagano, 1982, 1985; Graves, Krejci & Graves, 1988; Bousquet, 2012).
Notes. All coastal Southern California C. hirticollis belong to the subspecies C. h. gravida LeConte, 1851 (Nagano, 1982, 1985; Graves, Krejci & Graves, 1988). Recorded from the “Channel Islands” by Bousquet (2012: 362).
Cicindela oregona LeConte, 1856
Nomenclatural Authority: Bousquet (2012), Pearson et al. (2015)
Literature Records: Anacapa (Nagano, 1985: 106), San Miguel (Nagano, 1982: 36; Nagano, 1985: 105), San Nicolas (Nagano, 1982: 35; Nagano, 1985: 106), Santa Catalina (Nagano, 1982: 36; Nagano, 1985: 106), Santa Cruz (Freitag, 1965: 136; Nagano, 1982: 35; Nagano, 1985: 105), Santa Rosa (Fall, 1897: 236; Nagano, 1982: 36; Nagano, 1985: 105)
Digitized Records: Anacapa (1 LACM), San Miguel (1 SBMNH; 14 LACM), San Nicolas (26 LACM), Santa Cruz (20 SBMNH; 1 UCSB; 36 LACM; 12 YPMC), Santa Rosa (2 SBMNH; 17 LACM)
Range: Also known from mainland (Freitag, 1965; Nagano, 1982, 1985; Bousquet, 2012).
Notes. All coastal Southern California C. oregona belong to the nominate subspecies, C. o. oregona LeConte, 1856 (Nagano, 1982, 1985). Recorded from the “Channel Islands” by Bousquet (2012: 367).
Cicindela senilis Horn, 1867
Nomenclatural Authority: Bousquet (2012), Pearson et al. (2015)
Literature Records: San Clemente (Nagano, 1982: 37; Nagano, 1985: 107)
Digitized Records: San Clemente (60 LACM; 3 SBMNH)
Range: Also known from mainland (Nagano, 1982, 1985; Bousquet, 2012).
Notes. The Channel Islands form of this species was referred to as the subspecies C. s. frosti Varas Arangua, 1928 by Nagano (1982, 1985), but subspecies were not recognized for this species by Bousquet (2012). Pearson et al. (2015) noted that “some authors” recognize these subspecies. This species was recorded from the “Channel Islands” by Bousquet (2012: 325).
Cicindelidia Rivalier, 1954
Nomenclatural Authority: Bousquet (2012), Pearson et al. (2015)
Notes. Four species of Cicindelidia are known to occur in California (Bousquet, 2012; Pearson et al., 2015). Bousquet (2012) treated this taxon as a subgenus of Cicindela.
Cicindelidia hemorrhagica (LeConte, 1851)
Nomenclatural Authority: Bousquet (2012), Pearson et al. (2015)
Literature Records: San Nicolas (Nagano, 1982: 39; Nagano, 1985: 109), Santa Cruz (Nagano, 1982: 39; Nagano, 1985: 109)
Digitized Records: San Nicolas (19 SBMNH), Santa Cruz (2 SBMNH; 2 UCSB; 62 YPMC)
Range: Also known from mainland (Nagano, 1982, 1985; Bousquet, 2012).
Notes. All Channel Islands C. hemorrhagica belong to the nominate subspecies, C. h. hemorrhagica (LeConte, 1851) (Nagano, 1982, 1985). Recorded from the “Channel Islands” by Bousquet (2012: 314); Nagano (1982, 1985) listed this species as Cicindela haemorrhagica, and Bousquet (2012) as Cicindela hemorrhagica.
Cicindelidia trifasciata (Fabricius, 1781)
Nomenclatural Authority: Bousquet (2012), Pearson et al. (2015)
Literature Records: Santa Catalina (Nagano, 1982: 38; Nagano, 1985: 109)
Digitized Records: Santa Catalina (3 SBMNH; 132 LACM; 1 SDNHM)
Range: Also known from mainland (Nagano, 1982, 1985; Bousquet, 2012).
Notes. All Southern California C. trifasciata belong to the subspecies C. t. sigmoidea (LeConte, 1851) (Nagano, 1982, 1985). Recorded from the “Channel Islands” by Bousquet (2012: 326). Nagano (1982, 1985) and Bousquet (2012) listed this species as Cicindela trifasciata sigmoidea.
Dytiscidae
Notes. Seven subfamilies, 36 genera, and 156 species of this family are known to occur in California (Challet & Brett, 1998; M. L. Gimmel, 2022, unpublished data). Challet & Brett (1998) provided an excellent county-by-county summary of their distribution in California, though the nomenclature of some of the genera is now out of date. Likewise, Larson, Alarie & Roughley (2000) provided excellent illustrated keys for most North American species of the family, but the generic classification has changed in many groups. Nilsson & Hájek (2018) provided an updated world catalog for the family.
Agabinae: Agabini
Notes. Two tribes, six genera, and 38 species of Agabinae are known from California, of which just one species belongs to Hydrotrupini and the remainder belong to Agabini (Challet & Brett, 1998; M. L. Gimmel, 2022, unpublished data).
Agabinus Crotch, 1873
Nomenclatural Authority: Nilsson & Hájek (2018)
Digitized Records (genus-only): Santa Cruz (1 UCSB)
Notes. Two species of Agabinus are known to occur in California (Challet & Brett, 1998).
Agabinus glabrellus (Motschulsky, 1859)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Catalina (Fall, 1897: 236; Fall, 1901: 53; Challet, 1987: 13), Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: Santa Catalina (1 LACM; 8 SBMNH), Santa Cruz (3 SBMNH), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Agabinus sculpturellus Zimmermann, 1919
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: none
Range: Also known from mainland (Challet & Brett, 1998).
Agabus Leach, 1817
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. Sixteen species of Agabus are known to occur in California (Challet & Brett, 1998; M. L. Gimmel, 2022, unpublished data). These species are best separated by examination of the male genitalia (Larson, Alarie & Roughley, 2000).
Agabus obsoletus LeConte, 1858
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: none
Digitized Records: San Miguel (5 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Ilybiosoma Crotch, 1873
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. Eight species of Ilybiosoma are known to occur in California (Challet & Brett, 1998; M. L. Gimmel, 2022, unpublished data).
Ilybiosoma lugens (LeConte, 1852)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Catalina (Challet, 1987: 13), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Nicolas (2 SBMNH), Santa Catalina (58 SBMNH), Santa Cruz (9 SBMNH), Santa Rosa (6 LACM; 10 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Notes. Reported as Agabus lugens by Fall (1897) and Challet (1987). Specimens of this species are indistinguishable from another California species, Ilybiosoma perplexum (Sharp, 1882), except by male genitalia (Larson, Alarie & Roughley, 2000). The San Nicolas Island specimens were both female; consequently, once discovered, male specimens matching the description of I. lugens from that island should be dissected to confirm their identity.
Ilybiosoma regulare (LeConte, 1852)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: none
Digitized Records: Santa Cruz (5 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Ilybiosoma seriatum (Say, 1823)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Ilybius Erichson, 1832
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. Ten species of Ilybius are known to occur in California (Challet & Brett, 1998; M. L. Gimmel, 2022, unpublished data). These species are best separated by examination of the male genitalia (Larson, Alarie & Roughley, 2000).
Ilybius discors (LeConte, 1861)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: none
Range: Also known from mainland (Challet & Brett, 1998).
Notes. Reported by Furlong & Wenner (2002) as Agabus discors.
Ilybius lineellus (LeConte, 1861)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Ilybius walsinghami (Crotch, 1873)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: San Clemente (Larson, 1996: 665 [map only])
Digitized Records: Santa Rosa (17 SBMNH)
Range: Also known from mainland (Larson, 1996; Challet & Brett, 1998).
Notes. Reported by Larson (1996) as Agabus walsinghami.
Colymbetinae
Notes. Two genera, and 10 species of Colymbetinae have been recorded from California (Challet & Brett, 1998; M. L. Gimmel, 2022, unpublished data), all belonging to the tribe Colymbetini.
Rhantus Dejean, 1833
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. Seven species of Rhantus are known to occur in California (Challet & Brett, 1998; M. L. Gimmel, 2022, unpublished data).
Rhantus gutticollis (Say, 1830)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: San Clemente (Zimmerman & Smith, 1975: 52), Santa Catalina (Challet, 1987: 13), Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: San Clemente (28 SBMNH), San Nicolas (1 SBMNH), Santa Catalina (57 LACM; 6 SBMNH), Santa Cruz (5 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Zimmerman & Smith, 1975; Challet & Brett, 1998).
Dytiscinae
Notes. Five tribes, seven genera, and 14 species of Dytiscinae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Dytiscini
Notes. One genus and four species of Dytiscini are known to occur in California (Challet & Brett, 1998).
Dytiscus Linnaeus, 1758
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. Four species of Dytiscus are known to occur in California (Challet & Brett, 1998).
Dytiscus marginicollis LeConte, 1845
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Catalina (Challet, 1987: 13)
Digitized Records: San Clemente (6 SBMNH), Santa Catalina (4 LACM)
Range: Also known from mainland (Challet & Brett, 1998).
Eretini
Notes. One genus and species of Eretini is known to occur in California (Challet & Brett, 1998).
Eretes Laporte, 1833
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. One species of Eretes has been reported from California (Challet & Brett, 1998). This genus was revised by Miller (2002).
Eretes sticticus (Linnaeus, 1767)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: none
Digitized Records: Santa Catalina (2 LACM)
Range: Also known from mainland (Challet & Brett, 1998; Miller, 2002).
Hydroporinae
Notes. Six tribes, 19 genera, and 85 species of Hydroporinae are known to occur in California (Challet & Brett, 1998; M. L. Gimmel, 2022, unpublished data).
Bidessini
Notes. Three genera and 10 species of Bidessini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Liodessus Guignot, 1939
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. Two species of Liodessus have been reported from California (Miller, 1998). The L. affinis group, containing both California species, was revised by Miller (1998).
Liodessus obscurellus (LeConte, 1852)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: none
Digitized Records: San Miguel (3 SBMNH), Santa Catalina (1 SBMNH), Santa Cruz (11 SBMNH), Santa Rosa (8 SBMNH)
Range: Also known from mainland (Miller, 1998).
Notes. This species was treated as a junior synonym of Liodessus affinis (Say, 1823) until recently (Miller, 1998).
Neoclypeodytes Young, 1967
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. Seven species of Neoclypeodytes have been reported from California (Miller, 2001). The genus was revised by Miller (2001).
Neoclypeodytes pictodes (Sharp, 1882)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: none
Digitized Records: Santa Rosa (16 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998; Miller, 2001).
Uvarus Guignot, 1939
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. One species of Uvarus is known to occur in California (Challet & Brett, 1998).
Uvarus subtilis (LeConte, 1852)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: none
Digitized Records: Santa Cruz (6 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Hydroporini
Notes. Eleven genera and 49 species of Hydroporini are known to occur in California (Challet & Brett, 1998; M. L. Gimmel, 2022, unpublished data).
Leconectes Fery & Ribera, 2018
Nomenclatural Authority: Fery & Ribera (2018)
Notes. One species of Leconectes is known to occur in California (M. L. Gimmel, 2022, unpublished data).
Leconectes striatellus (LeConte, 1852)
Nomenclatural Authority: Fery & Ribera (2018)
Literature Records: Santa Catalina (Challet, 1987: 13), Santa Cruz (Short & Caterino, 2009: 405), Santa Rosa (Fall, 1897: 236; Short & Caterino, 2009: 405)
Digitized Records: San Clemente (1 SBMNH), San Miguel (2 SBMNH), Santa Catalina (54 LACM; 2 SBMNH), Santa Cruz (14 SBMNH; 1 UCSB), Santa Rosa (11 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998; Short & Caterino, 2009; Fery & Ribera, 2018).
Notes. Fall (1897) and Challet (1987) reported this species as Deronectes striatellus, while Short & Caterino (2009) reported it as Stictotarsus striatellus; it has also been known as Boreonectes striatellus until recently (Fery & Ribera, 2018).
Sanfilippodytes Franciscolo, 1979
Nomenclatural Authority: Nilsson & Hájek (2018)
Digitized Records (genus-only): Santa Cruz (8 EMEC)
Notes. Thirteen species of Sanfilippodytes are known to occur in California (Challet & Brett, 1998).
Sanfilippodytes barbarensis (Wallis, 1933)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: none
Digitized Records: Santa Catalina (7 SBMNH), Santa Cruz (9 SBMNH), Santa Rosa (34 SBMNH)
Range: Also known from mainland (Nilsson & Hájek, 2018).
Sanfilippodytes latebrosus (LeConte, 1852)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (3 SBMNH), Santa Rosa (12 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Sanfilippodytes vilis (LeConte, 1852)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Catalina (Challet, 1987: 13), Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: San Miguel (7 SBMNH), San Nicolas (8 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Notes. According to Challet (1987: 13), Fall (1897) originally reported this species as Hydroporus vilis from Santa Catalina and Santa Rosa, but later (Fall, 1923) indicated that these records referred to H. belfragei (see S. williami below). All of the foregoing species were subsequently transferred to Sanfilippodytes.
Sanfilippodytes williami (Rochette, 1986)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Catalina (Fall, 1897: 236; Fall, 1923: 59; Rochette, 1986: 341; Challet, 1987: 13), Santa Rosa (Fall, 1897: 236)
Digitized Records: Santa Cruz (3 SBMNH)
Range: Also known from mainland (Rochette, 1986; Challet & Brett, 1998).
Notes. Fall (1897: 236) misidentified this species as Hydroporus vilis LeConte. Fall (1923) and Challet (1987) corrected the identification to Hydroporus belfragei Sharp, 1882. Finally, Rochette (1986) described the new species H. williami that included the island material. All of the foregoing species were subsequently transferred to Sanfilippodytes.
Hydrovatini
Notes. One genus and two species of Hydrovatini are known to occur in California (Challet & Brett, 1998).
Hydrovatus Motschulsky, 1853
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. Two species of Hydrovatus are known to occur in California (Challet & Brett, 1998).
Hydrovatus brevipes Sharp, 1882
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: none
Range: Also known from mainland (Challet & Brett, 1998).
Hygrotini
Notes. Two genera and 21 species of Hygrotini are known to occur in California (Challet & Brett, 1998).
Hygrotus Stephens, 1828
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. Twenty species of Hygrotus are known to occur in California (Challet & Brett, 1998).
Hygrotus (Leptolambus) lutescens (LeConte, 1852)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Catalina (Challet, 1987: 13)
Digitized Records: San Clemente (14 SBMNH), San Nicolas (7 SBMNH), Santa Catalina (63 LACM; 10 SBMNH), Santa Cruz (12 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Laccophilinae
Notes. One genus and six species of Laccophilinae are known to occur in California (Challet & Brett, 1998), all belonging to the tribe Laccophilini.
Laccophilus Leach, 1815
Nomenclatural Authority: Nilsson & Hájek (2018)
Notes. Six species of Laccophilus are known to occur in California (Challet & Brett, 1998).
Laccophilus fasciatus Aubé, 1838
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Catalina (Challet, 1987: 13)
Digitized Records: San Clemente (16 SBMNH), Santa Catalina (34 LACM; 1 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Notes. Challet (1987) indicated that the subspecies occurring on the islands is L. f. terminalis Sharp, 1882.
Laccophilus maculosus (Germar, 1823)
Nomenclatural Authority: Nilsson & Hájek (2018)
Literature Records: Santa Catalina (Challet, 1987: 13)
Digitized Records: Santa Catalina (24 LACM; 6 SBMNH)
Range: Also known from mainland (Challet & Brett, 1998).
Notes. Challet (1987) indicated the subspecies occurring on the islands is L. m. decipiens LeConte, 1852.
Gyrinidae
Notes. Two genera and 11 species of this family are known to occur in California, all of which belong to the subfamily Gyrininae (M. L. Gimmel, 2022, unpublished data).
Gyrinus Müller, 1764
Nomenclatural Authority: Oygur & Wolfe (1991)
Notes. Nine species of Gyrinus are known to occur in California (Oygur & Wolfe, 1991). Additional species of Gyrinus are likely to occur on the Channel Islands. Oygur & Wolfe (1991) revised the genus for North America and provided an identification key to species.
Gyrinus plicifer LeConte, 1852
Nomenclatural Authority: Oygur & Wolfe (1991)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: Santa Catalina (6 SBMNH), Santa Cruz (1 CASC; 20 SBMNH; 3 UCSB), Santa Rosa (8 SBMNH)
Range: Also known from mainland (Oygur & Wolfe, 1991).
Haliplidae
Notes. Three genera and 20 species of this family are known to occur in California (van Vondel, 2021). van Vondel (2021) provided a recent revision of the North American species. Leech & Chandler (1956) provided keys to the California species known at the time.
Haliplus Latreille, 1802
Nomenclatural Authority: van Vondel (2021)
Notes. Thirteen species of Haliplus have been reported to occur in California (van Vondel, 2021).
Haliplus undetermined species
Literature Records: none
Digitized Records: Santa Catalina (4 LACM)
Peltodytes Régimbart, 1879
Nomenclatural Authority: van Vondel (2021)
Notes. Five species of Peltodytes have been reported to occur in California (van Vondel, 2021).
Peltodytes (Neopeltodytes) simplex (LeConte, 1852)
Nomenclatural Authority: van Vondel (2021)
Literature Records: Santa Catalina (van Vondel, 2021: 258), Santa Cruz (Furlong & Wenner, 2002: 250; van Vondel, 2021: 258)
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (17 SBMNH; 1 UCSB), Santa Rosa (9 SBMNH)
Range: Also known from mainland (van Vondel, 2021).
MYXOPHAGA
Hydroscaphidae
Notes. Only one species of this family is known to occur in California (Reichardt, 1973). The New World Hydroscaphidae were treated by Reichardt & Hinton (1976).
Hydroscapha LeConte, 1874
Nomenclatural Authority: Reichardt & Hinton (1976)
Notes. One species of Hydroscapha is known from California (Reichardt, 1973).
Hydroscapha natans LeConte, 1874
Nomenclatural Authority: Reichardt & Hinton (1976)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: Santa Cruz (9 SBMNH), Santa Rosa (9 SBMNH)
Range: Also known from mainland (Reichardt & Hinton, 1976).
Sphaeriusidae, NEW FAMILY RECORD
Notes. One species of this family is known to occur in California (Reichardt, 1973). This family has also been known as Sphaeridae, Sphaeriidae, and Microsporidae in the literature.
Sphaerius Waltl, 1838
Nomenclatural Authority: Reichardt (1973)
Notes. One species of Sphaerius has been recorded from California (Reichardt, 1973).
Sphaerius politus Horn, 1868
Nomenclatural Authority: Reichardt (1973)
Literature Records: none
Digitized Records: Santa Catalina (4 SBMNH), Santa Cruz (11 SBMNH)
Range: Also known from mainland (Horn, 1868).
Notes. This species was described from Visalia, Tulare County, California (Horn, 1868).
POLYPHAGA
SCIRTOIDEA
Scirtidae
Notes. Four genera and 14 species of Scirtidae are known from California (M. L. Gimmel, 2022, unpublished data).
Scirtidae undetermined genus and species
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: none
Notes. Furlong & Wenner (2002) did not indicate number of specimens, adult or larva, or specimen deposition of their record of “Scirtidae” from Santa Cruz Island.
CLAMBOIDEA
Clambidae
Notes. There are two genera and five species of Clambidae recorded from California (Endrődy-Younga, 1981). This family was treated for North America by Endrődy-Younga (1981), who provided keys to all known species.
Clambus Fischer von Waldheim, 1820
Nomenclatural Authority: Endrődy-Younga (1981)
Notes. Four species of Clambus have been recorded from California (Endrődy-Younga, 1981).
Clambus undetermined species
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Notes. The Santa Cruz Island specimen housed in SBMNH is a female, and therefore cannot presently be determined to species (M. L. Gimmel, 2021, personal observation).
Loricaster Mulsant & Rey, 1861
Nomenclatural Authority: Endrődy-Younga (1981)
Notes. One species of Loricaster has been recorded from California (Endrődy-Younga, 1981).
Loricaster rotundus Grigarick & Schuster, 1961
Nomenclatural Authority: Endrődy-Younga (1981)
Literature Records: none
Digitized Records: San Clemente (2 SBMNH), Santa Catalina (10 SBMNH)
Range: Also known from mainland (Endrődy-Younga, 1981).
Notes. The genus Loricaster was recorded from Santa Catalina Island by Caterino & Chandler (2010: 191); that record certainly refers to this species.
DASCILLOIDEA
Dascillidae, NEW FAMILY RECORD
Notes. Two subfamilies, two genera, and three species of Dascillidae are known to occur in California (Johnston & Gimmel, 2020). Johnston & Gimmel (2020) reviewed the family for North America.
Karumiinae
Notes. One genus and two species of Karumiinae have been recorded from California (Johnston & Gimmel, 2020).
Anorus LeConte, 1859
Nomenclatural Authority: Johnston & Gimmel (2020)
Notes. Two species of Anorus have been recorded from California (Johnston & Gimmel, 2020).
Anorus piceus LeConte, 1859
Nomenclatural Authority: Johnston & Gimmel (2020)
Literature Records: none
Digitized Records: San Clemente (1 LACM), Santa Catalina (3 LACM), Santa Cruz (4 LACM), Santa Rosa (23 LACM)
Range: Also known from mainland (Johnston & Gimmel, 2020).
Notes. Johnston & Gimmel (2020) saw no specimens from the Channel Islands during the course of their study. The specimens listed above were examined by us and are certainly within the A. piceus concept of the recent revision and are reliably labeled from the islands from multiple collecting events. The flightless morphology and life history of females of this species (Johnston & Gimmel, 2020) presents a fascinating question of how these island populations became established.
Rhipiceridae, NEW FAMILY RECORD
Notes. One genus and two species of Rhipiceridae are known to occur in California (Schnepp & Powell, 2018).
Sandalus Knoch, 1801
Nomenclatural Authority: Schnepp & Powell (2018)
Notes. Two species of Sandalus have been recorded from California (Schnepp & Powell, 2018).
Sandalus cribricollis Van Dyke, 1923
Nomenclatural Authority: Schnepp & Powell (2018)
Literature Records: none
Digitized Records: Santa Catalina (1 UCRC)
Range: Also known from mainland (Van Dyke, 1923).
BUPRESTOIDEA
Buprestidae
Notes. Four subfamilies, 35 genera, and 319 species of Buprestidae are known to occur in California (Nelson et al., 2008; M. L. Gimmel, 2022, unpublished data). This family was the subject of an extensive distributional catalog and bibliography by Nelson et al. (2008).
Agrilinae: Agrilini
Notes. Two tribes, three genera, and 33 species of Agrilinae are known to occur in California, of which just one species belongs to Tracheini and the remainder to Agrilini (Nelson et al., 2008; M. L. Gimmel, 2022, unpublished data).
Agrilus Curtis, 1825
Nomenclatural Authority: Nelson et al. (2008)
Notes. Thirty-one species of Agrilus have been reported from California (Nelson et al., 2008; M. L. Gimmel, 2022, unpublished data).
Agrilus quadriguttatus Gory, 1841
Nomenclatural Authority: Nelson et al. (2008)
Literature Records: none
Digitized Records: Santa Cruz (14 SBMNH)
Range: Also known from mainland (Nelson et al., 2008).
Notes. The subspecies of A. quadriguttatus occurring on the Channel Islands is A. q. niveiventris Horn, 1891 (N. Woodley, 2021, personal communication).
Buprestinae
Notes. Seven tribes, 15 genera, and 132 species of Buprestinae are known to occur in California (Nelson et al., 2008; M. L. Gimmel, 2022, unpublished data).
Anthaxiini
Notes. One genus and 24 species of Anthaxiini are known to occur in California (Nelson et al., 2008).
Anthaxia Eschscholtz, 1829
Nomenclatural Authority: Nelson et al. (2008)
Digitized Records (genus-only): Santa Cruz (9 SBMNH; 6 UCRC; 1 UCSB)
Notes. Twenty-four species of Anthaxia have been reported from California (Nelson et al., 2008). All but one of the undetermined specimens reported above belong to the A. aeneogaster species group, a taxonomic complex that needs revision (N. Woodley, 2021, personal communication).
Anthaxia (Melanthaxia) aeneogaster Gory & Laporte, 1839
Nomenclatural Authority: Nelson et al. (2008)
Literature Records: none
Digitized Records: Santa Cruz (4 SBMNH)
Range: Also known from mainland (Nelson et al., 2008).
Buprestini
Notes. Three genera and 19 species of Buprestini are known to occur in California (Nelson et al., 2008).
Buprestis Linnaeus, 1758
Nomenclatural Authority: Nelson et al. (2008)
Notes. Fourteen species of Buprestis have been reported from California (Nelson et al., 2008). The genus was revised for North America by Helfer (1941).
Buprestis (Cypriacis) aurulenta Linnaeus, 1767
Nomenclatural Authority: Nelson et al. (2008)
Literature Records: Santa Cruz (Cockerell, 1940: 286)
Digitized Records: Santa Cruz (6 CASC; 6 SBMNH)
Range: Also known from mainland (Nelson et al., 2008).
Chrysobothrini
Notes. Three genera and 68 species of Chrysobothrini are known to occur in California (Nelson et al., 2008; M. L. Gimmel, 2022, unpublished data).
Chrysobothris Eschscholtz, 1829
Nomenclatural Authority: Nelson et al. (2008)
Notes. Sixty-five species of Chrysobothris have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Chrysobothris mali Horn, 1886
Nomenclatural Authority: Nelson et al. (2008)
Literature Records: none
Digitized Records: Santa Catalina (1 CASC), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Nelson et al., 2008).
Melanophilini
Notes. Four genera and 13 species of Melanophilini have been recorded from California (Nelson et al., 2008).
Melanophila Eschscholtz, 1829
Nomenclatural Authority: Nelson et al. (2008)
Notes. Four species of Melanophila have been recorded from California (Nelson et al., 2008). The genus was revised for North America by Sloop (1937).
Melanophila consputa LeConte, 1857
Nomenclatural Authority: Nelson et al. (2008)
Literature Records: none
Digitized Records: Santa Catalina (4 iNat)
Range: Also known from mainland (Nelson et al., 2008).
Polycestinae: Acmaeoderini
Notes. Five tribes, eight genera, and 119 species of Polycestinae are known to occur in California, of which four genera and 102 species belong to Acmaeoderini (Nelson et al., 2008).
Acmaeodera Eschscholtz, 1829
Nomenclatural Authority: Nelson et al. (2008)
Digitized Records (genus-only): Santa Cruz (4 UCSB)
Notes. Cockerell (1940: 286) relayed information from H.C. Fall that Acmaeodera connexa LeConte, 1859 (which he misspelled as “A. convexa”) was erroneously recorded from Santa Rosa (Fall, 1897: 237). A total of 78 species of Acmaeodera has been recorded from California (Nelson et al., 2008).
Acmaeodera (Acmaeodera) hepburnii LeConte, 1860
Nomenclatural Authority: Nelson et al. (2008)
Literature Records: Santa Catalina (Bellamy, 1982: 359), Santa Cruz (Nelson, 1962: 56; Bellamy, 1982: 359), Santa Rosa (Fall, 1897: 237; Cockerell, 1940: 286)
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (41 CASC; 10 LACM; 29 SBMNH; 6 UCSB; 1 iNat), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Nelson, 1962; Nelson et al., 2008).
Notes. Often misspelled Acmaeodera hepburni.
Acmaeodera (Acmaeodera) mariposa Horn, 1878
Nomenclatural Authority: Nelson et al. (2008)
Literature Records: none
Digitized Records: Santa Cruz (3 SBMNH)
Range: Also known from mainland (Nelson et al., 2008).
Notes. The subspecies occurring on the islands is A. m. dohrni Horn, 1878 (N. Woodley, 2021, personal communication).
Acmaeodera (Acmaeodera) prorsa Fall, 1899
Nomenclatural Authority: Nelson et al. (2008)
Literature Records: Santa Cruz (Bellamy, 1982: 359)
Digitized Records: Santa Catalina (1 CASC), Santa Cruz (1 CASC; 1 SBMNH; 1 UCSB)
Range: Also known from mainland (Bellamy, 1982; Nelson et al., 2008).
DRYOPOIDEA
Dryopidae
Notes. Three genera and five species of Dryopidae are known to occur in California (Shepard, 1993). Shepard (1993) provided the most recent published checklist of California Dryopidae. Brown (1972), although outdated, remains the best identification guide for North America.
Postelichus Nelson, 1989
Nomenclatural Authority: Shepard (1993)
Notes. Two species of Postelichus have been recorded from California (Shepard, 1993; Barr & Shepard, 2022). A key to the species of Postelichus was provided by Barr & Shepard (2022).
Postelichus productus (LeConte, 1852)
Nomenclatural Authority: Barr & Shepard (2022)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (Brown, 1972; Barr & Shepard, 2022).
Notes. Fall (1897) reported this species as Dryops productus.
Elmidae
Notes. Fourteen genera and 24 species of Elmidae are known to occur in California (M. L. Gimmel, 2022, unpublished data). Shepard (1993) provided the most recent published checklist of California Elmidae. Brown (1972), although outdated, remains the best identification guide for North America.
Ordobrevia Sanderson, 1953
Nomenclatural Authority: Shepard (1993)
Notes. One species of Ordobrevia has been recorded from California (Shepard, 1993).
Ordobrevia nubifera (Fall, 1901)
Nomenclatural Authority: Shepard (1993)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: none
Range: Also known from mainland (Brown, 1972).
Heteroceridae
Notes. Three genera and 11 species of Heteroceridae are known to occur in California (M. L. Gimmel, 2022, unpublished data). Shepard (1993) provided the most recent published checklist of California Heteroceridae. Pacheco (1964) monographed the species for the New World, though most of his genera were not recognized by King, Starr & Lago (2011).
Heterocerus Fabricius, 1792
Nomenclatural Authority: Shepard (1993)
Digitized Records (genus-only): San Clemente (11 SBMNH), San Nicolas (5 SBMNH), Santa Catalina (7 SBMNH), Santa Cruz (7 SBMNH), Santa Rosa (16 SBMNH)
Notes. Eight species of Heterocerus have been recorded from California (Shepard, 1993).
Heterocerus mexicanus Sharp, 1882
Nomenclatural Authority: Shepard (1993), King, Starr & Lago (2011)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: Santa Cruz (3 SBMNH)
Range: Also known from mainland (Brown, 1972).
Notes. This species has been known in the literature (Pacheco, 1964; Shepard, 1993) as Dampfius mexicanus. However, the genus Dampfius Pacheco, 1964 was synonymized with Heterocerus by King, Starr & Lago (2011).
Limnichidae, NEW FAMILY RECORD
Notes. Six genera and 13 species of Limnichidae are known to occur in California (Shepard, 1993). Shepard (1993) provided the most recent published checklist of California Limnichidae.
Limnichites Casey, 1889
Nomenclatural Authority: Wooldridge (1977)
Notes. Three species of Limnichites have been recorded from California (Shepard, 1993). Wooldridge (1977) provided the most recent revision of this genus.
Limnichites nebulosus (LeConte, 1879)
Nomenclatural Authority: Wooldridge (1977)
Literature Records: none
Digitized Records: Santa Cruz (5 SBMNH)
Range: Also known from mainland (Wooldridge, 1977).
ELATEROIDEA
Cantharidae
Notes. Four subfamilies, 13 genera, and 157 species of Cantharidae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Cantharinae
Notes. Two tribes, seven genera, and 51 species of Cantharinae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Cantharini
Notes. Five genera and 16 species of Cantharini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Cultellunguis McKey-Fender, 1950
Nomenclatural Authority: Ramsdale (2002)
Digitized Records (genus-only): Santa Catalina (3 SBMNH), Santa Cruz (8 SBMNH)
Notes. This genus contains nine species restricted to the Pacific coast of North America (Ramsdale, 2002), all of which occur in California (McKey-Fender, 1950).
Cultellunguis americanus (Pic, 1906)
Nomenclatural Authority: McKey-Fender (1950), Ramsdale (2002)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (McKey-Fender, 1950).
Notes. Listed as “Telephorus notatus Mann. var.” by Fall (1897). Telephorus notatus Mannerheim, 1843 is an unavailable homonym which was replaced by C. americanus (see McKey-Fender, 1950).
Cultellunguis hatchi (McKey-Fender, 1950)
Nomenclatural Authority: McKey-Fender (1950), Fender (1968), Ramsdale (2002)
Literature Records: Santa Catalina (McKey-Fender, 1950: 65; Fender, 1968: 301; Miller, 1985a: 21), Santa Cruz (McKey-Fender, 1950: 65; Naughton et al., 2014: 303)
Digitized Records: Santa Cruz (13 SBMNH)
Range: Also known from mainland (McKey-Fender, 1950; Fender, 1968).
Notes. This species, as Cantharis (Cultellunguis) hatchi McKey-Fender, was originally recorded from both Santa Catalina and Santa Cruz by McKey-Fender (1950); however, a Santa Catalina subspecies, as Cantharis (Cultellunguis) hatchi dorothyae Fender, 1968, was later split off by Fender (1968). This subspecies, considered to be endemic to Santa Catalina, is now known as Cultellunguis h. dorothyae (Fender, 1968), while the nominate subspecies, Cultellunguis h. hatchi (McKey-Fender, 1950), occurs on Santa Cruz Island (Fender, 1968; Miller, 1985a). The latter subspecies also occurs on the mainland (Fender, 1968).
Pacificanthia Kazantsev, 2002
Nomenclatural Authority: Kazantsev (2002)
Notes. One species of the genus Pacificanthia occurs in California (Kazantsev, 2002). Kazantsev (2002) provided a generic description and key to species for the genus.
Pacificanthia consors (LeConte, 1851)
Nomenclatural Authority: Kazantsev (2002)
Literature Records: Santa Cruz (Fall & Davis, 1934: 144)
Digitized Records: Santa Catalina (32 LACM; 3 SBMNH; 1 iNat), Santa Cruz (11 LACM; 15 SBMNH; 8 UCSB), Santa Rosa (1 LACM)
Range: Also known from mainland (Kazantsev, 2002).
Notes. Fall & Davis (1934) recorded this species as Cantharis consors.
Podabrini
Notes. The two California genera of tribe Podabrini, Dichelotarsus Motschulsky, 1860 and Podabrus Westwood, 1838, while both valid, have not had their species properly assigned yet (Ramsdale, 2002). A collective total of 35 species belonging to both genera is known to occur in California (M. L. Gimmel, 2022, unpublished data).
Podabrus pruinosus LeConte, 1851
Nomenclatural Authority: Fall (1927), Fender (1949)
Literature Records: none
Digitized Records: Santa Cruz (13 SBMNH)
Range: Also known from mainland (Fall, 1927).
Notes. This species was included in Fender’s (1949) “Group I”, and therefore will probably stay in the genus Podabrus (see Ramsdale, 2002). The subspecies occurring on the islands is P. p. pruinosus LeConte, 1851.
Podabrus undetermined species
Nomenclatural Authority: Fender (1948), Fender (1949)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Notes. This species keyed to Fender’s (1949) “Group VIII”, and therefore may end up in the genus Dichelotarsus (see Ramsdale, 2002). However, it did not fit any of the species concepts in Fender’s (1948) revision of that group (M. L. Gimmel, 2021, personal observation).
Malthininae
Notes. Two genera and 60 species of Malthininae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Frostia Fender, 1951
Nomenclatural Authority: Fender (1951)
Notes. Four species of Frostia have been recorded from California (Fender, 1951). This genus was described and revised by Fender (1951).
Frostia laticollis (LeConte, 1866)
Nomenclatural Authority: Fender (1951)
Literature Records: Santa Cruz (LeConte, 1861: 351; Fall, 1897: 237; Fall, 1919: 35; Fall & Davis, 1934: 144; Naughton et al., 2014: 303)
Digitized Records: Santa Cruz (20 SBMNH)
Range: Also known from mainland (Fall, 1919; Fender, 1951).
Notes. LeConte (1861) recorded this species as Malthodes transversus LeConte, 1861, but the name was subsequently corrected to Malthodes laticollis by LeConte (1866b: 53). At that time the species was considered to be endemic to Santa Cruz Island. Fall (1897, 1919) and Fall & Davis (1934) recorded this species as Malthodes laticollis. Fender (1951: 524) transferred this species to Frostia. Naughton et al. (2014) reported two specimens of the genus Frostia from Santa Cruz Island; one voucher (in SBMNH) was identified by MLG as F. laticollis.
Silinae
Notes. Three genera and 43 species of Silinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Silis Charpentier, 1825
Nomenclatural Authority: Ramsdale (2002)
Digitized Records (genus-only): Santa Cruz (1 SBMNH)
Notes. Forty species of Silis are known to occur in California (M. L. Gimmel, 2022, unpublished data). Green (1966) revised the species of Silis known from North America at the time. The specimen the above record is based on is female, and therefore not identifiable to species (Green, 1966). However, it is likely that it belongs to S. carmelita, cited below.
Silis carmelita Green, 1966
Nomenclatural Authority: Green (1966)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Green, 1966).
Notes. The single specimen is a male, which was dissected for examination of the genitalia for identification (M. L. Gimmel, 2021, personal observation).
Elateridae
Notes. Eight subfamilies, 67 genera, and 362 species of Elateridae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Agrypninae: Oophorini
Notes. Three tribes, nine genera, and 23 species of Agrypninae are known to occur in California, of which four genera and 11 species belong to Oophorini (M. L. Gimmel, 2022, unpublished data).
Heteroderes Latreille, 1834
Nomenclatural Authority: Kundrata et al. (2019)
Notes. Two species of Heteroderes are known from California (M. L. Gimmel, 2022, unpublished data).
Heteroderes amplicollis (Gyllenhal, 1808)
Nomenclatural Authority: Stone (1975)
Literature Records: none
Digitized Records: Santa Rosa (1 LACM)
Range: Also known from mainland (Stone, 1975).
Notes. This species is a destructive pest introduced from South America (Stone, 1975). It has often been included in Conoderus Eschscholtz, 1829 in the literature (including by Stone (1975)), but is now commonly treated as a Heteroderes and will soon be moved to a new genus (P. Johnson, 2021, personal communication).
Cardiophorinae
Notes. Five genera and 52 species of Cardiophorinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Cardiophorus Eschscholtz, 1829
Nomenclatural Authority: Douglas (2003)
Digitized Records (genus-only): Santa Cruz (1 SBMNH), Santa Rosa (5 SBMNH)
Notes. One of the Santa Rosa Island specimens above was identified by Hume Douglas during 2006 as “Cardiophorus tenebrosus group”. The other specimens from both islands are similar in appearance and likely represent the same species, which may be conspecific with the specimens identified as C. tenebrosus below. However, this is a highly diverse genus with 28 described species of Cardiophorus recorded from California in the literature (M. L. Gimmel, 2022, unpublished data), even after the recent splitting off of Paracardiophorus Schwarz, 1895 by Douglas (2017) (13 species recorded from California; M. L. Gimmel, 2022, unpublished data).
Cardiophorus tenebrosus LeConte, 1853
Nomenclatural Authority: Douglas (2003)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (M. L. Gimmel, 2022, unpublished data).
Notes. A specimen in the H.C. Fall collection in the Museum of Comparative Zoology, Harvard University from San Clemente Island was questionably identified as this species (S. Miller, 2022, personal communication). This record needs verification.
Horistonotus Candèze, 1860
Nomenclatural Authority: Wells (2000)
Notes. Five species of this genus have been recorded from California (M. L. Gimmel, 2022, unpublished data). Wells (2000) provided a key to all North American species.
Horistonotus inanus (LeConte, 1853)
Nomenclatural Authority: Wells (2000)
Literature Records: none
Digitized Records: Santa Catalina (10 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Wells, 2000).
Notes. This is a dimorphic species with regard to dorsal color pattern, with some specimens showing distinct light coloration at the base of the elytra. However, all examined specimens from the Channel Islands are not or very weakly bicolored.
Dendrometrinae; Dendrometrini
Notes. Two tribes, 23 genera, and 138 species of Dendrometrinae have been recorded from California, of which eight genera and 62 species belong to Dendrometrini (M. L. Gimmel, 2022, unpublished data).
Athous Eschscholtz, 1829
Nomenclatural Authority: Etzler (2020b)
Notes. Seventeen species of Athous have been recorded from California (Becker, 1979).
Athous axillaris Horn, 1871
Nomenclatural Authority: Becker (1979)
Literature Records: none
Digitized Records: Santa Cruz (17 SBMNH), Santa Rosa (7 SBMNH)
Range: Also known from mainland (Becker, 1979).
Athous nigropilis Motschulsky, 1859
Nomenclatural Authority: Becker (1979)
Literature Records: none
Digitized Records: Santa Catalina (3 LACM)
Range: Also known from mainland (Becker, 1979).
Athous rufiventris (Eschscholtz, 1822)
Nomenclatural Authority: Becker (1979)
Literature Records: none
Digitized Records: Santa Catalina (3 SBMNH), Santa Cruz (3 SBMNH)
Range: Also known from mainland (Becker, 1979).
Hemicrepidius Germar, 1839
Nomenclatural Authority: Etzler (2020b)
Notes. Twelve species of Hemicrepidius have been recorded from California (Etzler, 2020b).
Hemicrepidius californicus Becker, 1979
Nomenclatural Authority: Etzler (2020b)
Literature Records: none
Digitized Records: San Miguel (2 SBMNH), San Nicolas (4 LACM; 2 SBMNH)
Range: Also known from mainland (Etzler, 2020b).
Hemicrepidius tumescens (LeConte, 1861)
Nomenclatural Authority: Etzler (2020b)
Literature Records: Santa Cruz (LeConte, 1861: 348; Fall, 1897: 237; Fall, 1901: 114; Van Dyke, 1932: 444; Etzler, 2020b: 86, 87)
Range: Also known from mainland (Van Dyke, 1932; Etzler, 2020b).
Notes. Recorded by LeConte (1861) and Fall (1897, 1901) as Asaphes tumescens. At the time of LeConte (1861), this species was considered to be endemic to Santa Cruz Island.
Limonius Eschscholtz, 1829
Nomenclatural Authority: Etzler (2019)
Notes. Eleven species of Limonius are known to occur in California (M. L. Gimmel, 2022, unpublished data). While Al Dhafer (2009) revised the North American species of Limonius, Etzler (2019) reclassified the world species to various genera.
Limonius canus LeConte, 1853
Nomenclatural Authority: Al Dhafer (2009), Etzler (2019)
Literature Records: none
Digitized Records: San Clemente (1 LACM), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Al Dhafer, 2009).
Elaterinae
Notes. Five tribes, 17 genera, and 127 species of Elaterinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Agriotini
Notes. Four genera and 41 species of Agriotini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Dalopius Eschscholtz, 1829
Nomenclatural Authority: Brown (1934)
Digitized Records: Santa Catalina (1 LACM), Santa Cruz (1 LACM)
Notes. Twenty-four species of Dalopius have been recorded from California (M. L. Gimmel, 2022, unpublished data). This genus was revised for North America in a multi-part paper by Brown (1934).
Dalopius luteolus Brown, 1934
Nomenclatural Authority: Brown (1934)
Literature Records: none
Digitized Records: Santa Cruz (2 LACM)
Range: Also known from mainland (Brown, 1934).
Dalopius undetermined species
Literature Records: none
Digitized Records: Santa Cruz (17 SBMNH), Santa Rosa (1 SBMNH)
Notes. The rather uniform specimens above (SBMNH) were examined and they do not match the description of D. luteolus. One male was dissected and the median lobe does not match any of the illustrated species in Brown (1934), though it is fairly close to that of Dalopius partitus Brown, 1934. The species may be undescribed.
Ampedini
Notes. Four genera and 35 species of Ampedini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Ampedus Dejean, 1833
Nomenclatural Authority: Ramberg (1979)
Notes. Twenty-four species of Ampedus have been recorded from California (Ramberg, 1979). Ramberg (1979) revised the species for North America. Unfortunately, most of the taxonomic acts in this thesis work have not been validly published.
Ampedus longicornis (LeConte, 1884)
Nomenclatural Authority: Ramberg (1979)
Literature Records: Santa Catalina (Ramberg, 1979: 318)
Digitized Records: Santa Catalina (2 LACM; 2 SBMNH)
Range: Also known from mainland (Ramberg, 1979).
Ampedus rhodopus (LeConte, 1884)
Nomenclatural Authority: Ramberg (1979)
Literature Records: none
Digitized Records: Santa Rosa (3 SBMNH)
Range: Also known from mainland (Ramberg, 1979).
Anchastus LeConte, 1853
Nomenclatural Authority: Johnson (2002)
Notes. Six species of Anchastus are known to occur in California (M. L. Gimmel, 2022, unpublished data). Van Dyke (1932) provided a key to the North American species of Anchastus.
Anchastus cinereipennis (Eschscholtz, 1829)
Nomenclatural Authority: Van Dyke (1932)
Literature Records: San Nicolas (Miller & Miller, 1985: 126), Santa Barbara (Miller & Miller, 1985: 126)
Digitized Records: San Clemente (3 SBMNH), San Nicolas (6 SBMNH), Santa Catalina (1 SBMNH), Santa Rosa (1 iNat)
Range: Also known from mainland (M. L. Gimmel, 2022, unpublished data).
Melanotus Eschscholtz, 1829
Nomenclatural Authority: Johnson (2002)
Notes. Four species of Melanotus have been recorded from California (Quate & Thompson, 1967). The genus was revised for North America by Quate & Thompson (1967).
Melanotus longulus (LeConte, 1853)
Nomenclatural Authority: Quate & Thompson (1967)
Literature Records: Santa Catalina (Fall, 1897: 237; Fall, 1901: 111; Quate & Thompson, 1967: 61)
Digitized Records: Santa Catalina (1 LACM; 12 SBMNH; 6 TAMU), Santa Cruz (10 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Quate & Thompson, 1967).
Notes. The nominate subspecies, M. l. longulus (LeConte, 1853), is the only subspecies of M. longulus known from south of the Tehachapi Mountains in southern California (Quate & Thompson, 1967). This species was recorded as Melanotus variolatus LeConte, 1861 by Fall (1897, 1901), which was synonymized with M. longulus by Quate & Thompson (1967).
Aplastini
Notes. Three genera and 28 species of Aplastini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Euthysanius LeConte, 1853
Nomenclatural Authority: Johnson (2002)
Notes. Seven species of Euthysanius are known to occur in California (Johnson, 2002). Van Dyke (1932) provided a key to the North American species of Euthysanius.
Euthysanius lautus LeConte, 1853
Nomenclatural Authority: Van Dyke (1932)
Literature Records: none
Digitized Records: Santa Cruz (9 LACM), Santa Rosa (1 LACM; 1 SBMNH)
Range: Also known from mainland (Van Dyke, 1932).
Octinodes Candèze, 1863
Nomenclatural Authority: Johnson (2002)
Notes. Nine species of Octinodes are known to occur in California (Johnson, 2002). Van Dyke (1932) provided a key to separate some North American species of Octinodes (as Plastocerus LeConte, 1853).
Octinodes frater (LeConte, 1859)
Nomenclatural Authority: Van Dyke (1932)
Literature Records: none
Digitized Records: Santa Cruz (1 LACM; 1 SBMNH)
Range: Also known from mainland (Van Dyke, 1932).
Elaterini
Notes. Five genera and 12 species of Elaterini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Elater Linnaeus, 1758
Nomenclatural Authority: Johnson (2002)
Notes. Four species of Elater are known from California (M. L. Gimmel, 2022, unpublished data). The genus was revised for North America by Roache (1961).
Elater lecontei (Horn, 1871)
Nomenclatural Authority: Roache (1961)
Literature Records: none
Digitized Records: Santa Cruz (3 SBMNH)
Range: Also known from mainland (Roache, 1961).
Negastriinae
Notes. Five genera and 11 species of Negastriinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Paradonus Stibick, 1971
Nomenclatural Authority: Etzler (2020a)
Notes. Three species of Paradonus have been recorded from California (Etzler, 2020a).
Paradonus inops (LeConte, 1853)
Nomenclatural Authority: Etzler (2020a)
Literature Records: none
Digitized Records: Santa Cruz (20 SBMNH)
Range: Also known from mainland (Etzler, 2020a).
Oxynopterinae
Notes. Only two genera and two species of Oxynopterinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Melanactes LeConte, 1853
Nomenclatural Authority: Mathieu (1961)
Notes. Only one species of the widespread genus Melanactes has been recorded from California (Mathieu, 1961). The genus was revised by Mathieu (1961).
Melanactes densus LeConte, 1853
Nomenclatural Authority: Mathieu (1961)
Literature Records: none
Digitized Records: Santa Catalina (2 USNM)
Range: Also known from mainland (Mathieu, 1961).
Eucnemidae
Notes. Six subfamilies, 14 genera, and 22 species of Eucnemidae have been recorded from California (Muona, 2000; M. L. Gimmel, 2022, unpublished data). The North American fauna was revised by Muona (2000).
Macraulacinae
Notes. Three genera and seven species of Macraulacinae have been recorded from California (Muona, 2000).
Asiocnemis Mamaev, 1976
Nomenclatural Authority: Muona (2000).
Notes. Five species of Asiocnemis have been reported from California (Muona, 2000).
Asiocnemis hospitalis (Blanchard, 1904)
Nomenclatural Authority: Muona (2000)
Literature Records: Santa Rosa (Muona, 2000: 81)
Digitized Records: Santa Rosa (1 LACM)
Range: Also known from mainland (Muona, 2000).
Lampyridae, NEW FAMILY RECORD
Notes. Three subfamilies, nine genera, and 22 species of Lampyridae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Lampyrinae
Notes. Six genera and 16 species of Lampyrinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Pyropyga Motschulsky, 1852
Nomenclatural Authority: Green (1961)
Notes. One species of Pyropyga has been recorded from California (Green, 1961).
Pyropyga nigricans (Say, 1823)
Nomenclatural Authority: Green (1961)
Literature Records: none
Digitized Records: Santa Cruz (1 UCSB), Santa Rosa (2 LACM)
Range: Also known from mainland (Green, 1961).
Pterotinae
Notes. One genus and two species of Pterotinae have been recorded from California (Chemsak, 1978).
Pterotus LeConte, 1859
Nomenclatural Authority: Chemsak (1978)
Notes. Two species of Pterotus are known, both of them occurring in California (Chemsak, 1978).
Pterotus obscuripennis LeConte, 1859
Nomenclatural Authority: Chemsak (1978)
Literature Records: none
Digitized Records: Santa Catalina (1 LACM; 3 SBMNH)
Range: Also known from mainland (Chemsak, 1978).
Phengodidae, NEW FAMILY RECORD
Notes. Two subfamilies, four genera, and seven species of Phengodidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Phengodinae
Notes. Two genera and four species of Phengodinae have been recorded from California (M. L. Gimmel, 2022, unpublished data)
Zarhipis LeConte, 1880
Nomenclatural Authority: Linsdale (1964)
Notes. Three species of Zarhipis have been recorded from California (Linsdale, 1964). The genus was revised by Linsdale (1964).
Zarhipis integripennis (LeConte, 1874)
Nomenclatural Authority: Linsdale (1964)
Literature Records: none
Digitized Records: Santa Catalina (1 LACM)
Range: Also known from mainland (Linsdale, 1964).
Throscidae, NEW FAMILY RECORD
Notes. Three genera and five species of Throscidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Trixagus Kugelann, 1794
Nomenclatural Authority: Yensen (1975)
Notes. Three species of Trixagus have been recorded from California (Yensen, 1975).
Trixagus sericeus (LeConte, 1868)
Nomenclatural Authority: Yensen (1975)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Yensen, 1975).
HISTEROIDEA
Histeridae
Notes. Seven subfamilies, 39 genera, and 141 species of Histeridae are known to occur in California (M. L. Gimmel, 2022, unpublished data). Although no histerid taxa below are recorded for Anacapa Island, LACM has undetermined material from that island.
Abraeinae
Notes. Three tribes, six genera, and 13 species of Abraeinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Abraeini
Notes. Two genera and five species of Abraeini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Plegaderus Erichson, 1834
Nomenclatural Authority: Mazur (2011)
Notes. Four species of Plegaderus are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Plegaderus undetermined species
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Acritini
Notes. Three genera and four species of Acritini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Halacritus Schmidt, 1893
Nomenclatural Authority: Mazur (2011)
Notes. One species of Halacritus has been recorded from California (Mazur, 2011).
Halacritus maritimus (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: none
Digitized Records: San Clemente (1 SBMNH), San Nicolas (2 SBMNH)
Range: Also known from mainland.
Dendrophilinae
Notes. Four tribes, five genera, and 20 species of Dendrophilinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Bacaniini
Notes. One genus and two species of Bacaniini have been recorded from California (Mazur, 2011).
Bacanius LeConte, 1853
Nomenclatural Authority: Mazur (2011)
Notes. Two species of Bacanius have been recorded from California (Mazur, 2011).
Bacanius undetermined species
Literature Records: Santa Catalina (Caterino & Chandler, 2010: 191)
Digitized Records: Santa Catalina (1 SBMNH)
Notes. The male specimen (SBMNH) from Santa Catalina Island does not appear to match either of the two species previously recorded for California, Bacanius (Gomyister) acuminatus Casey, 1893 or Bacanius (s.str.) globulinus Casey, 1893. It may represent an undescribed species.
Paromalini
Notes. Two genera and 12 species of Paromalini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Carcinops Marseul, 1855
Nomenclatural Authority: Mazur (2011); Reese & Swanson (2017)
Notes. Ten species of Carcinops are known to occur in California (M. L. Gimmel, 2022, unpublished data). This genus was reported from Santa Barbara Island by Miller & Miller (1985: 123) from stems of Coreopsis gigantea (Kellogg) H.M. Hall (Asteraceae).
Carcinops opuntiae (LeConte, 1851)
Nomenclatural Authority: Reese & Swanson (2017)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (Reese & Swanson, 2017).
Histerinae
Notes. Four tribes, nine genera, and 30 species of Histerinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Histerini
Notes. Five genera and 20 species of Histerini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Margarinotus Marseul, 1854
Nomenclatural Authority: Mazur (2011)
Notes. Eleven species of Margarinotus have been recorded from California (Caterino, 2010), belonging to two subgenera, Paralister Bickhardt, 1917 and Ptomister Houlbert & Monnot, 1922. The California species were revised by Caterino (2010).
Margarinotus (Ptomister) sexstriatus (LeConte, 1851)
Nomenclatural Authority: Caterino (2010)
Literature Records: Santa Cruz (Caterino, 2010: 10), Santa Rosa (Caterino, 2010: 10)
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Caterino, 2010).
Hololeptini
Notes. Two genera and six species of Hololeptini have been recorded from California (Mazur, 2011).
Hololepta Paykull, 1811
Nomenclatural Authority: Mazur (2011)
Notes. Five species of Hololepta have been recorded from California (Mazur, 2011) in two subgenera, Hololepta (s.str.) and Leionota Marseul, 1853.
Hololepta (Leionota) vicina LeConte, 1851
Nomenclatural Authority: Mazur (2011)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (Fall, 1901).
Iliotona Carnochan, 1917
Nomenclatural Authority: Mazur (2011)
Notes. One species of Iliotona has been recorded from California (Mazur, 2011).
Iliotona cacti (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: none
Digitized Records: San Miguel (1 SBMNH)
Range: Also known from mainland (Fall, 1901).
Saprininae
Notes. Eleven genera and 53 species of Saprininae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Aphelosternus Wenzel, 1962
Nomenclatural Authority: Mazur (2011)
Notes. Only one species is contained in the genus Aphelosternus (Mazur, 2011).
Aphelosternus interstitialis (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: Santa Catalina (Fall, 1897: 237; Fall, 1901: 96)
Digitized Records: none
Range: Also known from mainland (Fall, 1901).
Notes. This species was recorded as Saprinus interstitialis by Fall (1897, 1901).
Euspilotus Lewis, 1907
Nomenclatural Authority: Mazur (2011)
Digitized Records (genus-only): Santa Catalina (2 SBMNH), Santa Rosa (1 SBMNH)
Notes. Eleven species of Euspilotus are known to occur in California (M. L. Gimmel, 2022, unpublished data), distributed among three subgenera, Hesperosaprinus Wenzel, 1962, Neosaprinus Bickhardt, 1909, and Platysaprinus Bickhardt, 1916.
Euspilotus (Hesperosaprinus) scissus (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: none
Digitized Records: San Miguel (9 SBMNH), San Nicolas (8 SBMNH), Santa Cruz (5 SBMNH), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Mazur, 2011).
Euspilotus (Hesperosaprinus) species near laridus (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Unknown.
Notes. This species was recorded as “Saprinus sp. near laridus” by Fall (1897).
Geomysaprinus Ross, 1940
Nomenclatural Authority: Mazur (2011)
Notes. Six species of Geomysaprinus have been recorded from California (Mazur, 2011). All of these belong to the subgenus Priscosaprinus Wenzel, 1962 (Mazur, 2011).
Geomysaprinus undetermined species
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH), Santa Rosa (1 SBMNH)
Hypocaccus Thomson, 1867
Nomenclatural Authority: Mazur (2011)
Notes. Seven species of Hypocaccus are known to occur in California (M. L. Gimmel, 2022, unpublished data), distributed among two subgenera, Baeckmanniolus Reichardt, 1926 and Hypocaccus (s.str.).
Hypocaccus (Baeckmanniolus) gaudens (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: none
Digitized Records: San Miguel (4 SBMNH), San Nicolas (1 LACM; 1 SBMNH), Santa Catalina (2 SBMNH), Santa Cruz (3 SBMNH), Santa Rosa (10 SBMNH)
Range: Also known from mainland (Mazur, 2011).
Hypocaccus (Baeckmanniolus) serrulatus (LeConte, 1851)
Nomenclatural Authority: [none]
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland.
Notes. This name is missing from the catalog of Mazur (2011).
Hypocaccus (Hypocaccus) bigemmeus (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: none
Digitized Records: San Clemente (1 SBMNH), San Miguel (10 SBMNH), San Nicolas (7 SBMNH), Santa Cruz (8 SBMNH), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Mazur, 2011).
Hypocaccus (Hypocaccus) lucidulus (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: San Clemente (Caterino, Chatzimanolis & Richmond, 2015: 278), San Miguel (Caterino, Chatzimanolis & Richmond, 2015: 278), San Nicolas (Cockerell, 1940: 285; Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Cruz (Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Rosa (Caterino, Chatzimanolis & Richmond, 2015: 278)
Digitized Records: San Clemente (11 SBMNH), San Miguel (19 SBMNH), San Nicolas (12 SBMNH), Santa Cruz (21 SBMNH), Santa Rosa (13 SBMNH)
Range: Also known from mainland (Cockerell, 1940; Mazur, 2011).
Notes. Cockerell (1940) reported this species as Saprinus lucidulus.
Neopachylopus Reichardt, 1926
Nomenclatural Authority: Mazur (2011)
Notes. Two species of Neopachylopus have been recorded from California (Mazur, 2011).
Neopachylopus sulcifrons (Mannerheim, 1843)
Nomenclatural Authority: Mazur (2011)
Literature Records: San Nicolas (Cockerell, 1940: 285)
Digitized Records: San Clemente (1 SBMNH), San Miguel (5 SBMNH), San Nicolas (1 SBMNH), Santa Catalina (1 SBMNH), Santa Cruz (14 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Cockerell, 1940; Mazur, 2011).
Notes. Cockerell (1940) reported this species as Saprinus sulcifrons.
Saprinus Erichson, 1834
Nomenclatural Authority: Mazur (2011)
Digitized Records (genus-only): Santa Rosa (2 LACM)
Notes. Four species of Saprinus have been recorded from California, all of them belonging to the nominate subgenus (M. L. Gimmel, 2022, unpublished data).
Saprinus (Saprinus) lugens Erichson, 1834
Nomenclatural Authority: Mazur (2011)
Literature Records: San Clemente (Fall, 1897: 237; Miller & Miller, 1985: 123), San Nicolas (Fall, 1897: 237; Miller & Miller, 1985: 123), Santa Barbara (Fall, 1897: 237; Miller & Miller, 1985: 123), Santa Cruz (Fall & Davis, 1934: 144; Miller & Miller, 1985: 123), Santa Rosa (Fall, 1897: 237; Miller & Miller, 1985: 123)
Digitized Records: San Miguel (4 SBMNH), San Nicolas (1 SBMNH), Santa Barbara (1 SBMNH), Santa Catalina (3 SBMNH; 1 TAMU), Santa Cruz (7 SBMNH), Santa Rosa (26 SBMNH)
Range: Also known from mainland (Mazur, 2011).
Saprinus (Saprinus) oregonensis LeConte, 1844
Nomenclatural Authority: Mazur (2011)
Literature Records: Santa Cruz (Fall & Davis, 1934: 144)
Digitized Records: none
Range: Also known from mainland (Mazur, 2011).
Xerosaprinus Wenzel, 1962
Nomenclatural Authority: Mazur (2011)
Digitized Records (genus-only): Santa Rosa (2 LACM)
Notes. Thirteen species of Xerosaprinus are known from California (M. L. Gimmel, 2022, unpublished data), belonging to two subgenera, Vastosaprinus Wenzel, 1962 and Xerosaprinus (s.str.).
Xerosaprinus (Xerosaprinus) fimbriatus (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland.
Notes. This species was recorded as Saprinus fimbriatus by Fall (1897).
Xerosaprinus (Xerosaprinus) lubricus (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: San Clemente (Fall, 1897: 237), Santa Catalina (Fall, 1897: 237), Santa Cruz (Fall & Davis, 1934: 144), Santa Rosa (Fall, 1897: 237)
Digitized Records: Santa Catalina (3 SBMNH), Santa Cruz (5 SBMNH; 3 UCSB), Santa Rosa (13 SBMNH)
Range: Also known from mainland (Mazur, 2011).
Notes. This species was recorded as Saprinus lubricus by Fall (1897) and Fall & Davis (1934).
Xerosaprinus (Xerosaprinus) vitiosus (LeConte, 1851)
Nomenclatural Authority: Mazur (2011)
Literature Records: Santa Catalina (Seavey, 1892: 262; Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (Mazur, 2011).
Notes. This species was recorded as Saprinus vitiosus by Seavey (1892) and Fall (1897).
HYDROPHILOIDEA
Helophoridae, NEW FAMILY RECORD
Notes. The family Helophoridae contains a single genus, Helophorus. The species (as Hydrophilidae subfamily Helophorinae) were revised for North America by Smetana (1985).
Helophorus Fabricius, 1775
Nomenclatural Authority: Hansen (1999)
Notes. Nineteen species of Helophorus have been recorded from California, all of them belonging to the subgenus Rhopalohelophorus Kuwert, 1886 (Hansen, 1999).
Helophorus (Rhopalohelophorus) linearis LeConte, 1855
Nomenclatural Authority: Smetana (1985)
Literature Records: none
Digitized Records: San Clemente (15 SBMNH)
Range: Also known from mainland (Smetana, 1985).
Hydrophilidae
Notes. Five subfamilies, 21 genera, and 117 species of Hydrophilidae are known to occur in California (Hansen, 1999; M. L. Gimmel, 2022, unpublished data).
Acidocerinae
Notes. One species of Acidocerinae has been recorded from California (Hansen, 1999).
Helochares Mulsant, 1844
Nomenclatural Authority: Hansen (1999)
Notes. One species of Helochares has been recorded from California (Hansen, 1999), belonging to the subgenus Hydrobaticus MacLeay, 1871.
Helochares (Hydrobaticus) normatus (LeConte, 1861)
Nomenclatural Authority: Hansen (1999); Short & Girón (2018)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: Santa Cruz (16 SBMNH), Santa Rosa (12 SBMNH)
Range: Also known from mainland (Hansen, 1999; Short & Girón, 2018).
Chaetarthriinae
Notes. Two tribes, three genera, and 20 species of Chaetarthriinae are known to occur in California (Hansen, 1999; M. L. Gimmel, 2022, unpublished data).
Anacaenini
Notes. Two genera and nine species of Anacaenini are known to occur in California (Hansen, 1999; M. L. Gimmel, 2022, unpublished data).
Anacaena Thomson, 1859
Nomenclatural Authority: Hansen (1999)
Notes. Three species of Anacaena are known from California (M. L. Gimmel, 2022, unpublished data).
Anacaena signaticollis (Fall, 1924)
Nomenclatural Authority: Hansen (1999)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250; Short & Caterino, 2009: 405)
Digitized Records: San Miguel (3 SBMNH), Santa Cruz (50 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Hansen, 1999; Short & Caterino, 2009).
Chaetarthriini
Notes. One genus and 11 species of Chaetarthriini have been recorded from California (Hansen, 1999).
Chaetarthria Stephens, 1835
Nomenclatural Authority: Hansen (1999)
Digitized Records (genus-only): Santa Cruz (37 SBMNH), Santa Rosa (1 SBMNH)
Notes. Eleven species of Chaetarthria have been recorded from California (Hansen, 1999). The species of this genus were revised for the New World by Miller (1974).
Chaetarthria hespera Miller, 1974
Nomenclatural Authority: Miller (1974), Hansen (1999)
Literature Records: Santa Catalina (Fall, 1897: 236; Miller, 1974: 43)
Digitized Records: Santa Catalina (17 SBMNH), Santa Cruz (2 SBMNH)
Range: Also known from mainland (Miller, 1974; Hansen, 1999).
Notes. Based on material examined in Miller (1974) (who accidentally indicated Santa Catalina as being in Orange Co.), Fall’s (1897) record of C. nigrella apparently refers to this species.
Chaetarthria nigrella (LeConte, 1861)
Nomenclatural Authority: Miller (1974), Hansen (1999)
Literature Records: none
Digitized Records: Santa Cruz (4 SBMNH), Santa Rosa (5 SBMNH)
Range: Also known from mainland (Miller, 1974; Hansen, 1999).
Notes. Fall’s (1897) island record of C. nigrella refers to C. hespera (see that species).
Chaetarthria punctulata Sharp, 1882
Nomenclatural Authority: Miller (1974), Hansen (1999)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Miller, 1974; Hansen, 1999).
Chaetarthria pusilla Sharp, 1882
Nomenclatural Authority: Miller (1974), Hansen (1999)
Literature Records: none
Digitized Records: Santa Cruz (12 SBMNH)
Range: Also known from mainland (Miller, 1974; Hansen, 1999).
Enochrinae
Notes. Two genera and 21 species of Enochrinae have been recorded from California (Hansen, 1999; M. L. Gimmel, 2022, unpublished data).
Cymbiodyta Bedel, 1881
Nomenclatural Authority: Hansen (1999)
Notes. Nine species of Cymbiodyta have been recorded from California (Hansen, 1999). The genus was revised by Smetana (1974).
Cymbiodyta columbiana Leech, 1948
Nomenclatural Authority: Smetana (1974), Hansen (1999)
Literature Records: none
Digitized Records: Santa Cruz (5 SBMNH)
Range: Also known from mainland (Smetana, 1974; Hansen, 1999).
Notes. This species is morphologically extremely similar to C. dorsalis (see Smetana, 1974: 44). Consequently, Channel Island records of these two species should be interpreted with caution.
Cymbiodyta dorsalis (Motschulsky, 1859)
Nomenclatural Authority: Smetana (1974), Hansen (1999)
Literature Records: San Miguel (Smetana, 1974: 38 [map]), Santa Catalina (Fall, 1897: 236; Smetana, 1974: 38 [map]), Santa Cruz (Winters, 1927: 27; Leech, 1948: 449; Smetana, 1974: 38 [map]), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Miguel (22 SBMNH), San Nicolas (8 SBMNH), Santa Catalina (8 SBMNH), Santa Cruz (13 SBMNH), Santa Rosa (37 SBMNH)
Range: Also known from mainland (Winters, 1927; Leech, 1948; Smetana, 1974; Hansen, 1999).
Notes. See note under C. columbiana above.
Cymbiodyta punctatostriata (Horn, 1873)
Nomenclatural Authority: Smetana (1974), Hansen (1999)
Literature Records: Santa Cruz (Smetana, 1974: 24)
Digitized Records: Santa Cruz (10 SBMNH)
Range: Also known from mainland (Smetana, 1974; Hansen, 1999).
Enochrus Thomson, 1859
Nomenclatural Authority: Hansen (1999)
Notes. Twelve species of Enochrus are known from California (M. L. Gimmel, 2022, unpublished data); these belong to three subgenera, Enochrus (s.str.), Lumetus Zaitzev, 1908, and Methydrus Rey, 1885. The genus was revised for North America by Gundersen (1978).
Enochrus (Enochrus) carinatus (LeConte, 1855)
Nomenclatural Authority: Gundersen (1978), Hansen (1999)
Literature Records: San Miguel (Cockerell, 1940: 285)
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Gundersen, 1978).
Notes. The nominate subspecies, E. c. carinatus (LeConte, 1855), is the only subspecies occurring in California (Gundersen, 1978).
Enochrus (Enochrus) piceus Miller, 1964
Nomenclatural Authority: Gundersen (1978), Hansen (1999)
Literature Records: none
Digitized Records: San Nicolas (1 SBMNH), Santa Catalina (5 SBMNH), Santa Cruz (21 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Gundersen, 1978; Hansen, 1999).
Notes. The nominate subspecies, E. p. piceus Miller, 1964, is the only subspecies occurring in California (Gundersen, 1978).
Enochrus (Lumetus) hamiltoni (Horn, 1890)
Nomenclatural Authority: Gundersen (1978), Hansen (1999)
Literature Records: none
Digitized Records: San Miguel (5 SBMNH)
Range: Also known from mainland (Gundersen, 1978; Hansen, 1999).
Enochrus (Methydrus) cristatus (LeConte, 1855)
Nomenclatural Authority: Gundersen (1978), Hansen (1999)
Literature Records: none
Digitized Records: Santa Cruz (5 SBMNH)
Range: Also known from mainland (Gundersen, 1978; Hansen, 1999).
Enochrus (Methydrus) pygmaeus (Fabricius, 1792)
Nomenclatural Authority: Gundersen (1978), Hansen (1999)
Literature Records: none
Digitized Records: Santa Catalina (8 SBMNH)
Range: Also known from mainland (Gundersen, 1978; Hansen, 1999).
Notes. Enochrus p. pectoralis (LeConte, 1855) is the subspecies occurring in coastal California (Gundersen, 1978).
Hydrophilinae
Notes. Four tribes, eight genera, and 47 species of Hydrophilinae are known to occur in California (Hansen, 1999; M. L. Gimmel, 2022, unpublished data).
Berosini
Notes. One genus and 11 species of Berosini are known to occur in California (Van Tassell, 1966; Hansen, 1999).
Berosus Leach, 1817
Nomenclatural Authority: Hansen (1999)
Notes. Eleven species of Berosus in two subgenera, Berosus (s.str.) and Enoplurus Hope, 1838, are known to occur in California (M. L. Gimmel, 2022, unpublished data). The species were revised for North America by Van Tassell (1966).
Berosus (Berosus) fraternus LeConte, 1855
Nomenclatural Authority: Van Tassell (1966), Hansen (1999)
Literature Records: Santa Catalina (Van Tassell, 1966: 223 [map])
Digitized Records: none
Range: Also known from mainland (Van Tassell, 1966; Hansen, 1999).
Berosus (Berosus) hatchi Miller, 1965
Nomenclatural Authority: Van Tassell (1966), Hansen (1999)
Literature Records: Santa Catalina (Van Tassell, 1966: 214)
Digitized Records: none
Range: Also known from mainland (Van Tassell, 1966; Hansen, 1999).
Berosus (Berosus) infuscatus LeConte, 1855
Nomenclatural Authority: Van Tassell (1966), Hansen (1999)
Literature Records: none
Digitized Records: San Clemente (1 SBMNH)
Range: Also known from mainland (Van Tassell, 1966; Hansen, 1999).
Berosus (Enoplurus) punctatissimus LeConte, 1852
Nomenclatural Authority: Van Tassell (1966), Hansen (1999)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: Santa Catalina (10 SBMNH), Santa Cruz (11 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Van Tassell, 1966; Hansen, 1999).
Hydrobiusini
Notes. Two genera and three species of Hydrobiusini have been recorded from California (Hansen, 1999).
Hydrobius Leach, 1815
Nomenclatural Authority: Hansen (1999)
Digitized Records (genus-only): Santa Cruz (1 UCSB)
Notes. Only one species of Hydrobius is known to occur in California (Hansen, 1999).
Hydrobius fuscipes (Linnaeus, 1758)
Nomenclatural Authority: Hansen (1999)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: none
Range: Also known from mainland (Hansen, 1999).
Hydrophilini
Notes. Three genera and 12 species of Hydrophilini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Hydrochara Berthold, 1827
Nomenclatural Authority: Hansen (1999)
Notes. Two species of Hydrochara have been recorded from California (Hansen, 1999). The genus was revised by Smetana (1980).
Hydrochara lineata (LeConte, 1855)
Nomenclatural Authority: Smetana (1980), Hansen (1999)
Literature Records: Santa Cruz (LeConte, 1876: 298; Fall, 1897: 236; Fall, 1901: 56; Fall & Davis, 1934: 144; Smetana, 1980: 67), Santa Rosa (Fall, 1897: 236; Fall, 1901: 56; Smetana, 1980: 67)
Digitized Records: Santa Cruz (16 LACM; 9 SBMNH; 2 UCSB)
Range: Also known from mainland (Fall, 1901; Smetana, 1980; Hansen, 1999).
Notes. This species was recorded as Hydrocharis glaucus LeConte by LeConte (1876) and Fall (1897, 1901); this name is now a junior synonym of H. lineata (see Smetana, 1980). Fall & Davis (1934) recorded this species as Hydrophilus lineatus (LeConte). The dot on Smetana’s (1980: 23) “Map 3” in the Pacific Ocean south of San Clemente Island is in error (Miller & Menke, 1981: 68).
Hydrophilus Geoffroy, 1762
Nomenclatural Authority: Hansen (1999)
Notes. Two species of Hydrophilus, both belonging to the nominate subgenus, have been recorded from California (Short & McIntosh, 2014). The North American species were reviewed by Short & McIntosh (2014).
Hydrophilus (Hydrophilus) triangularis Say, 1823
Nomenclatural Authority: Hansen (1999), Short & McIntosh (2014)
Literature Records: Santa Cruz (Furlong & Wenner, 2002: 250)
Digitized Records: San Clemente (1 iNat), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Short & McIntosh, 2014).
Tropisternus Solier, 1834
Nomenclatural Authority: Hansen (1999)
Digitized Records (genus-only): Santa Cruz (3 UCSB)
Notes. Eight species of Tropisternus are known to occur in California, all belonging to the nominate subgenus (M. L. Gimmel, 2022, unpublished data).
Tropisternus (Tropisternus) affinis Motschulsky, 1859
Nomenclatural Authority: Hansen (1999)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: Santa Catalina (8 SBMNH; 66 LACM), Santa Cruz (40 SBMNH; 12 LACM; 6 UCSB), Santa Rosa (18 SBMNH; 42 LACM)
Range: Also known from mainland (Hansen, 1999).
Notes. This species has been reported as Tropisternus ellipticus (LeConte, 1855) by most prior workers, including Fall (1897). However, that name is a junior primary homonym and is permanently invalid (Hansen, 1999).
Tropisternus (Tropisternus) californicus (LeConte, 1855)
Nomenclatural Authority: Hansen (1999)
Literature Records: Santa Catalina (Seavey, 1892: 262; Fall, 1897: 236), Santa Cruz (LeConte, 1876: 298; Fall, 1897: 236; Fall & Davis, 1934: 144)
Digitized Records: none
Range: Also known from mainland (Hansen, 1999).
Laccobiini
Notes. Two genera and 21 species of Laccobiini are known to occur in California (Hansen, 1999; M. L. Gimmel, 2022, unpublished data).
Laccobius Erichson, 1837
Nomenclatural Authority: Hansen (1999)
Notes. The species of Laccobius were revised for the United States in two publications by Gentili (1986a, 1986b). Fifteen species of the genus have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Laccobius (Hydroxenus) californicus d’Orchymont, 1942
Nomenclatural Authority: Hansen (1999)
Literature Records: Santa Cruz (Gentili, 1986b: 47)
Digitized Records: Santa Cruz (4 SBMNH)
Range: Also known from mainland (Gentili, 1986b).
Laccobius (Hydroxenus) ellipticus LeConte, 1855
Nomenclatural Authority: Hansen (1999)
Literature Records: Santa Catalina (Fall, 1897: 236), Santa Cruz (Winters, 1926: 50; d’Orchymont, 1942: 4; Gentili, 1986b: 49), Santa Rosa (Fall, 1897: 236)
Digitized Records: Santa Catalina (4 SBMNH), Santa Cruz (14 SBMNH), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Gentili, 1986b).
Notes. Fall (1897) mistakenly recorded this species as “Laccophilus ellipticus”.
Laccobius (Microlaccobius) insolitus d’Orchymont, 1942
Nomenclatural Authority: Hansen (1999)
Literature Records: none
Digitized Records: San Nicolas (4 SBMNH), Santa Rosa (7 SBMNH)
Range: Also known from mainland (Gentili, 1986b).
Sphaeridiinae
Notes. Three tribes, seven genera, and 28 species of Sphaeridiinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Megasternini
Notes. Five genera and 23 species of Megasternini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Agna Smetana, 1978
Nomenclatural Authority: Hansen (1999)
Notes. One species of Agna has been recorded from California (Hansen, 1999). The species were reviewed by Smetana (1978) and Arriaga-Varela, Cortés-Aguilar & Fikáček (2019).
Agna capillata (LeConte, 1855)
Nomenclatural Authority: Smetana (1978), Hansen (1999)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (Smetana, 1978; Arriaga-Varela, Cortés-Aguilar & Fikáček, 2019).
Notes. This species apparently develops exclusively in rotting cacti and other succulents (Arriaga-Varela, Cortés-Aguilar & Fikáček, 2019).
Cercyon Leach, 1817
Nomenclatural Authority: Smetana (1978), Hansen (1999)
Digitized Records (genus-only): San Miguel (223 LACM), San Nicolas (53 LACM)
Notes. Eighteen species of Cercyon are known to occur in California, belonging to three subgenera, Cercyon (s.str.), Paracercyon Seidlitz, 1888, and Prostercyon Smetana, 1978 (M. L. Gimmel, 2022, unpublished data). Smetana (1978) revised the North American species, and Suzumura, Kobayashi & Ôhara (2019) provided an update to certain western coastal species. “Cercyon sp. larvae” were reported from Santa Catalina Island by Straughan & Hadley (1980: 392).
Cercyon (Cercyon) fimbriatus Mannerheim, 1852
Nomenclatural Authority: Smetana (1978), Hansen (1999), Suzumura, Kobayashi & Ôhara (2019)
Literature Records: San Miguel (Smetana, 1978: 145 [map]), Santa Catalina (Smetana, 1978: 145 [map])
Digitized Records: San Clemente (5 SBMNH), San Miguel (19 SBMNH), San Nicolas (2 SBMNH), Santa Catalina (7 SBMNH), Santa Cruz (33 SBMNH), Santa Rosa (8 SBMNH)
Range: Also known from mainland (Smetana, 1978; Hansen, 1999; Suzumura, Kobayashi & Ôhara, 2019).
Cercyon (Cercyon) haemorrhoidalis (Fabricius, 1775)
Nomenclatural Authority: Smetana (1978), Hansen (1999)
Literature Records: Santa Barbara (Miller & Miller, 1985: 123)
Digitized Records: Santa Rosa (4 SBMNH)
Range: Also known from mainland (Smetana, 1978; Hansen, 1999).
Notes. Introduced to North America from the Palearctic realm (Smetana, 1978). Misspelled by Miller & Miller (1985) as “Cercyon haemorrhoides”.
Cercyon (Cercyon) luniger Mannerheim, 1853
Nomenclatural Authority: Suzumura, Kobayashi & Ôhara (2019)
Literature Records: San Miguel (Straughan & Hadley, 1980: 392), Santa Catalina (Fall, 1897: 236; Fall, 1901: 58; Blackwelder, 1931: 24; Leech, 1948: 458; Smetana, 1978: 149), Santa Cruz (Smetana, 1978: 149)
Digitized Records: San Clemente (3 SBMNH), Santa Cruz (8 SBMNH)
Range: Also known from mainland (Suzumura, Kobayashi & Ôhara, 2019).
Notes. This species was originally referred to as C. luniger by Fall (1897, 1901), Blackwelder (1931), Leech (1948), and Straughan & Hadley (1980). Smetana (1978) did not report his concept of C. luniger from the islands, but included Fall and CASC island material among the paratypes of his Cercyon spathifer Smetana, 1978. However, Suzumura, Kobayashi & Ôhara (2019) synonymized C. spathifer with C. luniger; what was previously considered C. luniger was given a new name. The distribution map in Suzumura, Kobayashi & Ôhara (2019: 480) also shows a Channel Island record for C. luniger, presumably from an earlier instance of the California Beetle Project pages, but the resolution does not allow identification of the island.
Cercyon (Cercyon) quisquilius (Linnaeus, 1761)
Nomenclatural Authority: Smetana (1978), Hansen (1999)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Smetana, 1978; Hansen, 1999).
Notes. Introduced to North America from the Palearctic realm (Smetana, 1978).
Sphaeridiini
Notes. One genus and three species of Sphaeridiini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Sphaeridium Fabricius, 1775
Nomenclatural Authority: Hansen (1999)
Notes. Three introduced species of Sphaeridium are known to occur in California (M. L. Gimmel, 2022, unpublished data). These species were treated by Smetana (1978).
Sphaeridium scarabaeoides (Linnaeus, 1758)
Nomenclatural Authority: Smetana (1978), Hansen (1999)
Literature Records: none
Digitized Records: Santa Catalina (2 LACM), Santa Cruz (4 SBMNH), Santa Rosa (44 LACM; 3 SBMNH)
Range: Also known from mainland (Smetana, 1978; Hansen, 1999).
Notes. Introduced to North America from the Palearctic realm (Smetana, 1978)
SCARABAEOIDEA
Geotrupidae
Notes. Two subfamilies, five genera, and seven species of Geotrupidae have been recorded from California (Howden, 1984; M. L. Gimmel, 2022, unpublished data).
Bolboceratinae
Notes. Four genera and six species of Bolboceratinae have been recorded from California (Howden, 1984; M. L. Gimmel, 2022, unpublished data).
Bolbocerastes Cartwright, 1953
Nomenclatural Authority: Smith (2009)
Notes. Two species of Bolbocerastes are known from California (Howden, 1984). The species of the genus were treated by Cartwright (1953).
Bolbocerastes regalis Cartwright, 1953
Nomenclatural Authority: Cartwright (1953), Howden (1984)
Literature Records: San Clemente (von Bloeker, 1939b: 153; Cartwright, 1953: 108)
Digitized Records: none
Range: Also known from mainland (Cartwright, 1953; Howden, 1984).
Notes. This species was misidentified as Bolboceras serratus LeConte by von Bloeker (1939b: 153).
Odonteus Samouelle, 1819
Nomenclatural Authority: Smith (2009)
Notes. One species of Odonteus is known from California (Howden, 1984). This genus has been known in recent literature as Bolboceras Kirby, 1818.
Odonteus obesus LeConte, 1859
Nomenclatural Authority: Smith (2009)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Howden, 1984).
Notes. This species was formerly known as Bolboceras obesus (e.g., in Howden (1984)).
Scarabaeidae
Notes. Seven subfamilies, 76 genera, and 321 species of Scarabaeidae are known to occur in California (M. L. Gimmel, 2022, unpublished data). The subfamily Rutelinae has yet to be recorded from the Channel Islands.
Aphodiinae
Notes. Five tribes, 36 genera, and 102 species of Aphodiinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Aegialiini
Notes. Two genera and 14 species of Aegialiini have been recorded from California (Gordon & Cartwright, 1988).
Aegialia Latreille, 1807
Nomenclatural Authority: Smith (2009)
Notes. Thirteen species of Aegialia are known to occur in California (Gordon & Cartwright, 1988); these are distributed among two subgenera, Aegialia (s.str.) and Psammoporus Thomson, 1863. The North American fauna was treated by Gordon & Cartwright (1988).
Aegialia (Aegialia) convexa Fall, 1932
Nomenclatural Authority: Gordon & Cartwright (1988)
Literature Records: San Clemente (Gordon & Cartwright, 1988: 22)
Digitized Records: none
Range: Also known from mainland (Gordon & Cartwright, 1988).
Aegialia (Aegialia) crassa LeConte, 1860
Nomenclatural Authority: Gordon & Cartwright (1988)
Literature Records: San Clemente (von Bloeker, 1939b: 153)
Digitized Records: none
Range: Also known from mainland (Gordon & Cartwright, 1988).
Aegialia (Aegialia) nigrella Brown, 1931
Nomenclatural Authority: Gordon & Cartwright (1988)
Literature Records: San Nicolas (Gordon & Cartwright, 1988: 24)
Digitized Records: none
Range: Also known from mainland (Gordon & Cartwright, 1988).
Aegialia (Aegialia) punctata Brown, 1931
Nomenclatural Authority: Gordon & Cartwright (1988)
Literature Records: San Nicolas (Gordon & Cartwright, 1988: 25)
Digitized Records: none
Range: Also known from mainland (Gordon & Cartwright, 1988).
Aphodiini
Notes. Twenty-eight genera and 66 species of Aphodiini are known to occur in California (Gordon & Skelley, 2007; M. L. Gimmel, 2022, unpublished data). The tribe was monographed for North America by Gordon & Skelley (2007).
Aphodius Illiger, 1798
Nomenclatural Authority: Gordon & Skelley (2007)
Digitized Records (genus-only): Santa Catalina (1 LACM)
Notes. Prior to Gordon & Skelley (2007), this genus was considered to encompass most of the North American fauna of Aphodiini. Only a single, adventive species occurs in North America (Gordon & Skelley, 2007).
Aphodius fimetarius (Linnaeus, 1758)
Nomenclatural Authority: Gordon & Skelley (2007)
Literature Records: none
Digitized Records: Anacapa (4 LACM; 1 SBMNH), Santa Catalina (9 LACM; 1 SBMNH; 1 iNat), Santa Cruz (2 SBMNH; 3 UCSB)
Range: Also known from mainland (Gordon & Skelley, 2007).
Notes. Adventive in North America, originating in Europe (Gordon & Skelley, 2007).
Calamosternus Motschulsky, 1859
Nomenclatural Authority: Gordon & Skelley (2007)
Notes. Only a single, adventive species occurs in North America (Gordon & Skelley, 2007).
Calamosternus granarius (Linnaeus, 1767)
Nomenclatural Authority: Gordon & Skelley (2007)
Literature Records: Santa Rosa (von Bloeker, 1939b: 153)
Digitized Records: Anacapa (1 SBMNH), San Nicolas (3 LACM; 1 SBMNH), Santa Catalina (1 LACM), Santa Rosa (2 LACM)
Range: Also known from mainland (Gordon & Skelley, 2007).
Notes. von Bloeker (1939b) recorded this species as Aphodius granarius and incorrectly attributed the species to LeConte. This species was introduced to North America from Europe (Gordon & Skelley, 2007).
Cinacanthus Schmidt, 1913
Nomenclatural Authority: Gordon & Skelley (2007)
Notes. Three species of this genus are known to occur in California (Gordon & Skelley, 2007).
Cinacanthus militaris (LeConte, 1858)
Nomenclatural Authority: Gordon & Skelley (2007)
Literature Records: San Nicolas (von Bloeker, 1939b: 153)
Digitized Records: none
Range: Also known from mainland (Gordon & Skelley, 2007).
Notes. This species was recorded by von Bloeker (1939b) as Aphodius militaris.
Labarrus Mulsant & Rey, 1869
Nomenclatural Authority: Gordon & Skelley (2007)
Notes. Two species of this genus occur in North America (Gordon & Skelley, 2007). At least one occurs in California, while the status of the other in the state remains unknown (see below).
Labarrus pseudolividus (Balthasar, 1941)
Nomenclatural Authority: Gordon & Skelley (2007)
Literature Records: San Clemente (von Bloeker, 1939b: 153), San Miguel (von Bloeker, 1939b: 153), Santa Catalina (Cockerell, 1940: 286), Santa Cruz (von Bloeker, 1939b: 153), Santa Rosa (von Bloeker, 1939b: 153)
Digitized Records: Anacapa (1 SBMNH; 24 LACM), San Clemente (7 LACM), San Miguel (22 LACM), San Nicolas (32 LACM), Santa Cruz (5 SBMNH; 198 LACM), Santa Rosa (5 LACM)
Range: Also known from mainland (Gordon & Skelley, 2007).
Notes. von Bloeker (1939b) and Cockerell (1940) recorded this species as Aphodius lividus [now Labarrus lividus (Olivier, 1789)]. Gordon & Skelley (2007: 263) noted that most North American specimens identified as L. lividus probably represent L. pseudolividus. After examination of the von Bloeker (1939b) vouchers housed in SBMNH, we have determined that these indeed represent L. pseudolividus. There are no confirmed published vouchers of true L. lividus reported from California. Labarrus lividus is probably adventive in North America, while L. pseudolividus is probably native (Gordon & Skelley, 2007).
Otophorus Mulsant, 1842
Nomenclatural Authority: Gordon & Skelley (2007)
Notes. Only a single, adventive species of this genus is known from North America (Gordon & Skelley, 2007).
Otophorus haemorrhoidalis (Linnaeus, 1758)
Nomenclatural Authority: Gordon & Skelley (2007)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Gordon & Skelley, 2007).
Notes. Introduced to North America from the Palearctic realm (Gordon & Skelley, 2007).
Planolinellus Dellacasa & Dellacasa, 2005
Nomenclatural Authority: Gordon & Skelley (2007)
Notes. Only one species of this genus occurs in North America (Gordon & Skelley, 2007).
Planolinellus vittatus (Say, 1825)
Nomenclatural Authority: Gordon & Skelley (2007)
Literature Records: San Nicolas (von Bloeker, 1939b: 153), Santa Rosa (von Bloeker, 1939b: 153)
Digitized Records: San Nicolas (41 LACM; 1 SBMNH), Santa Rosa (43 LACM; 1 SBMNH)
Range: Also known from mainland (Gordon & Skelley, 2007).
Notes. Recorded by von Bloeker (1939b) as Aphodius vittatus.
Rugaphodius Gordon & Skelley, 2007
Nomenclatural Authority: Gordon & Skelley (2007)
Notes. Only one species of this genus occurs in North America (Gordon & Skelley, 2007).
Rugaphodius rugatus (Schmidt, 1907)
Nomenclatural Authority: Gordon & Skelley (2007)
Literature Records: San Nicolas (von Bloeker, 1939b: 153), Santa Rosa (von Bloeker, 1939b: 153)
Digitized Records: San Nicolas (1 LACM), Santa Rosa (1 LACM)
Range: Also known from mainland (Gordon & Skelley, 2007).
Notes. This species was recorded by von Bloeker (1939b) as Aphodius rugatus.
Psammodiini
Notes. Six genera and 11 species of Psammodiini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Tesarius Rakovič, 1981
Nomenclatural Authority: Rakovič (1984)
Notes. Four species of Tesarius have been recorded from California (Rakovič, 1984). The species of the genus were treated by Rakovič (1984).
Tesarius mcclayi (Cartwright, 1955)
Nomenclatural Authority: Rakovič (1984)
Literature Records: none
Digitized Records: San Nicolas (1 SBMNH), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Cartwright, 1955).
Cetoniinae
Notes. Four tribes, five genera, and 14 species of Cetoniinae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Cremastocheilini
Notes. Two genera and 10 species of Cremastocheilini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Cremastocheilus Knoch, 1801
Nomenclatural Authority: Smith (2009)
Digitized Records (genus-only): Santa Catalina (2 SBMNH)
Notes. Nine species of Cremastocheilus are known from California (M. L. Gimmel, 2022, unpublished data).
Cremastocheilus schaumii LeConte, 1853
Nomenclatural Authority: Smith (2009)
Literature Records: Santa Catalina (von Bloeker, 1939b: 156; Cockerell, 1940: 286)
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland.
Notes. von Bloeker (1939b) reported “one adult collected at edge of ant-hill”. Reported as Cremastocheilus schaumi by von Bloeker (1939b) and Cockerell (1940).
Gymnetini
Notes. One species of Gymnetini has been recorded from California (Goodrich, 1966).
Cotinis Burmeister, 1842
Nomenclatural Authority: Smith (2009)
Notes. One species of Cotinis is known from California (Goodrich, 1966). The genus was revised by Goodrich (1966).
Cotinis mutabilis (Gory & Percheron, 1833)
Nomenclatural Authority: Goodrich (1966); Smith (2009)
Literature Records: none
Digitized Records: Santa Catalina (1 iNat)
Range: Also known from mainland (Goodrich, 1966).
Dynastinae
Notes. Five tribes, seven genera, and 12 species of Dynastinae have been recorded from California (Ratcliffe & Cave, 2017). The North American fauna of the subfamily was treated by Ratcliffe & Cave (2017).
Cyclocephalini
Notes. One genus and five or six species of Cyclocephalini have been recorded from California (Ratcliffe & Cave, 2017).
Cyclocephala Dejean, 1821
Nomenclatural Authority: Ratcliffe & Cave (2017)
Notes. Five or six species of Cyclocephala have been recorded from California (Ratcliffe & Cave, 2017).
Cyclocephala borealis Arrow, 1911
Nomenclatural Authority: Ratcliffe & Cave (2017)
Literature Records: San Clemente (von Bloeker, 1939b: 156; Ratcliffe & Cave (2017: 61), Santa Catalina (Fall, 1897: 238; von Bloeker, 1939b: 156)
Digitized Records: none
Range: Also known from mainland (Ratcliffe & Cave, 2017).
Notes. von Bloeker (1939b) recorded this species as Ochrosidia villosa (Burmeister, 1855); this name is preoccupied in Cyclocephala and C. borealis is the currently valid name for this taxon. This species is otherwise known to occur only in the eastern USA; the Channel Islands records are suspect. Ratcliffe & Cave (2017: 61) wrote: “We have a strange record of one male specimen collected in April, 1939 from San Clemente Island in the Channel Islands…”.
Cyclocephala hirta LeConte, 1861
Nomenclatural Authority: Ratcliffe & Cave (2017)
Literature Records: none
Digitized Records: Santa Catalina (18 LACM)
Range: Also known from mainland (Ratcliffe & Cave, 2017).
Notes. According to Ratcliffe & Cave (2017), the subspecies present in southern California is the nominate subspecies, C. h. hirta LeConte, 1861.
Cyclocephala longula LeConte, 1863
Nomenclatural Authority: Ratcliffe & Cave (2017)
Literature Records: San Clemente (von Bloeker, 1939b: 156), San Nicolas (von Bloeker, 1939b: 156), Santa Cruz (von Bloeker, 1939b: 156), Santa Rosa (von Bloeker, 1939b: 156)
Digitized Records: none
Range: Also known from mainland (Ratcliffe & Cave, 2017).
Notes. von Bloeker (1939b) recorded this species as Ochrosidia longula from Santa Cruz and Santa Rosa; he also recorded it as Ochrosidia obesula Casey, 1915 from San Clemente and San Nicolas. The latter name is now considered a synonym of C. longula (see Ratcliffe & Cave, 2017).
Cyclocephala melanocephala (Fabricius, 1775)
Nomenclatural Authority: Ratcliffe & Cave (2017)
Literature Records: San Miguel (von Bloeker, 1939b: 156), Santa Cruz (von Bloeker, 1939b: 156), Santa Rosa (von Bloeker, 1939b: 156)
Digitized Records: none
Range: Also known from mainland (Ratcliffe & Cave, 2017).
Notes. von Bloeker (1939b) recorded this species as Dichromina dimidiata (Burmeister, 1847) (now considered a junior synonym of C. melanocephala, see Ratcliffe & Cave, 2017) and noted that it was “fairly common in flowers of Datura metalloides, August”.
Cyclocephala pasadenae (Casey, 1915)
Nomenclatural Authority: Ratcliffe & Cave (2017)
Literature Records: San Miguel (von Bloeker, 1939b: 156), Santa Cruz (von Bloeker, 1939b: 156), Santa Rosa (von Bloeker, 1939b: 156)
Digitized Records: none
Range: Also known from mainland (Ratcliffe & Cave, 2017).
Notes. von Bloeker (1939b) recorded this species as Ochrosidia pasadenae.
Pentodontini
Notes. Two genera and three species of Pentodontini have been recorded from California (Ratcliffe & Cave, 2017).
Ligyrus Casey, 1915
Nomenclatural Authority: López-García & Deloya (2022)
Notes. One species of Ligyrus has been recorded from California (Ratcliffe & Cave, 2017; López-García & Deloya, 2022).
Ligyrus gibbosus (DeGeer, 1774)
Nomenclatural Authority: López-García & Deloya (2022)
Literature Records: San Clemente (von Bloeker, 1939b: 156), San Miguel (von Bloeker, 1939b: 156), San Nicolas (von Bloeker, 1939b: 156), Santa Rosa (von Bloeker, 1939b: 156)
Digitized Records: San Clemente (40 LACM; 2 SBMNH), San Miguel (18 LACM; 1 SBMNH), San Nicolas (26 LACM; 1 SBMNH), Santa Cruz (13 LACM; 1 SBMNH), Santa Rosa (13 LACM; 1 SBMNH)
Range: Also known from mainland (Ratcliffe & Cave, 2017).
Notes. von Bloeker (1939b) recorded this species as two different taxa, Ligyrus californicus Casey, 1909 and L. scitulus Casey, 1915. The former was reported from San Clemente, San Miguel, San Nicolas, and Santa Rosa and was noted to be “common in loose sand under Abronia maritima, A. alba, Franseria bipinnatifida, Astragalus nevinii and A. miguelensis, and occasional at lights” (von Bloeker, 1939b: 156); the latter was reported from San Clemente, San Miguel, and San Nicolas and was noted to be “occasional at lights, much less common than the preceding species [L. scitulus]” (von Bloeker, 1939b: 156). Both taxa are now considered junior synonyms of L. gibbosus (see Ratcliffe & Cave, 2017), and the species has been recently placed in the genus Tomarus Erichson, 1847 but subsequently transferred back to Ligyrus (López-García & Deloya, 2022).
Melolonthinae
Notes. Nine tribes, 15 genera, and 172 species of Melolonthinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Dichelonychini
Notes. Three genera and 46 species of Dichelonychini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Coenonycha Horn, 1876
Nomenclatural Authority: Evans & Smith (2009)
Notes. Twenty-six species of Coenonycha have been recorded from California (Evans & Smith, 1986). The species of the genus were keyed by Evans & Smith (1986).
Coenonycha clementina Casey, 1909
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: San Clemente (Casey, 1909: 281; von Bloeker, 1939b: 155; Cazier & McClay, 1943: 17; Evans, 1985: 86; Miller, 1985a: 19; Evans & Smith, 1986: 86; Evans & d’Hotman, 1988: 207)
Digitized Records: San Clemente (2 ASUHIC; 5 LACM; 6 SBMNH; 6 USNM)
Range: Endemic (Casey, 1909; Cazier & McClay, 1943; Evans, 1985; Miller, 1985a; Evans & Smith, 1986; Evans & d’Hotman, 1988).
Notes. A flightless species (Cazier & McClay, 1943: 7; Evans, 1985: 86). Adults were reported from among the roots of a perennial Lupinus (Fabaceae) species during March (Evans, 1985: 86). This species was assessed by the US Fish and Wildlife Service and available data did not suggest Endangered or Threatened status at the time of assessment (Greenwalt, 1977).
Coenonycha clypeata McClay, 1943
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: Santa Catalina (Cazier & McClay, 1943: 23; Miller, 1985a: 19; Caterino & Chandler, 2010: 187)
Digitized Records: Santa Catalina (3 LACM; 1 SBMNH)
Range: Endemic (Cazier & McClay, 1943; Miller, 1985a; Caterino & Chandler, 2010).
Notes. Early Santa Catalina records of Coenonycha rotundata LeConte, 1856 (Fall, 1897: 238; von Bloeker, 1939b: 155) probably referred to this species or to C. fulva.
Coenonycha fulva McClay, 1943
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: Santa Catalina (Cazier & McClay, 1943: 23; Evans, 1985: 86; Miller, 1985a; Evans & Smith, 1986: 90; Caterino & Chandler, 2010: 187)
Digitized Records: Anacapa (1 LACM), Santa Catalina (2 ASUHIC; 43 LACM; 12 SBMNH)
Range: Endemic (Cazier & McClay, 1943; Miller, 1985a; Evans & Smith, 1986; Caterino & Chandler, 2010).
Notes. Early Santa Catalina records of C. rotundata (Fall, 1897: 238; von Bloeker, 1939b: 155) probably referred to this species or to C. clypeata. Evans (1985: 86) reported a large number of individuals from Adenostoma fasciculatum Hook. & Arn. (Rosaceae).
Coenonycha santacruzae Evans, 1986
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: Santa Cruz (Evans & Smith, 1986: 82)
Digitized Records: Santa Cruz (2 ASUHIC; 1 SBMNH)
Range: Endemic (Evans & Smith, 1986).
Notes. Evans & Smith (1986) reported adults from Adenostoma fasciculatum, Cercocarpus traskae Eastw. (Rosaceae), and Artemisia californica Less. (Asteraceae).
Dichelonyx Harris, 1827
Nomenclatural Authority: Evans & Smith (2009)
Notes. Seventeen species of Dichelonyx are known from California (M. L. Gimmel, 2022, unpublished data).
Dichelonyx backii Kirby, 1837
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: none
Digitized Records: Santa Cruz (5 LACM)
Range: Also known from mainland (Evans & Smith, 2009).
Dichelonyx fulgida LeConte, 1856
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: Santa Cruz (von Bloeker, 1939b: 155)
Digitized Records: none
Range: Also known from mainland (Evans & Smith, 2009).
Notes. The subspecies occurring on the islands is D. f. crotchii Horn, 1876; it was recorded as Dichelonyx crotchi by von Bloeker (1939b).
Dichelonyx pusilla LeConte, 1856
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: San Miguel (von Bloeker, 1939b: 155), Santa Cruz (von Bloeker, 1939b: 155), Santa Rosa (von Bloeker, 1939b: 155)
Digitized Records: San Miguel (2 LACM), Santa Cruz (3 LACM), Santa Rosa (2 LACM)
Range: Also known from mainland (Evans & Smith, 2009).
Diplotaxini
Notes. One genus and 22 species of Diplotaxini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Diplotaxis Kirby, 1837
Nomenclatural Authority: Evans & Smith (2009)
Notes. Twenty-two species of Diplotaxis are known to occur in California (M. L. Gimmel, 2022, unpublished data). The genus was revised in two papers by Vaurie (1958, 1960).
Diplotaxis fimbriata Fall, 1909
Nomenclatural Authority: Vaurie (1960), Evans & Smith (2009)
Literature Records: San Clemente (von Bloeker, 1939b: 154), Santa Cruz (von Bloeker, 1939b: 154)
Digitized Records: none
Range: Also known from mainland (Vaurie, 1960).
Diplotaxis subangulata LeConte, 1856
Nomenclatural Authority: Vaurie (1960), Evans & Smith (2009)
Literature Records: San Clemente (von Bloeker, 1939b: 154), Santa Cruz (von Bloeker, 1939b: 154), Santa Rosa (von Bloeker, 1939b: 154)
Digitized Records: San Clemente (1 LACM; 1 SBMNH), Santa Cruz (3 LACM), Santa Rosa (1 LACM; 1 SBMNH)
Range: Also known from mainland (Vaurie, 1960).
Hopliini
Notes. One genus and four species of Hopliini have been recorded from California (Hardy, 1977).
Hoplia Illiger, 1803
Nomenclatural Authority: Evans & Smith (2009)
Notes. Four species of Hoplia have been recorded from California (Hardy, 1977). The species were revised by Hardy (1977).
Hoplia callipyge LeConte, 1856
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: Santa Cruz (Hardy, 1977: 14)
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Hardy, 1977).
Melolonthini
Notes. Five genera and 28 species of Melolonthini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Amblonoxia Reitter, 1902
Nomenclatural Authority: Evans & Smith (2009)
Notes. Six species of Amblonoxia have been recorded from California (Hardy, 1974). The species were revised by Hardy (1974). Members of this genus were until recently placed in the genus Parathyce Hardy, 1974, which is now a junior synonym of Amblonoxia (Evans & Smith, 2009).
Amblonoxia palpalis (Horn, 1880)
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: San Clemente (von Bloeker, 1939b: 155; Hardy, 1974: 20; Miller & Miller, 1985: 124), San Nicolas (von Bloeker, 1939b: 155; Hardy, 1974: 20; Miller & Miller, 1985: 124), Santa Barbara (von Bloeker, 1939b: 155; Miller & Miller, 1985: 124)
Digitized Records: Anacapa (2 SBMNH), San Nicolas (1 SBMNH)
Range: Also known from mainland (Hardy, 1974).
Notes. This species was recorded as Thyce blaisdelli Casey by von Bloeker (1939b), which is now considered a junior synonym of A. palpalis (see Evans & Smith, 2009). It was recorded as Parathyce palpalis by Hardy (1974) and Miller & Miller (1985).
Polyphylla Harris, 1841
Nomenclatural Authority: Evans & Smith (2009)
Literature Records (genus-only): San Miguel (von Bloeker, 1939b: 155), Santa Cruz (von Bloeker, 1939b: 155)
Digitized Records (genus-only): Santa Cruz (1 UCSB)
Notes. Seventeen species of Polyphylla have been recorded from California (M. L. Gimmel, 2022, unpublished data). The genus was revised by Young (1988). The above literature records were deemed unidentifiable to species (because of specimen damage) by von Bloeker (1939b).
Polyphylla crinita LeConte, 1856
Nomenclatural Authority: Young (1988), Evans & Smith (2009)
Literature Records: Santa Cruz (von Bloeker, 1939a: 148; von Bloeker, 1939b: 154; Young, 1967: 307; Young, 1988: 55), Santa Rosa (von Bloeker, 1939a: 149; von Bloeker, 1939b: 154; Young, 1967: 307; Young, 1988: 55)
Digitized Records: Santa Catalina (16 LACM), Santa Cruz (47 LACM; 1 SBMNH)
Range: Also known from mainland (Young, 1967, 1988).
Notes. This species was first recorded from the Channel Islands as the Santa Cruz-endemic Polyphylla ona von Bloeker, 1939 and the Santa Rosa-endemic Polyphylla santarosae von Bloeker, 1939 by von Bloeker (1939a, 1939b). These were synonymized with P. crinita by Cazier (1940: 137), who also synonymized Polyphylla nigra Casey, 1914, Polyphylla martini von Bloeker, 1939, and Polyphylla santacruzae von Bloeker, 1939 under the same name; Young (1967: 305) followed this arrangement. Later, however, Young (1988) removed the latter three taxa from synonymy with P. crinita (see entry for P. nigra below).
Polyphylla nigra Casey, 1914
Nomenclatural Authority: Young (1988), Evans & Smith (2009)
Literature Records: Santa Cruz (von Bloeker, 1939a: 149; von Bloeker, 1939b: 154; Young, 1988: 68), Santa Rosa (von Bloeker, 1939a: 149; von Bloeker, 1939b: 155; Young, 1988: 68)
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (11 SBMNH)
Range: Also known from mainland (Young, 1988).
Notes. This species was first recorded from the Channel Islands as the Santa Rosa-endemic P. martini and the Santa Cruz-endemic P. santacruzae by von Bloeker (1939a, 1939b). These were synonymized with P. crinita by Cazier (1940: 137), which was followed by Young (1967: 305). Later, Young (1988: 67) synonymized these two names with the newly resurrected P. nigra. The larva of “P. santacruzae” was reported from “beneath roots of Eucalyptus, August” (von Bloeker, 1939b).
Phobetusini
Notes. One genus and nine species of Phobetusini have been recorded from California (Hardy, 1978).
Phobetus LeConte, 1856
Nomenclatural Authority: Evans & Smith (2009)
Literature Records (genus-only): San Clemente (Doyen, 1974: 87), Santa Barbara (Miller & Miller, 1985: 125)
Digitized Records (genus-only): San Clemente (4 SBMNH), San Nicolas (11 SBMNH), Santa Rosa (26 SBMNH)
Notes. Nine species of Phobetus have been recorded from California (Hardy, 1978). Barrett (1935) and Hardy (1978) provided keys to species. The San Clemente, San Miguel, and Santa Barbara records of Phobetus comatus LeConte, 1856 in von Bloeker (1939b: 155) and San Clemente, San Nicolas, and Santa Rosa specimens in the SBMNH collection apparently represent one or more new species in the Phobetus testaceus group; the southern island specimens appear to be brachypterous (M. L. Gimmel, 2021, personal observation). The genus needs revision (A. Evans, 2021, personal communication).
Phobetus ciliatus Barrett, 1935
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: Santa Catalina (Barrett, 1935: 51; Cazier, 1937: 84; Cockerell, 1940: 286; Miller, 1985a: 19; Caterino & Chandler, 2010: 187)
Digitized Records: Santa Catalina (17 LACM; 4 SBMNH)
Range: Endemic (Barrett, 1935; Cazier, 1937; Hardy, 1978; Miller, 1985a; Caterino & Chandler, 2010).
Notes. Recorded simply from “Channel Islands, California” by Hardy (1978). However, this distinctive species appears to be restricted to Santa Catalina Island (A. Evans, 2021, personal communication). Fall’s (1897: 238; 1901: 141) and von Bloeker’s (1939b: 155) records of P. comatus from that island apparently represent P. ciliatus (Cockerell, 1940: 286).
Phobetus testaceus LeConte, 1862
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: Santa Cruz (LeConte, 1861: 346; Casey, 1909: 282; Fall & Davis, 1934: 144; Cazier, 1937: 85; Cockerell, 1940: 286; Evans, 1985: 87; Miller, 1985a: 19)
Digitized Records: Santa Cruz (22 SBMNH; 3 UCSB)
Range: Endemic (LeConte, 1861; Cazier, 1937; Hardy, 1978; Evans, 1985; Miller, 1985a).
Notes. Casey (1909: 282) opined that this was “probably… a well-marked subspecies of comatus.” von Bloeker (1939b: 155) noted that Fall (1897: 238) recorded this species from Santa Cruz Island under P. comatus, and did not provide any new records or comments on its validity. Evans (1985: 87) reported adults copulating and feeding on Cercocarpus betuloides blancheae (C. K. Schneid.) Little. Recorded simply from “Channel Islands, California” by Hardy (1978). Apparently endemic to Santa Cruz Island (A. Evans, 2021, personal communication).
Rhizotrogini
Notes. One genus and 11 species of Rhizotrogini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Phyllophaga Harris, 1827
Nomenclatural Authority: Evans & Smith (2009)
Notes. Eleven species of Phyllophaga are known from California, belonging to two subgenera, Listrochelus Blanchard, 1851 and Phyllophaga (s.str.) (M. L. Gimmel, 2022, unpublished data).
Phyllophaga (Listrochelus) mucorea (LeConte, 1856)
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: San Clemente (von Bloeker, 1939b: 154)
Digitized Records: none
Range: Also known from mainland (Evans & Smith, 2009).
Notes. Reported by von Bloeker (1939b) as Listrochelus mucoreus.
Sericini
Notes. One genus and 49 species of Sericini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Serica MacLeay, 1819
Nomenclatural Authority: Evans & Smith (2009)
Literature Records (genus-only): San Clemente (Doyen, 1974: 87), Santa Barbara (Miller & Miller, 1985: 125)
Digitized Records (genus-only): Santa Cruz (1 UCSB)
Notes. Forty-nine species of Serica have been recorded from California (M. L. Gimmel, 2022, unpublished data). The genus is under revision by P. Lago (2022, personal communication).
Serica alternata LeConte, 1856
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: San Clemente (von Bloeker, 1939b: 154), San Nicolas (von Bloeker, 1939b: 154), Santa Barbara (von Bloeker, 1939b: 154)
Digitized Records: none
Range: Also known from mainland (Dawson, 1933).
Notes. Miller & Miller (1985) rejected von Bloeker’s (1939b) island records of this species, since Dawson (1933) had revised the species previously included under S. alternata, and von Bloeker (1939b) apparently ignored this.
Serica catalina Dawson, 1947
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: Santa Catalina (Dawson, 1947: 234; Miller, 1985a: 19; Caterino & Chandler, 2010: 187)
Digitized Records: none
Range: Endemic (Dawson, 1947; Miller, 1985a; Caterino & Chandler, 2010).
Serica cruzi Saylor, 1939
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: Santa Cruz (Saylor, 1939: 55; Miller, 1985a: 19)
Digitized Records: Santa Cruz (5 LACM; 1 SBMNH; 1 UCSB; 1 USNM)
Range: Endemic (Saylor, 1939; Miller, 1985a).
Serica mixta LeConte, 1856
Nomenclatural Authority: Evans & Smith (2009)
Literature Records: San Clemente (von Bloeker, 1939b: 154), San Miguel (von Bloeker, 1939b: 154), San Nicolas (von Bloeker, 1939b: 154), Santa Cruz (von Bloeker, 1939b: 154), Santa Rosa (von Bloeker, 1939b: 154)
Digitized Records: none
Range: Also known from mainland (Dawson, 1947).
Scarabaeinae
Notes. Five tribes, seven genera, and nine species of Scarabaeinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Canthon Hoffmannsegg, 1817
Nomenclatural Authority: Smith (2009)
Notes. One species of Canthon has been recorded from California (Robinson, 1948), belonging to the subgenus Boreocanthon Halffter, 1958.
Canthon (Boreocanthon) simplex LeConte, 1857
Nomenclatural Authority: Robinson (1948)
Literature Records: San Clemente (von Bloeker, 1939b: 153)
Digitized Records: none
Range: Also known from mainland (Robinson, 1948).
Trogidae
Notes. This family is represented by two genera and eight species in California (Vaurie, 1955). The family was treated for North America by Vaurie (1955).
Trox Fabricius, 1775
Nomenclatural Authority: Smith (2009)
Notes. Six species of Trox have been recorded from California (Vaurie, 1955).
Trox atrox LeConte, 1854
Nomenclatural Authority: Vaurie (1955)
Literature Records: San Clemente (von Bloeker, 1939b: 154)
Digitized Records: San Clemente (2 LACM)
Range: Also known from mainland (Vaurie, 1955).
Trox gemmulatus Horn, 1874
Nomenclatural Authority: Vaurie (1955)
Literature Records: San Clemente (von Bloeker, 1939b: 154)
Digitized Records: San Clemente (5 LACM; 1 SBMNH)
Range: Also known from mainland (Vaurie, 1955).
STAPHYLINOIDEA
Colonidae, NEW FAMILY RECORD
Notes. One genus and 14 species of Colonidae have been recorded from California (Peck & Stephan, 1996; Peck & Newton, 2017). Until recently, this family was treated as a subfamily of Leiodidae (see Cai et al., 2022).
Colon Herbst, 1797
Nomenclatural Authority: Peck & Newton (2017)
Notes. Peck & Stephan (1996) reported 14 species of Colon from California belonging to three subgenera (Colon (s.str.), Eurycolon Ganglbauer, 1899, and Myloechus Latreille, 1806) in their revision of the North American species of the genus.
Colon (Myloechus) forceps Hatch, 1957
Nomenclatural Authority: Peck & Newton (2017)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Peck & Stephan, 1996).
Hydraenidae
Notes. Two subfamilies, five genera, and 49 species of Hydraenidae have been recorded from California (Perkins, 1980). This family was treated for the New World by Perkins (1980).
Hydraeninae
Notes. Two genera and 16 species of Hydraeninae have been recorded from California (Perkins, 1980).
Hydraena Kugelann, 1794
Nomenclatural Authority: Perkins (1980)
Digitized Records (genus-only): San Clemente (6 SBMNH), Santa Cruz (23 SBMNH), Santa Rosa (10 SBMNH)
Notes. Twelve species of Hydraena have been recorded from California (Perkins, 1980).
Hydraena arenicola Perkins, 1980
Nomenclatural Authority: Perkins (1980)
Literature Records: Santa Cruz (Perkins, 1980: 485)
Digitized Records: none
Range: Also known from mainland (Perkins, 1980).
Hydraena circulata Perkins, 1980
Nomenclatural Authority: Perkins (1980)
Literature Records: Santa Cruz (Perkins, 1980: 483)
Digitized Records: none
Range: Also known from mainland (Perkins, 1980).
Hydraena vandykei d’Orchymont, 1923
Nomenclatural Authority: Perkins (1980)
Literature Records: Santa Cruz (Perkins, 1980: 493)
Digitized Records: none
Range: Also known from mainland (Perkins, 1980).
Ochthebiinae
Notes. Three genera and 33 species of Ochthebiinae have been recorded from California (Perkins, 1980).
Ochthebius Leach, 1815
Nomenclatural Authority: Perkins (1980)
Digitized Records (genus-only): San Clemente (52 SBMNH), San Miguel (50 SBMNH), San Nicolas (22 SBMNH), Santa Catalina (14 SBMNH), Santa Cruz (30 SBMNH), Santa Rosa (103 SBMNH)
Notes. Thirty species of Ochthebius have been recorded from California (Perkins, 1980).
Ochthebius discretus LeConte, 1878
Nomenclatural Authority: Perkins (1980)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Perkins, 1980).
Ochthebius interruptus LeConte, 1852
Nomenclatural Authority: Perkins (1980)
Literature Records: Santa Cruz (Perkins, 1980: 507; Furlong & Wenner, 2002: 250)
Digitized Records: none
Range: Also known from mainland (Perkins, 1980).
Ochthebius puncticollis LeConte, 1852
Nomenclatural Authority: Perkins (1980)
Literature Records: Santa Cruz (Perkins, 1980: 524)
Digitized Records: Santa Cruz (3 SBMNH)
Range: Also known from mainland (Perkins, 1980).
Leiodidae
Notes. There are four subfamilies, 19 genera, and 101 species of Leiodidae known from California (Peck & Newton, 2017; M. L. Gimmel, 2022, unpublished data). A distributional catalog of the North American fauna of the family was provided by Peck & Newton (2017).
Catopocerinae
Notes. One genus and 23 species of Catopocerinae have been recorded from California (Peck & Cook, 2011).
Pinodytes Horn, 1880
Nomenclatural Authority: Peck & Newton (2017)
Notes. A total of 23 species of this genus is known from California (Peck & Cook, 2011), and it is likely that additional species will be discovered in the Channel Islands. The genus was revised by Peck & Cook (2011).
Pinodytes gibbosus Peck & Cook, 2011
Nomenclatural Authority: Peck & Newton (2017)
Literature Records: Santa Catalina (Peck & Cook, 2011: 26), Santa Cruz (Peck & Cook, 2011: 26), Santa Rosa (Peck & Cook, 2011: 26)
Digitized Records: Santa Catalina (7 SBMNH), Santa Cruz (4 SBMNH), Santa Rosa (36 SBMNH)
Range: Also known from mainland (Peck & Cook, 2011).
Notes. Caterino & Chandler (2010: 191) reported the genus Catopocerus Motschulsky, 1870 from Santa Catalina Island; this record presumably refers to this species.
Leiodinae
Notes. Three tribes, 13 genera, and 63 species of Leiodinae have been recorded from California (Peck & Newton, 2017; M. L. Gimmel, 2022, unpublished data).
Agathidium Panzer, 1797
Nomenclatural Authority: Peck & Newton (2017)
Notes. Twenty-three species of Agathidium have been recorded from California (Miller & Wheeler, 2005; Wheeler & Miller, 2005). This genus was monographed for North America in two papers by Miller & Wheeler (2005) and Wheeler & Miller (2005).
Agathidium pulchrum LeConte, 1853
Nomenclatural Authority: Peck & Newton (2017)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Miller & Wheeler, 2005).
Agathidium virile Fall, 1901
Nomenclatural Authority: Peck & Newton (2017)
Literature Records: none
Digitized Records: San Clemente (29 SBMNH), Santa Catalina (2 SBMNH)
Range: Also known from mainland (Wheeler & Miller, 2005).
Leiodes Latreille, 1797
Nomenclatural Authority: Peck & Newton (2017)
Literature Records (genus-only): Santa Catalina (Caterino & Chandler, 2010: 191)
Notes. Sixteen species of Leiodes have been recorded from California (Baranowski, 1993). This genus was revised for North America by Baranowski (1993). Caterino & Chandler (2010: 191) reported this genus as a new record from Santa Catalina Island.
Leiodes antennata (Fall, 1910)
Nomenclatural Authority: Peck & Newton (2017)
Literature Records: none
Digitized Records: Santa Catalina (3 SBMNH)
Range: Also known from mainland (Baranowski, 1993).
Leiodes paludicola (Crotch, 1874)
Nomenclatural Authority: Peck & Newton (2017)
Literature Records: none
Digitized Records: Santa Catalina (3 SBMNH)
Range: Also known from mainland (Baranowski, 1993).
Ptiliidae, NEW FAMILY RECORD
Notes. Two subfamilies, 11 genera, and 29 species of Ptiliidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Nossidiinae
Notes. Two genera and two species of Nossidiinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Motschulskium Matthews, 1872
Nomenclatural Authority: Hall (2000)
Notes. One species of Motschulskium has been recorded from the west coast of North America, including California (Hall, 2000).
Motschulskium sinuatocolle Matthews, 1872
Nomenclatural Authority: Hall (2000)
Literature Records: none
Digitized Records: San Clemente (6 SBMNH), San Nicolas (6 SBMNH), Santa Catalina (1 SBMNH)
Range: Also known from mainland (Hall, 2000).
Ptiliinae
Notes. Five tribes, nine genera, and 27 species of Ptiliinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Acrotrichini
Notes. Two genera and 11 species of Acrotrichini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Acrotrichis Motschulsky, 1848
Nomenclatural Authority: Hall (2000)
Notes. Ten species of Acrotrichis have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Acrotrichis undetermined species
Literature Records: none
Digitized Records: Santa Cruz (3 SBMNH), Santa Rosa (1 SBMNH)
Ptenidiini
Notes. One genus and four species of Ptenidiini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Ptenidium Erichson, 1845
Nomenclatural Authority: Hall (2000)
Notes. Four species of Ptenidium have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Ptenidium undetermined species
Literature Records: none
Digitized Records: Santa Rosa (33 SBMNH)
Ptiliini
Notes. Three genera and nine species of Ptiliini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Actidium Matthews, 1869
Nomenclatural Authority: Hall (2000)
Notes. Four species of Actidium have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Actidium undetermined species
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Ptiliolum Flach, 1888
Nomenclatural Authority: Hall (2000)
Notes. No species of Ptiliolum have been recorded from California in the literature (M. L. Gimmel, 2022, unpublished data); this represents a new state record for the genus.
Ptiliolum undetermined species
Literature Records: none
Digitized Records: San Clemente (9 SBMNH), Santa Catalina (38 SBMNH), Santa Rosa (30 SBMNH)
Ptinellini
Notes. Two genera and two species of Ptinellini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Pteryx Matthews, 1859
Nomenclatural Authority: Hall (2000)
Notes. No species of Pteryx have been recorded from California in the literature (M. L. Gimmel, 2022, unpublished data); this represents a new state record for the genus.
Pteryx undetermined species
Literature Records: none
Digitized Records: Santa Cruz (27 SBMNH), Santa Rosa (24 SBMNH)
Staphylinidae
Notes. Twenty-five subfamilies, 306 genera, and, 1,349 species of Staphylinidae are known to occur in California, making it the largest family of beetles in the state (M. L. Gimmel, 2022, unpublished data). The staphylinid subfamilies Dasycerinae, Euaesthetinae, Micropeplinae, Osoriinae, Proteininae, Scaphidiinae, Steninae, Trichophyinae, and Trigonurinae occur on nearby mainland but no Channel Islands records are known.
Aleocharinae
Notes. Nineteen tribes, 103 genera, and 360 species of this enormously diverse and poorly understood subfamily are known from California (M. L. Gimmel, 2022, unpublished data). “Aleocharinae, genus near Oxypoda” was reported from Santa Barbara Island by Miller & Miller (1985: 124); the specimen is housed in SBMNH but was on loan during this study.
Aleocharini
Notes. Three genera and 36 species of Aleocharini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Aleochara Gravenhorst, 1802
Nomenclatural Authority: Newton et al. (2000)
Notes. Thirty-three species of Aleochara have been recorded from California, belonging to seven subgenera, Aleochara (s.str.), Calochara Casey, 1906, Coprochara Mulsant & Rey, 1874, Echocara Casey, 1906, Emplenota Casey, 1884, Maseochara Sharp, 1883, and Xenochara Mulsant & Rey, 1874 (M. L. Gimmel, 2022, unpublished data). One specimen from Santa Catalina Island in the H.C. Fall collection in the Museum of Comparative Zoology, Harvard University was identified more recently as Aleochara (Emplenota) pacifica (Casey, 1893) by J. Klimaszewski in 1982 (S. Miller, 2022, personal communication). However, this specimen did not appear in Klimaszewski’s (1984) revision and needs to be verified.
Aleochara (Coprochara) bimaculata Gravenhorst, 1802
Nomenclatural Authority: Klimaszewski (1984)
Literature Records: San Clemente (Fall, 1897: 236), Santa Catalina (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Klimaszewski, 1984).
Aleochara (Coprochara) densissima Bernhauer, 1906
Nomenclatural Authority: Klimaszewski (1984)
Literature Records: Santa Catalina (Cockerell, 1940: 285; Klimaszewski, 1984: 27)
Digitized Records: none
Range: Also known from mainland (Klimaszewski, 1984).
Aleochara (Coprochara) sulcicollis Mannerheim, 1843
Nomenclatural Authority: Klimaszewski (1984)
Literature Records: San Miguel (Cockerell, 1940: 285), San Nicolas (Cockerell, 1940: 285), Santa Cruz (Klimaszewski, 1984: 33), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Clemente (5 SBMNH), San Miguel (17 SBMNH), San Nicolas (5 SBMNH), Santa Catalina (4 SBMNH), Santa Cruz (40 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Klimaszewski, 1984).
Notes. This species was recorded as Baryodma sulcicollis by Cockerell (1940).
Aleochara (Emplenota) curtidens Klimaszewski, 1984
Nomenclatural Authority: Klimaszewski (1984)
Literature Records: Santa Barbara (Klimaszewski, 1984: 102; Miller & Miller, 1985: 124)
Digitized Records: none
Range: Also known from mainland (Klimaszewski, 1984).
Notes. Misspelled by Miller & Miller (1985) as Aleochara curtedens.
Aleochara (Emplenota) littoralis (Mäklin, 1853)
Nomenclatural Authority: Klimaszewski (1984)
Literature Records: Santa Barbara (Fall, 1897: 236), Santa Catalina (Fall, 1897: 236; Klimaszewski, 1984: 99)
Digitized Records: none
Range: Also known from mainland (Fall, 1901; Klimaszewski, 1984).
Notes. Fall (1897) recorded this species as Polistoma arenaria Casey and Fall (1901) recorded it as Polystoma arenaria from “the islands off the coast”; P. arenaria was synonymized with A. littoralis by Klimaszewski (1984: 98).
Aleochara (Maseochara) valida LeConte, 1858
Nomenclatural Authority: Klimaszewski (1984)
Literature Records: Santa Catalina (Klimaszewski, 1984: 107)
Digitized Records: San Clemente (5 SBMNH), San Miguel (12 SBMNH), Santa Rosa (5 SBMNH)
Range: Also known from mainland (Klimaszewski, 1984).
Aleochara (Xenochara) fumata Gravenhorst, 1802
Nomenclatural Authority: Klimaszewski (1984)
Literature Records: Santa Catalina (Klimaszewski, 1984: 53)
Digitized Records: none
Range: Also known from mainland (Klimaszewski, 1984).
Notes. This species was introduced from the Palearctic region (Klimaszewski, 1984).
Aleochara (Xenochara) lanuginosa Gravenhorst, 1802
Nomenclatural Authority: Klimaszewski (1984)
Literature Records: Santa Rosa (Klimaszewski, 1984: 50)
Digitized Records: Santa Rosa (4 SBMNH)
Range: Also known from mainland (Klimaszewski, 1984).
Notes. This species was introduced from the Palearctic region (Klimaszewski, 1984).
Athetini
Notes. Twenty-eight genera and 124 species of Athetini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Acrotona Thomson, 1859
Nomenclatural Authority: Newton et al. (2000)
Digitized Records (genus-only): San Nicolas (1 SBMNH), Santa Cruz (4 SBMNH)
Notes. Six species of Acrotona have been reported from California (M. L. Gimmel, 2022, unpublished data).
Acrotona recondita (Erichson, 1839)
Nomenclatural Authority: Klimaszewski et al. (2015)
Literature Records: Santa Catalina (Casey, 1910: 136)
Digitized Records: none
Range: Also known from mainland (Casey, 1910; Klimaszewski et al., 2015).
Notes. Reported by Casey (1910) as Arisota umbrina Casey, 1910, currently a synonym of Ac. recondita (see Klimaszewski et al., 2015).
Acrotona sonomana (Casey, 1910)
Nomenclatural Authority: Gusarov (2003b)
Literature Records: Santa Catalina (Casey, 1911: 166; Miller, 1985a: 19; Gusarov, 2003b: 106)
Digitized Records: none
Range: Also known from mainland (Gusarov, 2003b).
Notes. Reported as the “endemic” Strigota (Eustrigota) seclusa Casey, 1911 by Casey (1911) and Miller (1985a). This species was synonymized with A. sonomana by Gusarov (2003b).
Adota Casey, 1910
Nomenclatural Authority: Newton et al. (2000), Gusarov (2003a)
Notes. Of the three Adota species in North America, two have been reported from California (Gusarov, 2003a), both from seashore environments.
Adota maritima (Mannerheim, 1843)
Nomenclatural Authority: Gusarov (2003a)
Literature Records: Santa Catalina (Gusarov, 2003a: 11)
Digitized Records: none
Range: Occurs along much of the west coast of North America, from southern California to Alaska (Gusarov, 2003a).
Atheta Thomson, 1858
Nomenclatural Authority: Newton et al. (2000)
Notes. Sixteen species of Atheta have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Atheta hampshirensis Bernhauer, 1909
Nomenclatural Authority: Gusarov (2003b)
Literature Records: none
Digitized Records: San Nicolas (1 SBMNH)
Range: Also known from mainland (Gusarov, 2003b).
Hydrosmecta Thomson, 1858
Nomenclatural Authority: Newton et al. (2000)
Notes. As many as nine species of Hydrosmecta may occur in California (M. L. Gimmel & M. S. Caterino, 2022, personal data), but the genus has not been recently revised. Several may not be congeneric, and some may be synonyms (Seevers, 1978).
Hydrosmecta undetermined species
Literature Records: none
Digitized Records: Santa Cruz (20 SBMNH)
Pontomalota Casey, 1885
Nomenclatural Authority: Newton et al. (2000)
Notes. Two species of Pontomalota have been recorded from California (Ahn & Ashe, 1992). The species of this genus were revised by Ahn & Ashe (1992).
Pontomalota opaca (LeConte, 1863)
Nomenclatural Authority: Ahn & Ashe (1992)
Literature Records: San Miguel (Ahn & Ashe, 1992: 352)
Digitized Records: San Miguel (30 SBMNH), San Nicolas (6 SBMNH), Santa Rosa (7 SBMNH)
Range: Also known from mainland (Ahn & Ashe, 1992).
“Sonomota” Casey, 1911
Nomenclatural Authority: Newton et al. (2000)
Notes. Currently this genus-group name is a synonym of Atheta (Microdota) Mulsant & Rey, 1873, not of Geostiba Thomson, 1858 as listed by Newton et al. (2000) (see Gusarov, 2002). However, it is being used here to designate a distinctive group of West Coast athetines (V. Gusarov, 2022, personal communication).
Sonomota undetermined species
Literature Records: none
Digitized Records: San Clemente (7 SBMNH), Santa Cruz (7 SBMNH)
Tarphiota Casey, 1893
Nomenclatural Authority: Newton et al. (2000)
Notes. Two species of Tarphiota have been recorded from California (Ahn, 1996b; Klimaszewski, Majka & Langor, 2006). The North American species were revised by Ahn (1996b); Klimaszewski, Majka & Langor (2006) provided an update.
Tarphiota fucicola (Mäklin, 1852)
Nomenclatural Authority: Ahn (1996b)
Literature Records: none
Digitized Records: San Miguel (24 SBMNH), San Nicolas (5 SBMNH), Santa Cruz (29 SBMNH), Santa Rosa (16 SBMNH)
Range: Also known from mainland (Ahn, 1996b).
Tarphiota geniculata (Mäklin, 1852)
Nomenclatural Authority: Ahn (1996b)
Literature Records: none
Digitized Records: San Clemente (9 SBMNH), San Miguel (17 SBMNH), San Nicolas (5 SBMNH), Santa Catalina (10 SBMNH), Santa Cruz (30 SBMNH), Santa Rosa (13 SBMNH)
Range: Also known from mainland (Ahn, 1996b).
Thinusa Casey, 1893
Nomenclatural Authority: Newton et al. (2000)
Digitized Records: Santa Cruz (1 UCRC)
Notes. Two species of Thinusa have been recorded from California (Ahn, 1997). The species of this genus were revised by Ahn (1997).
Thinusa fletcheri Casey, 1906
Nomenclatural Authority: Ahn (1997)
Literature Records: none
Digitized Records: San Clemente (2 SBMNH), San Nicolas (1 SBMNH), Santa Catalina (1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Ahn, 1997).
Thinusa maritima (Casey, 1885)
Nomenclatural Authority: Ahn (1997)
Literature Records: Santa Cruz (Ahn, 1997: 80)
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Ahn, 1997).
Falagriini
Notes. Six genera and seven species of Falagriini have been recorded from California (M. L. Gimmel, 2022, unpublished data). The species of Falagriini were revised for North America by Hoebeke (1985).
Falagriota Casey, 1906
Nomenclatural Authority: Newton et al. (2000)
Notes. One species of Falagriota has been recorded from California (Hoebeke, 1985).
Falagriota occidua (Casey, 1885)
Nomenclatural Authority: Hoebeke (1985)
Literature Records: Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: Santa Cruz (11 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Hoebeke, 1985).
Homalotini
Notes. Nine genera and 32 species of Homalotini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Diestota Mulsant & Rey, 1871
Nomenclatural Authority: Newton et al. (2000)
Notes. Two species, Diestota angustula (Casey, 1906) and Diestota spissula (Casey, 1911), have been recorded from California (Seevers, 1978, as Apheloglossa Casey, 1893).
Diestota undetermined species
Literature Records: none
Digitized Records: San Clemente (3 SBMNH), San Miguel (14 SBMNH), San Nicolas (4 SBMNH), Santa Catalina (2 SBMNH), Santa Cruz (2 SBMNH), Santa Rosa (1 SBMNH)
Stictalia Casey, 1906
Nomenclatural Authority: Newton et al. (2000)
Notes: The sixteen North American species of Stictalia are all western (Newton et al., 2000). Of these, 12 have been recorded from California, but essentially only from their original type localities (Seevers, 1978).
Stictalia undetermined species
Literature Records: none
Digitized Records: San Clemente (36 SBMNH), Santa Cruz (5 SBMNH), Santa Rosa (15 SBMNH)
Hypocyphtini
Notes. Three genera and five species of Hypocyphtini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Holobus Solier, 1849
Nomenclatural Authority: Newton et al. (2000)
Notes: Only one species, Holobus oviformis Casey, 1893, has been recorded from California, from Los Angeles and San Diego counties. It is likely that the undetermined Santa Catalina Island specimen below is referable to this species.
Holobus undetermined species
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH)
Oligota Mannerheim, 1830
Nomenclatural Authority: Newton et al. (2000)
Notes: Three of the 10 North American species of Oligota have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Oligota undetermined species
Literature Records: none
Digitized Records: San Nicolas (1 SBMNH), Santa Cruz (4 SBMNH), Santa Rosa (15 SBMNH)
Liparocephalini
Notes. Four genera and seven species of Liparocephalini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Diaulota Casey, 1893
Nomenclatural Authority: Newton et al. (2000)
Notes. Four species of Diaulota have been recorded from California (Ahn, 1996a). The species of this genus were reviewed by Ahn (1996a).
Diaulota fulviventris Moore, 1956
Nomenclatural Authority: Ahn (1996a)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Ahn, 1996a).
Myllaenini
Notes. Two genera and eight species of Myllaenini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Bryothinusa Casey, 1904
Nomenclatural Authority: Newton et al. (2000)
Notes. One species of Bryothinusa has been recorded from California (Moore & Orth, 1979).
Bryothinusa catalinae Casey, 1904
Nomenclatural Authority: Moore & Orth (1979)
Literature Records: Santa Catalina (Casey, 1904: 313; Baker, 1905: 57; Moore, 1956: 132)
Digitized Records: Santa Catalina (8 USNM)
Range: Also known from mainland (Moore, 1956; Moore & Orth, 1979).
Notes. Moore & Orth (1979) described the larva of this odd seashore species. The species was considered endemic to Santa Catalina Island until Moore (1956) reported on mainland specimens.
Myllaena Erichson, 1837
Nomenclatural Authority: Newton et al. (2000)
Notes. Seven species of Myllaena have been reported from California (Klimaszewski, 1982). At least four of these occur in coastal southern California, potentially conspecific with Myllaena from the Channel Islands. The species were revised for North America by Klimaszewski (1982).
Myllaena undetermined species
Literature Records: none
Digitized Records: San Miguel (3 SBMNH), Santa Catalina (1 SBMNH), Santa Cruz (11 SBMNH), Santa Rosa (1 SBMNH)
Oxypodini
Notes. Twenty genera and 78 species of Oxypodini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Meoticina undetermined genus and species
Literature Records: none
Digitized Records: Santa Cruz (8 SBMNH)
Notes: Four genera of this oxypodine subtribe are known from California, Gyronycha Casey, 1893, Alisalia Casey, 1911, Apimela Mulsant & Rey, 1874, and Bamona Sharp, 1883 (Newton et al., 2000), though none, yet, from the Channel Islands. This record is certainly referable to one of them.
Blepharhymenus Solier, 1849
Nomenclatural Authority: Newton et al. (2000)
Notes: Nineteen species of Blepharhymenus have been described from California (Seevers, 1978), all in need of revision.
Blepharhymenus undetermined species
Literature Records: none
Digitized Records: San Clemente (2 SBMNH), Santa Cruz (2 SBMNH), Santa Rosa (10 SBMNH)
Oxypoda Mannerheim, 1830
Nomenclatural Authority: Newton et al. (2000)
Notes. Twenty-eight species of Oxypoda have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Oxypoda undetermined species
Literature Records: none
Digitized Records: Santa Catalina (4 SBMNH), Santa Cruz (10 SBMNH), Santa Rosa (15 SBMNH)
Phloeopora Erichson, 1837
Nomenclatural Authority: Newton et al. (2000)
Notes: Two of the eight North American species of the cosmopolitan genus Phloeopora have been reported from California (Seevers, 1978).
Phloeopora undetermined species
Literature Records: none
Digitized Records: Santa Cruz (3 SBMNH)
Tachyusini
Notes. Seven genera and 27 species of Tachyusini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Gnypeta Thomson, 1858
Nomenclatural Authority: Newton et al. (2000)
Notes: Eighteen species of this large, widespread genus have been reported from California (Seevers, 1978), but they are in need of revision.
Gnypeta undetermined species
Literature Records: none
Digitized Records: Santa Cruz (4 SBMNH), Santa Rosa (2 SBMNH)
Habrocerinae
Notes. The subfamily Habrocerinae, which contains one genus and species in California, was revised by Assing & Wunderle (1995).
Habrocerus Erichson, 1839
Nomenclatural Authority: Herman (2001)
Notes. One adventive species of Habrocerus occurs in California (Herman, 2001).
Habrocerus capillaricornis (Gravenhorst, 1806)
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Assing & Wunderle, 1995).
Notes. This species was introduced to North America from the Western Palearctic (Assing & Wunderle, 1995).
Leptotyphlinae
Notes. The subfamily Leptotyphlinae contains seven genera and 12 described species in California (M. L. Gimmel, 2022, unpublished data).
Leptotyphlinae undetermined genus and species
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH).
Notes. The two specimen lots recorded above from SBMNH are currently housed in ethanol tubes and collectively represent >30 specimens. A large number of undescribed species of Leptotyphlinae are known to occur in California (V. Gusarov, 2022, personal communication); the Channel Island specimens are almost certainly undescribed.
Mycetoporinae
Notes. This subfamily was recently split from Tachyporinae by Yamamoto (2021). It contains nine genera and 29 species in California (M. L. Gimmel, 2022, unpublished data).
Bryoporus Kraatz, 1857
Nomenclatural Authority: Herman (2001)
Notes. One species of Bryoporus has been recorded from California (Campbell, 1993b). The genus was revised for North America by Campbell (1993b).
Bryoporus rufescens LeConte, 1863
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: San Clemente (32 SBMNH), Santa Cruz (2 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Campbell, 1993b).
Lordithon Thomson, 1859
Nomenclatural Authority: Herman (2001)
Notes. Four species of Lordithon have been recorded from California (Campbell, 1982). This genus was revised for North America by Campbell (1982). Additional species are likely to occur on the Channel Islands.
Lordithon thoracicus (Fabricius, 1777)
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Catalina (2 SBMNH), Santa Cruz (7 SBMNH)
Range: Also known from mainland (Campbell, 1982).
Notes. The nominate subspecies, L. t. thoracicus (Fabricius, 1777), is the only subspecies occurring in California (Campbell, 1982). The species has a Holarctic distribution (Campbell, 1982).
Mycetoporus Mannerheim, 1830
Nomenclatural Authority: Herman (2001)
Notes. Six species of Mycetoporus have been recorded from California (Campbell, 1991). The genus was revised for North America by Campbell (1991). Additional species of Mycetoporus possibly occur on the Channel Islands.
Mycetoporus neotomae Fall, 1910
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (5 SBMNH)
Range: Also known from mainland (Campbell, 1991).
Omaliinae: Omaliini
Notes. Five tribes, 33 genera, and 95 species of Omaliinae are known to occur in California, with 10 genera and 22 species of Omaliini known to occur in the state (Herman, 2001; M. L. Gimmel, 2022, unpublished data).
Omalium Gravenhorst, 1802
Nomenclatural Authority: Herman (2001)
Notes. Eight species of Omalium have been recorded from California (Herman, 2001).
Omalium algarum Casey, 1885
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: San Nicolas (1 LACM)
Range: Also known from mainland (Frank & Ahn, 2011).
Notes. One of the few seashore-inhabiting species of the genus, O. algarum occurs along much of the West Coast of North America, from southern California to British Columbia (Frank & Ahn, 2011).
Oxytelinae
Notes. Four tribes, 13 genera, and 79 species of Oxytelinae are known to occur in California (Herman, 2001; M. L. Gimmel, 2022, unpublished data). “Oxytelinae” was reported from Santa Cruz Island by Straughan & Hadley (1980: 392).
Blediini
Notes. One genus and 34 species of Blediini have been recorded from California (Herman, 2001).
Bledius Leach, 1819
Nomenclatural Authority: Herman (2001)
Literature Records (genus-only): Santa Catalina (Straughan & Hadley, 1980: 392)
Digitized Records (genus-only): Santa Cruz (1 SBMNH)
Notes. Thirty-four species of Bledius have been recorded from California (Herman, 2001). The species were revised in a series of papers by Herman (1972, 1976, 1983, 1986).
Bledius albonotatus Mäklin, 1853
Nomenclatural Authority: Herman (2001)
Literature Records: San Miguel (Herman, 1983: 123), San Nicolas (Herman, 1983: 123)
Digitized Records: San Miguel (16 SBMNH), San Nicolas (3 SBMNH), Santa Catalina (1 SBMNH), Santa Cruz (3 SBMNH), Santa Rosa (11 SBMNH)
Range: Also known from mainland (Herman, 1983).
Bledius fenyesi Bernhauer & Schubert, 1911
Nomenclatural Authority: Herman (2001)
Literature Records: San Miguel (Herman, 1976: 164), San Nicolas (Herman, 1976: 164), Santa Cruz (Herman, 1976: 164)
Digitized Records: San Clemente (7 SBMNH), San Miguel (38 SBMNH), San Nicolas (3 SBMNH), Santa Catalina (7 SBMNH), Santa Cruz (19 SBMNH), Santa Rosa (10 SBMNH)
Range: Also known from mainland (Herman, 1976).
Bledius opacifrons LeConte, 1877
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Cruz (6 SBMNH), Santa Rosa (16 SBMNH)
Range: Also known from mainland (Herman, 1976).
Bledius ruficornis LeConte, 1863
Nomenclatural Authority: Herman (2001)
Literature Records: San Clemente (Herman, 1983: 128)
Digitized Records: San Clemente (1 SBMNH), Santa Cruz (2 SBMNH), Santa Rosa (11 SBMNH)
Range: Also known from mainland (Herman, 1983).
Oxytelini
Notes. Ten genera and 42 species of Oxytelini have been recorded from California (Herman, 2001; M. L. Gimmel, 2022, unpublished data).
Aploderus Stephens, 1833
Nomenclatural Authority: Herman (2001)
Digitized Records (genus-only): Santa Cruz (15 SBMNH), Santa Rosa (20 SBMNH)
Notes. Seven species of Aploderus have been recorded from California (Herman, 2001).
Aploderus flavipennis Casey, 1889
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (Herman, 2001).
Notes. Reported by Fall (1897) as Haploderus flavipennis.
Apocellus Erichson, 1839
Nomenclatural Authority: Herman (2001)
Notes. Three species of Apocellus have been recorded from California (Herman, 2001).
Apocellus analis LeConte, 1877
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (Herman, 2001).
Carpelimus Leach, 1819
Nomenclatural Authority: Herman (2001)
Notes. Thirteen species of Carpelimus have been recorded from California (Herman, 2001).
Carpelimus undetermined species
Literature Records: none
Digitized Records: San Clemente (1 SBMNH), San Nicolas (3 SBMNH), Santa Catalina (2 SBMNH), Santa Cruz (19 SBMNH), Santa Rosa (2 SBMNH)
Platystethus Mannerheim, 1830
Nomenclatural Authority: Herman (2001)
Notes. Two species of Platystethus have been recorded from California (Moore & Legner, 1971). The North American species were reviewed by Moore & Legner (1971).
Platystethus americanus Erichson, 1840
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Rosa (3 SBMNH)
Range: Also known from mainland (Moore & Legner, 1971).
Thinobius Kiesenwetter, 1844
Nomenclatural Authority: Herman (2001)
Notes. Six species of Thinobius have been recorded from California (Herman, 2001).
Thinobius undetermined species
Literature Records: none
Digitized Records: San Nicolas (10 SBMNH), Santa Cruz (8 SBMNH), Santa Rosa (41 SBMNH)
Paederinae: Paederini
Notes. Two tribes, 19 genera, and 101 species of Paederinae are known to occur in California, of which 17 genera and 99 species belong to Paederini (M. L. Gimmel, 2022, unpublished data).
Astenus Dejean, 1833
Nomenclatural Authority: Newton et al. (2000)
Notes. This genus contains 24 species in North America (Newton et al., 2000); four of these have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Astenus undetermined species
Literature Records: none
Digitized Records: San Clemente (1 SBMNH), Santa Cruz (2 SBMNH)
Lobrathium Mulsant & Rey, 1878
Nomenclatural Authority: Newton et al. (2000)
Digitized Records (genus-only): Santa Cruz (35 SBMNH)
Notes. Seven species of Lobrathium have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Lobrathium jacobinum (LeConte, 1863)
Nomenclatural Authority: Casey (1905)
Literature Records: Santa Rosa (Fall, 1897: 237)
Range: Also known from mainland (Casey, 1905).
Notes. Reported by Fall (1897) as Lathrobium jacobinum.
Medon Stephens, 1833
Nomenclatural Authority: Newton et al. (2000)
Notes. Twenty-eight species of Medon have been recorded from California (M. L. Gimmel, 2022, unpublished data). Naughton et al. (2014) reported two separate, unidentified species of Medon from Santa Cruz Island.
Medon undetermined species
Literature Records: Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: San Clemente (18 SBMNH), San Miguel (3 SBMNH), Santa Catalina (4 SBMNH), Santa Cruz (80 SBMNH), Santa Rosa (51 SBMNH)
Orus Casey, 1884
Nomenclatural Authority: Newton et al. (2000)
Notes. Twelve species of Orus have been recorded from California (Herman, 1964, 1965; Moore & Legner, 1972). The species of this genus were revised by Herman (1964, 1965), with a modification by Moore & Legner (1972).
Orus undetermined species
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Notes. The one SBMNH specimen from Santa Cruz Island cited above appears to be a female, and therefore not determinable to species using the keys of Herman (1964, 1965).
Sunius Stephens, 1829
Nomenclatural Authority: Newton et al. (2000)
Literature Records (genus-only): Santa Barbara (Miller & Miller, 1985: 124), Santa Catalina (Fall, 1897: 237)
Notes. Fourteen species of Sunius have been recorded from California (M. L. Gimmel, 2022, unpublished data). The genus-only record reported by Fall (1897) as “Caloderma sp.” was presumably a species different from S. mobilis or S. reductus, which were reported in the same publication. The former record is presumably in reference to a specimen in the Museum of Comparative Zoology, Harvard University that was later identified by H.C. Fall as Sunius exilis (Casey, 1905) (S. Miller, 2022, personal communication). The species reported by Miller & Miller (1985) was said to be near Sunius cuneicollis (Casey, 1886); they noted that the genus needed revision.
Sunius mobilis (Casey, 1886)
Nomenclatural Authority: Casey (1905), Newton et al. (2000)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (Casey, 1905).
Notes. Reported by Fall (1897) as Caloderma mobile.
Sunius reductus (Casey, 1886)
Nomenclatural Authority: Casey (1905), Newton et al. (2000)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (Casey, 1905).
Notes. Reported by Fall (1897) as Caloderma reductum.
Pselaphinae
Notes. Twelve tribes, 35 genera, and 255 species of Pselaphinae are known to occur in California (Chandler, 1997; M. L. Gimmel, 2022, unpublished data).
Euplectini
Notes. Eleven genera and 58 species of Euplectini have been recorded from California (Chandler, 1997; M. L. Gimmel, 2022, unpublished data).
Actium Casey, 1886
Nomenclatural Authority: Chandler (1997)
Notes. Twenty-eight species of Actium have been recorded from California (Chandler, 1997; Caterino & Chandler, 2010). The genus was revised by Grigarick & Schuster (1971), with an addition by Caterino & Chandler (2010).
Actium californicum (LeConte, 1878)
Nomenclatural Authority: Grigarick & Schuster (1971)
Literature Records: Santa Cruz (Grigarick & Schuster, 1971: 26; Caterino & Chandler, 2010: 191)
Digitized Records: none
Range: Also known from mainland (Grigarick & Schuster, 1971).
Notes. Grigarick & Schuster (1971) reported that most specimens of this species collected on Santa Cruz Island were taken by stripping bark of fallen Quercus agrifolia Née (Fagaceae) lying close to small, intermittent streams during the spring.
Actium vestigialis Caterino & Chandler, 2010
Nomenclatural Authority: Caterino & Chandler (2010)
Literature Records: Santa Catalina (Caterino & Chandler, 2010: 188)
Digitized Records: Santa Catalina (5 SBMNH)
Range: Endemic (Caterino & Chandler, 2010).
Faronini
Notes. Two genera and 34 species of Faronini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Sonoma Casey, 1886
Nomenclatural Authority: Chandler (1997)
Literature Records (genus-only): San Clemente (Caterino & Chandler, 2010: 191), Santa Catalina (Caterino & Chandler, 2010: 191), Santa Cruz (Caterino & Chandler, 2010: 191)
Digitized Records (genus-only): San Clemente (27 SBMNH), Santa Catalina (6 SBMNH), Santa Cruz (1 SBMNH)
Notes. Twenty-eight species of Sonoma have been recorded from California (Ferro, 2016). The genus was revised for western North America by Ferro (2016). Caterino & Chandler (2010) reported occurrence of this genus in leaf litter on San Clemente, Santa Catalina, and Santa Cruz islands. The records of the former two, and perhaps all three, islands presumably refer to S. isabellae.
Sonoma isabellae (LeConte, 1851)
Nomenclatural Authority: Ferro (2016)
Literature Records: San Clemente (Ferro, 2016: 49), Santa Catalina (Ferro, 2016: 49)
Digitized Records: San Clemente (7 SBMNH), Santa Catalina (5 SBMNH)
Range: Also known from mainland (Ferro, 2016).
Trogastrini
Notes. Three genera and 37 species of Trogastrini have been recorded from California (Chandler, 1997; M. L. Gimmel, 2022, unpublished data).
Oropus Casey, 1886
Nomenclatural Authority: Chandler (1997)
Notes. Twenty-seven species of Oropus have been recorded from California (Chandler, 1997). The genus was revised by Schuster & Grigarick (1960).
Oropus undetermined species
Literature Records: Santa Catalina (Caterino & Chandler, 2010: 191), Santa Cruz (Caterino & Chandler, 2010: 191)
Digitized Records: Santa Cruz (2 SBMNH)
Notes. Caterino & Chandler (2010) reported occurrence of the genus Oropus in leaf litter on Santa Catalina and Santa Cruz islands.
Tychini
Notes. Two genera and 22 species of Tychini have been recorded from California (Chandler, 1997; M. L. Gimmel, 2022, unpublished data).
Hesperotychus Schuster & Marsh, 1958
Nomenclatural Authority: Chandler (1997)
Notes. Thirteen species of Hesperotychus have been recorded from California (M. L. Gimmel, 2022, unpublished data). The species were revised by Schuster & Marsh (1958).
Hesperotychus undetermined species
Literature Records: Santa Catalina (Caterino & Chandler, 2010: 191)
Digitized Records: Santa Catalina (1 SBMNH)
Notes. Caterino & Chandler (2010) reported occurrence of this genus in leaf litter on Santa Catalina Island.
Pseudopsinae
Notes. This subfamily is represented by four genera and eight species in California (Herman, 2001). So far only Pseudopsis has been identified from the Channel Islands, though Nanobius serricollis (LeConte, 1875), which occurs in the southern California coast ranges (Herman, 1977), may also occur there.
Pseudopsis Newman, 1834
Nomenclatural Authority: Herman (2001)
Literature Records (genus-only): Santa Catalina (Fall, 1897: 237)
Notes. Four species of Pseudopsis have been recorded from California (Herman, 2001). Fall (1897: 237) reported an undetermined species of this genus from Santa Catalina Island. This may not represent P. minuta, as he later (Fall, 1901: 227) described that species only from “the cañons of the southern Sierras”. This could represent either an undescribed species or the later-described P. montoraria Herman, 1975, which occurs in the coastal mountain ranges of southern California and has not yet been identified from the Channel Islands. Herman (1975) revised the genus.
Pseudopsis minuta Fall, 1901
Nomenclatural Authority: Herman (1975, 2001)
Literature Records: Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: Santa Cruz (15 SBMNH)
Range: Also known from mainland (Herman, 1975).
Notes. Naughton et al. (2014: 304) identified a single specimen only to “Pseudopsis sp.”. The voucher in SBMNH was examined and this represents P. minuta.
Scydmaeninae
Notes. Five tribes, 13 genera, and 57 species of Scydmaeninae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Cephenniini
Notes. Two genera and nine species of Cephenniini are known to occur in California (Hopp & Caterino, 2009; M. L. Gimmel, 2022, unpublished data).
Cephennium Müller & Kunze, 1822
Nomenclatural Authority: O’Keefe (2000)
Notes. Eight species of Cephennium have been recorded from California (Hopp & Caterino, 2009). The Californian species were revised by Hopp & Caterino (2009).
Cephennium urbanum Hopp & Caterino, 2009
Nomenclatural Authority: Hopp & Caterino (2009)
Literature Records: Santa Catalina (Caterino & Chandler, 2010: 191)
Digitized Records: Santa Catalina (9 SBMNH)
Range: Also known from mainland (Hopp & Caterino, 2009).
Notes. Only the genus Cephennium was reported from Santa Catalina Island by Caterino & Chandler (2010); that report presumably referred to this species.
Glandulariini
Notes. Seven genera and 41 species of Glandulariini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Brachycepsis Brendel, 1889
Nomenclatural Authority: O’Keefe (2000)
Notes. Two species of Brachycepsis have been recorded from California (M. L. Gimmel, 2022, unpublished data). Species of this genus are currently unidentifiable, as there are several undescribed species and the last treatment is over 120 years old (see O’Keefe, 2000).
Brachycepsis undetermined species
Literature Records: Santa Catalina (Caterino & Chandler, 2010: 191), Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: Santa Cruz (4 SBMNH), Santa Rosa (4 SBMNH)
Euconnus Thomson, 1862
Nomenclatural Authority: O’Keefe (2000)
Notes. Seven species of Euconnus have been recorded from California, belonging to two subgenera, Drastophus Casey, 1897 and Napochus Thomson, 1862 (M. L. Gimmel, 2022, unpublished data).
Euconnus undetermined species
Literature Records: none
Digitized Records: San Clemente (1 SBMNH), Santa Catalina (3 SBMNH), Santa Rosa (24 SBMNH)
Notes. All members of Euconnus observed from the Channel Islands belong to the subgenus Drastophus. Based on morphology, there are at least two species represented among this material, and probably more.
Stenichnus Thomson, 1859
Nomenclatural Authority: O’Keefe (2000)
Notes. There are several described species of this genus in North America, and the latest treatment is over 120 years old (see O’Keefe, 2000). Undescribed species probably exist.
Stenichnus undetermined species
Literature Records: Santa Catalina (Caterino & Chandler, 2010: 191)
Digitized Records: Santa Catalina (3 SBMNH)
Silphinae
Notes. Long known as the family Silphidae, this group was recently made a subfamily of Staphylinidae (Cai et al., 2022). There are two tribes, four genera, and nine species of Silphinae recorded from California (Miller & Peck, 1979; Peck & Miller, 1993). Miller & Peck (1979) provided a guide to the group for California, while Peck & Miller (1993) provided a catalog for North America. Members of this subfamily have been extensively surveyed and investigated for the Channel Islands, and it is doubtful that additional species will be discovered there.
Nicrophorini
Notes. One genus and four species of Nicrophorini have been recorded from California (Peck & Miller, 1993).
Nicrophorus Fabricius, 1775
Nomenclatural Authority: Sikes, Madge & Newton (2002)
Digitized Records (genus-only): Santa Cruz (2 EMEC)
Notes. Four species of Nicrophorus have been recorded from California (Peck & Miller, 1993).
Nicrophorus guttula Motschulsky, 1845
Nomenclatural Authority: Sikes, Madge & Newton (2002)
Literature Records: San Clemente (Fall, 1897: 236; Fall, 1901: 58; Miller & Peck, 1979: 97; Anderson & Peck, 1986: 296; Peck & Kaulbars, 1988: 72 [map])
Digitized Records: San Clemente (1 LACM), Santa Catalina (2 LACM), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Fall, 1901; Miller & Peck, 1979; Anderson & Peck, 1986; Peck & Kaulbars, 1988; Peck & Miller, 1993; Sikes, Madge & Newton, 2002).
Nicrophorus marginatus Fabricius, 1801
Nomenclatural Authority: Sikes, Madge & Newton (2002)
Literature Records: none
Digitized Records: San Miguel (1 LACM)
Range: Also known from mainland (Fall, 1901; Miller & Peck, 1979; Anderson & Peck, 1986; Peck & Kaulbars, 1988; Peck & Miller, 1993; Sikes, Madge & Newton, 2002).
Notes. While this is the most widespread species of Nicrophorus in North America, only one specimen has been reported from the Channel Islands, collected in 1985.
Nicrophorus nigrita Mannerheim, 1843
Nomenclatural Authority: Sikes, Madge & Newton (2002)
Literature Records: Anacapa (Miller & Peck, 1979: 96; Miller & Miller, 1985: 124), San Clemente (Fall, 1897: 236; Miller & Peck, 1979: 96; Miller & Miller, 1985: 124; Peck & Kaulbars, 1988: 69 [map]), Santa Barbara (Miller & Peck, 1979: 96; Anderson, 1982: 262; Miller & Miller, 1985: 124), Santa Catalina (Miller & Miller, 1985: 124), Santa Cruz (Miller & Peck, 1979: 96; Miller & Miller, 1985: 124; Peck & Kaulbars, 1988: 69 [map]), Santa Rosa (Fall, 1897: 236; Miller & Miller, 1985: 124; Sikes, Madge & Newton, 2002: 139)
Digitized Records: Santa Catalina (1 DMNS; 6 LACM; 1 SBMNH), Santa Cruz (4 LACM; 15 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Fall, 1901; Miller & Peck, 1979; Peck & Kaulbars, 1988; Peck & Miller, 1993; Sikes, Madge & Newton, 2002).
Notes. Fall (1901) recorded this species as Necrophorus pustulatus var. nigritus, and reported it from “both islands” (presumably San Clemente and Santa Rosa). Miller & Miller (1985) recorded it “under dead mice and at lanterns” on Santa Barbara Island.
Silphini
Notes. Three genera and five species of Silphini have been recorded from California (Miller & Peck, 1979; Peck & Miller, 1993).
Heterosilpha Portevin, 1926
Nomenclatural Authority: Peck & Miller (1993)
Notes. Two species of Heterosilpha have been recorded from California (Peck & Miller, 1993).
Heterosilpha ramosa (Say, 1823)
Nomenclatural Authority: Peck & Miller (1993)
Literature Records: San Miguel (Miller & Peck, 1979: 93), Santa Cruz (Fall & Davis, 1934: 144; Miller & Peck, 1979: 93; Peck & Kaulbars, 1988: 57 [map]), Santa Rosa (Fall, 1897: 236; Miller & Peck, 1979: 93)
Digitized Records: San Clemente (1 UCRC), Santa Cruz (2 LACM; 11 SBMNH; 23 TAMU; 2 UCRC), Santa Rosa (24 LACM; 11 SBMNH)
Range: Also known from mainland (Miller & Peck, 1979; Peck & Kaulbars, 1988; Peck & Miller, 1993).
Notes. Fall (1897) and Fall & Davis (1934) recorded this species as Silpha ramosa.
Thanatophilus Leach, 1815
Nomenclatural Authority: Peck & Miller (1993)
Notes. Two species of Thanatophilus have been recorded from California (Miller & Peck, 1979).
Thanatophilus lapponicus (Herbst, 1793)
Nomenclatural Authority: Peck & Miller (1993)
Literature Records: Santa Rosa (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Miller & Peck, 1979; Peck & Kaulbars, 1988; Peck & Miller, 1993).
Notes. Fall (1897) recorded this species as Silpha lapponica. Miller & Peck (1979: 91) stated that they had not seen Channel Island specimens of this species, and speculated that Fall’s specimens may have been destroyed in the 1906 San Francisco fire.
Staphylininae
Notes. Three tribes, 35 genera, and 232 species of Staphylininae are known to occur in California (Herman, 2001; M. L. Gimmel, 2022, unpublished data).
Staphylinini: Amblyopinina
Notes. Seven subtribes, 20 genera, and 171 species of Staphylinini are known to occur in California, of which one genus and 10 species belong to Amblyopinina (Herman, 2001; M. L. Gimmel, 2022, unpublished data).
Heterothops Stephens, 1829
Nomenclatural Authority: Herman (2001)
Digitized Records (genus-only): Santa Catalina (1 SBMNH), Santa Cruz (1 SBMNH), Santa Rosa (19 SBMNH)
Notes. Ten species of Heterothops have been recorded from California (Herman, 2001). The species were revised for North America by Smetana (1971).
Heterothops conformis Smetana, 1971
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (43 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Smetana, 1971).
Heterothops fusculus LeConte, 1863
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Smetana, 1971).
Notes. This species was recorded by Fall (1897) as Heterothops californicus LeConte, 1863, now considered a synonym of H. californicus (see Smetana, 1971: 26).
Staphylinini: Erichsoniina
Notes. One genus and two species of Erichsoniina have been recorded from California (Herman, 2001).
Erichsonius Fauvel, 1874
Nomenclatural Authority: Herman (2001)
Notes. Two species of Erichsonius have been recorded from California (Herman, 2001). The species were revised for the New World by Frank (1975).
Erichsonius puncticeps (Horn, 1884)
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: San Miguel (1 SBMNH), Santa Cruz (3 SBMNH), Santa Rosa (9 SBMNH)
Range: Also known from mainland (Frank, 1975).
Notes. Fall (1897) recorded this species as Actobius puncticeps.
Staphylinini: Philonthina
Notes. Six genera and 100 species of Philonthina are known to occur in California (Herman, 2001; M. L. Gimmel, 2022, unpublished data).
Belonuchus Nordmann, 1837
Nomenclatural Authority: Herman (2001)
Notes. Three species of Belonuchus have been recorded from California (Herman, 2001). The species were revised for North America by Smetana (1995).
Belonuchus ephippiatus (Say, 1830)
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: San Miguel (2 SBMNH), Santa Catalina (3 SBMNH)
Range: Also known from mainland (Smetana, 1995).
Bisnius Stephens, 1829
Nomenclatural Authority: Herman (2001)
Notes. Thirteen species of Bisnius have been recorded from California (Herman, 2001). The species were revised for North America by Smetana (1995).
Bisnius albionicus (Mannerheim, 1843)
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: San Miguel (7 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Smetana, 1995).
Bisnius sordidus (Gravenhorst, 1802)
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: San Nicolas (1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Smetana, 1995).
Notes. The distribution map in Smetana (1995: 526) shows a record from either San Miguel or Santa Rosa. This species was introduced to North America from the Palearctic realm (Smetana, 1995).
Cafius Curtis, 1829
Nomenclatural Authority: Herman (2001)
Notes. Eight species of Cafius have been recorded from California (Herman, 2001). The species were revised for the west coast of North America by Orth & Moore (1980).
Cafius canescens (Mäklin, 1852)
Nomenclatural Authority: Herman (2001)
Literature Records: San Nicolas (Fall, 1897: 236; Cockerell, 1940: 285), Santa Catalina (Orth & Moore, 1980: 186)
Digitized Records: San Miguel (34 LACM; 46 SBMNH), San Nicolas (76 LACM; 2 SBMNH), Santa Cruz (7 SBMNH), Santa Rosa (7 LACM; 15 SBMNH; 2 UTCI)
Range: Also known from mainland (Orth & Moore, 1980).
Cafius lithocharinus (LeConte, 1863)
Nomenclatural Authority: Herman (2001)
Literature Records: San Nicolas (Straughan & Hadley, 1980: 392), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Clemente (9 SBMNH), San Miguel (1 SBMNH), San Nicolas (5 SBMNH), Santa Catalina (3 SBMNH), Santa Cruz (4 SBMNH), Santa Rosa (47 SBMNH)
Range: Also known from mainland (Orth & Moore, 1980).
Cafius luteipennis Horn, 1884
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Orth & Moore, 1980: 192), Santa Rosa (Fall, 1897: 236)
Digitized Records: San Clemente (7 SBMNH), San Miguel (6 SBMNH), San Nicolas (5 SBMNH), Santa Catalina (5 SBMNH), Santa Cruz (14 SBMNH), Santa Rosa (11 SBMNH)
Range: Also known from mainland (Orth & Moore, 1980).
Cafius opacus (LeConte, 1863)
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (Orth & Moore, 1980).
Cafius seminitens Horn, 1884
Nomenclatural Authority: Herman (2001)
Literature Records: San Miguel (Cockerell, 1940: 285; Orth & Moore, 1980: 185), San Nicolas (Cockerell, 1940: 285)
Digitized Records: San Clemente (6 SBMNH), San Miguel (10 LACM; 21 SBMNH), San Nicolas (64 LACM; 9 SBMNH), Santa Cruz (1 LACM; 3 SBMNH), Santa Rosa (7 LACM; 12 SBMNH; 2 UTCI)
Range: Also known from mainland (Orth & Moore, 1980).
Cafius sulcicollis (LeConte, 1863)
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Cruz (Orth & Moore, 1980: 195), Santa Rosa (Fall, 1897: 237)
Digitized Records: San Clemente (2 SBMNH), Santa Rosa (6 SBMNH)
Range: Also known from mainland (Orth & Moore, 1980).
Gabrius Stephens, 1829
Nomenclatural Authority: Herman (2001)
Digitized Records (genus-only): San Nicolas (1 SBMNH), Santa Cruz (1 SBMNH)
Notes. Sixteen species of Gabrius have been recorded from California (Herman, 2001). The genus was revised for North America by Smetana (1995).
Gabrius nigritulus (Gravenhorst, 1802)
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: San Nicolas (4 SBMNH)
Range: Also known from mainland (Smetana, 1995).
Notes. This species was recorded as Philonthus nigritulus by Fall (1897). It was introduced to North America from the western Palearctic realm (Smetana, 1995).
Neobisnius Ganglbauer, 1895
Nomenclatural Authority: Herman (2001)
Notes. Eight species of Neobisnius have been recorded from California (Herman, 2001). The species were revised for the New World by Frank (1981).
Neobisnius occidentoides Frank, 1981
Nomenclatural Authority: Herman (2001)
Literature Records: San Clemente (Frank, 1981: 49)
Digitized Records: San Clemente (7 SBMNH; 4 UTCI), Santa Catalina (2 SBMNH), Santa Cruz (1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Frank, 1981).
Neobisnius sobrinus (Erichson, 1840)
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Frank, 1981).
Neobisnius terminalis (LeConte, 1863)
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Cruz (3 SBMNH)
Range: Also known from mainland (Frank, 1981).
Philonthus Stephens, 1829
Nomenclatural Authority: Herman (2001)
Notes. Fifty-two species of Philonthus have been recorded from California (M. L. Gimmel, 2022, unpublished data). The species were revised for North America by Smetana (1995).
Philonthus cruentatus (Gmelin, 1790)
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Cockerell, 1940: 285)
Digitized Records: Santa Barbara (1 LACM), Santa Catalina (1 SBMNH), Santa Cruz (1 SBMNH), Santa Rosa (26 LACM; 1 SBMNH)
Range: Also known from mainland (Smetana, 1995).
Notes. Species introduced from the Palearctic (Smetana, 1995). Cockerell (1940) recorded it as “Philonthus cruentus”.
Philonthus davus Smetana, 1995
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Cruz (3 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Smetana, 1995).
Philonthus flavolimbatus Erichson, 1840
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (Smetana, 1995).
Philonthus hepaticus Erichson, 1840
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Catalina (2 SBMNH)
Range: Also known from mainland (Smetana, 1995).
Philonthus lecontei Horn, 1884
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Rosa (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Smetana, 1995).
Philonthus longicornis Stephens, 1832
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Fall, 1897: 236)
Digitized Records: none
Range: Also known from mainland (Smetana, 1995).
Notes. This species was introduced from the Palearctic region (Smetana, 1995).
Philonthus quadrulus Horn, 1884
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Cruz (10 SBMNH)
Range: Also known from mainland (Smetana, 1995).
Philonthus triangulum Horn, 1884
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Smetana, 1995: 340)
Digitized Records: none
Range: Also known from mainland (Smetana, 1995).
Staphylinini: Quediina
Notes. Two genera and 40 species of Quediina have been recorded from California (Herman, 2001; M. L. Gimmel, 2022, unpublished data).
Quedius Stephens, 1829
Nomenclatural Authority: Herman (2001)
Literature Records (genus-only): Santa Cruz (Naughton et al., 2014: 304)
Digitized Records (genus-only): Santa Cruz (1 SBMNH)
Notes. Thirty-nine species of Quedius have been recorded from California, belonging to five subgenera, Distichalius Casey, 1915, Microsaurus Dejean, 1833, Paraquedius Casey, 1915, Quedius (s.str.), and Raphirus Stephens, 1829 (M. L. Gimmel, 2022, unpublished data). The species were revised for North America by Smetana (1971). One of the two voucher specimens (SBMNH, June specimen) of the “Quedius sp.” from the Naughton et al. (2014) study was examined by MLG; this specimen represents the digitized genus-only record above. It is a female of either Q. limbifer or Q. (Microsaurus) pellax Smetana, 1971.
Quedius (Microsaurus) limbifer Horn, 1878
Nomenclatural Authority: Smetana (1971), Herman (2001)
Literature Records: Santa Cruz (Smetana, 1971: 113)
Digitized Records: Santa Cruz (10 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Smetana, 1971).
Staphylinini: Staphylinina
Notes. Eight genera and 14 species of Staphylinina are known to occur in California (Herman, 2001; M. L. Gimmel, 2022, unpublished data).
Creophilus Leach, 1819
Nomenclatural Authority: Herman (2001)
Notes. One species of Creophilus has been recorded from California (Herman, 2001). The species were revised for the world by Clarke (2011).
Creophilus maxillosus (Linnaeus, 1758)
Nomenclatural Authority: Clarke (2011)
Literature Records: San Clemente (Fall, 1897: 236; Fall, 1901: 68), Santa Catalina (Fall, 1897: 236)
Digitized Records: San Clemente (1 LACM; 1 SBMNH), Santa Cruz (8 SBMNH), Santa Rosa (1 LACM; 6 SBMNH)
Range: Also known from mainland (Fall, 1901; Clarke, 2011).
Notes. This species was recorded as C. villosus by Fall (1897, 1901). Creophilus m. villosus (Gravenhorst, 1802) is the only subspecies of C. maxillosus (Linnaeus, 1758) over most of North America (Clarke, 2011); this is the subspecies occurring in the islands. The distribution map in Clarke (2011: 765) shows records in the Channel Islands, but these are not listed.
Hadrotes Mäklin, 1852
Nomenclatural Authority: Herman (2001)
Notes. One species of Hadrotes has been recorded from California (Herman, 2001).
Hadrotes crassus (Mannerheim, 1846)
Nomenclatural Authority: Herman (2001)
Literature Records: San Clemente (Caterino, Chatzimanolis & Richmond, 2015: 278), San Nicolas (Cockerell, 1940: 285; Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Catalina (Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Cruz (Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Rosa (Fall, 1897: 236; Caterino, Chatzimanolis & Richmond, 2015: 278)
Digitized Records: Anacapa (1 SBMNH), San Clemente (11 SBMNH), San Miguel (1 LACM; 8 SBMNH), San Nicolas (13 SBMNH), Santa Catalina (12 SBMNH), Santa Cruz (13 SBMNH), Santa Rosa (11 SBMNH; 2 UTCI)
Range: Also known from mainland (Caterino, Chatzimanolis & Richmond, 2015).
Tasgius Stephens, 1829
Nomenclatural Authority: Herman (2001)
Notes. Two adventive species of Tasgius have been recorded from California (Miller & Miller, 1985; Herman, 2001).
Tasgius ater (Gravenhorst, 1802)
Nomenclatural Authority: Herman (2001)
Literature Records: San Miguel (Miller & Miller, 1985: 124), Santa Barbara (Miller & Miller, 1985: 124)
Digitized Records: San Miguel (2 SBMNH), San Nicolas (1 SBMNH), Santa Rosa (6 SBMNH)
Range: Also known from mainland (Miller & Miller, 1985).
Notes. Introduced to North America from Europe (Miller & Miller, 1985). Recorded as Staphylinus ater (Gravenhorst) from Suaeda (Amaranthaceae) by Miller & Miller (1985).
Thinopinus LeConte, 1852
Nomenclatural Authority: Herman (2001)
Notes. Only one species of Thinopinus is known (Herman, 2001).
Thinopinus pictus LeConte, 1852
Nomenclatural Authority: Herman (2001)
Literature Records: San Miguel (Caterino, Chatzimanolis & Richmond, 2015: 278), San Nicolas (Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Catalina (Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Cruz (Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Rosa (Caterino, Chatzimanolis & Richmond, 2015: 278)
Digitized Records: San Miguel (12 SBMNH), San Nicolas (1 LACM; 13 SBMNH), Santa Catalina (1 SEMC; 11 SBMNH; 1 iNat), Santa Cruz (10 SBMNH), Santa Rosa (10 SBMNH; 2 UTCI; 1 iNat)
Range: Also known from mainland (Caterino, Chatzimanolis & Richmond, 2015).
Xantholinini
Notes. Fourteen genera and 55 species of Xantholinini are known to occur in California (Herman, 2001). The species of the tribe were revised for North America by Smetana (1982). A specimen deposited in LACM of an additional genus and species occurring on Santa Rosa Island, Neohypnus picipennis (LeConte, 1880), was captured by Scott Miller during the 1980s, but no specimen was located to substantiate this record (G-A. Kung, 2022, personal communication).
Linohesperus Smetana, 1982
Nomenclatural Authority: Herman (2001)
Digitized Records (genus-only): San Clemente (2 SBMNH), Santa Catalina (16 SBMNH), Santa Cruz (3 SBMNH)
Notes. Twenty-two species of Linohesperus have been recorded from California (Herman, 2001).
Linohesperus borealis (Casey, 1906)
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Smetana, 1982).
Linohesperus cuspifer Smetana, 1982
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Cruz (Smetana, 1988: 545)
Digitized Records: none
Range: Also known from mainland (Smetana, 1982, 1988).
Nudobius Thomson, 1860
Nomenclatural Authority: Herman (2001)
Notes. Only one species of Nudobius has been recorded from California (Herman, 2001).
Nudobius pugetanus Casey, 1906
Nomenclatural Authority: Smetana (1982), Herman (2001)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Smetana, 1982).
Tachyporinae
Notes. Three tribes, eight genera, and 32 species of Tachyporinae are known to occur in California (M. L. Gimmel, 2022, unpublished data). This subfamily was recently restricted (i.e., the previous tribe “Mycetoporini” excluded as a separate subfamily) by Yamamoto (2021).
Tachinusini
Notes. Three genera and 15 species of Tachinusini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Nitidotachinus Campbell, 1993
Nomenclatural Authority: Herman (2001)
Notes. Three species of Nitidotachinus have been recorded from California (Campbell, 1993a). This genus was reviewed by Campbell (1993a), who provided a key to species.
Nitidotachinus agilis (Horn, 1877)
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Campbell, 1993a).
Tachinus Gravenhorst, 1802
Nomenclatural Authority: Herman (2001)
Notes. Ten species of Tachinus have been recorded from California (M. L. Gimmel, 2022, unpublished data). Additional species of Tachinus are likely to occur on the Channel Islands. The genus was revised for North America by Campbell (1973) and updated by Campbell (1988).
Tachinus debilis Horn, 1877
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Cruz (9 SBMNH)
Range: Also known from mainland (Campbell, 1973).
Tachyporini
Notes. Three genera and 11 species of Tachyporini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Palporus Campbell, 1979
Nomenclatural Authority: Yamamoto (2021)
Notes. Only one species of Palporus occurs in California (Campbell, 1979). This genus was revised for North America by Campbell (1979), as Tachyporus (Palporus); the subgenus was subsequently elevated to genus by Yamamoto (2021).
Palporus nitidulus (Fabricius, 1781)
Nomenclatural Authority: Yamamoto (2021)
Literature Records: none
Digitized Records: San Nicolas (1 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Campbell, 1979).
Sepedophilus Gistel, 1856
Nomenclatural Authority: Herman (2001)
Notes. Three species of Sepedophilus have been recorded from California (Campbell, 1976). The genus was revised for North America by Campbell (1976).
Sepedophilus castaneus (Horn, 1877)
Nomenclatural Authority: Herman (2001)
Literature Records: none
Digitized Records: Santa Cruz (25 SBMNH), Santa Rosa (14 SBMNH)
Range: Also known from mainland (Campbell, 1976).
Tachyporus Gravenhorst, 1802
Nomenclatural Authority: Herman (2001)
Literature Records (genus-only): San Miguel (Miller & Davis, 1986: 550)
Notes. Seven species of Tachyporus have been recorded from California (Campbell, 1979). Additional species of Tachyporus are likely to occur on the Channel Islands. This genus was revised for North America by Campbell (1979).
Tachyporus californicus Horn, 1877
Nomenclatural Authority: Herman (2001)
Literature Records: Santa Catalina (Fall, 1897: 237), Santa Rosa (Fall, 1897: 237)
Digitized Records: San Clemente (1 SBMNH), San Nicolas (4 SBMNH), Santa Cruz (17 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Campbell, 1979).
BOSTRICHOIDEA
Bostrichidae
Notes. Five subfamilies, 19 genera, and 35 species of Bostrichidae are known to occur in California (M. L. Gimmel, 2022, unpublished data). The works of Fisher (1950; all subfamilies except Lyctinae) and Gerberg (1957; Lyctinae) adequately cover the North American fauna known at the time. Borowski & Węgrzynowicz (2007) provided a world catalog of this group; we follow their classification below.
Bostrichinae
Notes. Four tribes, 10 genera, and 15 species of Bostrichinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Bostrichini
Notes. Four genera and seven species of Bostrichini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Amphicerus LeConte, 1861
Nomenclatural Authority: Borowski & Węgrzynowicz (2007)
Notes. Three species of Amphicerus are known to occur in California (M. L. Gimmel, 2022, unpublished data). The species were keyed by Fisher (1950).
Amphicerus cornutus (Pallas, 1772)
Nomenclatural Authority: Fisher (1950), Borowski & Węgrzynowicz (2007)
Literature Records: none
Digitized Records: Santa Catalina (4 LACM), Santa Cruz (9 SBMNH; 1 UCSB)
Range: Also known from mainland (Fisher, 1950; Borowski & Węgrzynowicz, 2007).
Xyloperthini
Notes. Four genera and six species of Xyloperthini have been recorded from California (Fisher, 1950).
Scobicia Lesne, 1901
Nomenclatural Authority: Borowski & Węgrzynowicz (2007)
Notes. Two species of Scobicia are known to occur in California (Fisher, 1950). These species were keyed by Fisher (1950).
Scobicia declivis (LeConte, 1857)
Nomenclatural Authority: Fisher (1950), Borowski & Węgrzynowicz (2007)
Literature Records: San Nicolas (Fall, 1897: 238; Fall, 1901: 133; Burke, Hartman & Snyder, 1922: 12; Fisher, 1950: 111)
Digitized Records: Santa Cruz (4 SBMNH)
Range: Also known from mainland (Fall, 1901; Fisher, 1950; Borowski & Węgrzynowicz, 2007).
Notes. Fall (1897, 1901) recorded this species as Sinoxylon declive. Borowski & Węgrzynowicz (2007) falsely stated the year of publication of the species as 1859.
Scobicia suturalis (Horn, 1878)
Nomenclatural Authority: Fisher (1950), Borowski & Węgrzynowicz (2007)
Literature Records: none
Digitized Records: Anacapa (1 SBMNH), Santa Catalina (1 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Fisher, 1950; Borowski & Węgrzynowicz, 2007).
Dinoderinae
Notes. Four genera and eight species of Dinoderinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Stephanopachys Waterhouse, 1888
Nomenclatural Authority: Borowski & Węgrzynowicz (2007)
Notes. Four species of Stephanopachys have been reported from California (M. L. Gimmel, 2022, unpublished data). The species were keyed out by Fisher (1950).
Stephanopachys substriatus (Paykull, 1800)
Nomenclatural Authority: Fisher (1950), Borowski & Węgrzynowicz (2007)
Literature Records: none
Digitized Records: Santa Catalina (11 LACM)
Range: Also known from mainland (Fisher, 1950; Borowski & Węgrzynowicz, 2007).
Lyctinae
Notes. Two genera and six species of Lyctinae have been recorded from California (Gerberg, 1957).
Lyctus Fabricius, 1792
Nomenclatural Authority: Borowski & Węgrzynowicz (2007)
Notes. Five species have been reported from California (Gerberg, 1957). These were keyed out by Gerberg (1957).
Lyctus cavicollis LeConte, 1866
Nomenclatural Authority: Gerberg (1957), Borowski & Węgrzynowicz (2007)
Literature Records: none
Digitized Records: Santa Cruz (7 SBMNH)
Range: Also known from mainland (Gerberg, 1957; Borowski & Węgrzynowicz, 2007).
Lyctus linearis (Goeze, 1777)
Nomenclatural Authority: Gerberg (1957), Borowski & Węgrzynowicz (2007)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Gerberg, 1957; Borowski & Węgrzynowicz, 2007).
Notes. This species is cosmopolitan (Gerberg, 1957).
Lyctus planicollis LeConte, 1858
Nomenclatural Authority: Gerberg (1957), Borowski & Węgrzynowicz (2007)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Gerberg, 1957; Borowski & Węgrzynowicz, 2007).
Notes. This species was indicated as a synonym of the older Lyctus carbonarius Waltl, 1832 by Lesne (1916); however, the synonymy was seen as tentative by Gerberg (1957: 26). Borowski & Węgrzynowicz (2007) treated L. carbonarius as valid with priority over L. planicollis.
Polycaoninae
Notes. Two genera and four species of Polycaoninae have been recorded from California (Fisher, 1950).
Melalgus Dejean, 1833
Nomenclatural Authority: Borowski & Węgrzynowicz (2007)
Notes. Two species of Melalgus are known from California (Fisher, 1950). The species were keyed out by Fisher (1950).
Melalgus confertus (LeConte, 1866)
Nomenclatural Authority: Fisher (1950), Borowski & Węgrzynowicz (2007)
Literature Records: none
Digitized Records: Santa Catalina (22 LACM)
Range: Also known from mainland (Fisher, 1950; Borowski & Węgrzynowicz, 2007).
Polycaon Castelnau, 1836
Nomenclatural Authority: Borowski & Węgrzynowicz (2007)
Notes. Two species of Polycaon are known from California (Fisher, 1950). The species were keyed out by Fisher (1950).
Polycaon stoutii (LeConte, 1853)
Nomenclatural Authority: Fisher (1950), Borowski & Węgrzynowicz (2007)
Literature Records: Santa Cruz (LeConte, 1876: 299; Fall, 1897: 238; Fall & Davis, 1934: 144)
Digitized Records: Santa Catalina (6 LACM; 2 SBMNH; 1 iNat)
Range: Also known from mainland (Fisher, 1950; Borowski & Węgrzynowicz, 2007).
Notes. In addition to P. stoutii, LeConte (1876) also reported Polycaon ovicollis (LeConte, 1857) from Santa Cruz Island. The latter is now considered a junior synonym of the former (see Fisher, 1950).
Psoinae
Notes. One genus and two species of Psoinae have been recorded from California (Fisher, 1950), belonging to the tribe Psoini.
Psoa Herbst, 1797
Nomenclatural Authority: Borowski & Węgrzynowicz (2007)
Notes. Two species of Psoa are known from California (Fisher, 1950). The species were keyed out by Fisher (1950).
Psoa maculata (LeConte, 1852)
Nomenclatural Authority: Fisher (1950), Borowski & Węgrzynowicz (2007)
Literature Records: none
Digitized Records: Santa Catalina (4 LACM)
Range: Also known from mainland (Fisher, 1950; Borowski & Węgrzynowicz, 2007).
Psoa quadrisignata (Horn, 1868)
Nomenclatural Authority: Fisher (1950), Borowski & Węgrzynowicz (2007)
Literature Records: Santa Catalina (Cockerell, 1940: 286)
Digitized Records: none
Range: Also known from mainland (Fisher, 1950; Borowski & Węgrzynowicz, 2007).
Dermestidae
Notes. Six subfamilies, 18 genera, and 77 species are known to occur in California (Háva & Herrmann, 2021; M. L. Gimmel, 2022, unpublished data). Beal (2003) provided a distributional checklist of Dermestidae from North America, which was updated by Háva & Herrmann (2021). We use the classification of the latter publication below.
Dermestinae
Notes. One genus and 14 species of Dermestinae are known to occur in California, belonging to the tribe Dermestini (Háva & Herrmann, 2021).
Dermestes Linnaeus, 1758
Nomenclatural Authority: Beal (2003)
Digitized Records (genus-only): Anacapa (9 LACM), San Clemente (1 LACM; 22 YPMC), Santa Barbara (3 LACM), Santa Catalina (21 LACM; 2 YPMC), Santa Cruz (5 YPMC)
Notes. Fourteen species of Dermestes are known to occur in California, belonging to two subgenera, Dermestes (s.str.) and Dermestinus Zantiev, 1967 (Beal, 2003; Háva & Herrmann, 2021). These were all keyed out in the work of Lepesme (1949).
Dermestes (Dermestinus) caninus Germar, 1824
Nomenclatural Authority: Beal (2003), Háva & Herrmann (2021)
Literature Records: San Clemente (Fall, 1897: 237), San Nicolas (Fall, 1897: 237), Santa Barbara (Fall, 1897: 237; Miller & Miller, 1985: 125), Santa Rosa (Fall, 1897: 237)
Digitized Records: Santa Barbara (2 SBMNH)
Range: Also known from mainland (Beal, 2003; Háva & Herrmann, 2021).
Notes. Fall (1897) recorded this species as Dermestes mannerheimii LeConte, which is a junior synonym of D. caninus (see Miller & Miller, 1985). These taxa represent two valid subspecies according to Háva & Herrmann (2021).
Dermestes (Dermestinus) frischi Kugelann, 1792
Nomenclatural Authority: Beal (2003)
Literature Records: San Clemente (Miller & Miller, 1985: 125), San Miguel (Miller & Miller, 1985: 125), Santa Barbara (Miller & Miller, 1985: 125), Santa Cruz (Fall & Davis, 1934: 143; Miller & Miller, 1985: 125)
Digitized Records: San Clemente (5 LACM; 3 SBMNH), San Miguel (1 SBMNH), San Nicolas (8 SBMNH), Santa Barbara (3 SBMNH), Santa Cruz (5 SBMNH)
Range: Also known from mainland (Beal, 2003; Háva & Herrmann, 2021).
Notes. The species epithet was spelled frischii by Háva & Herrmann (2021), who did not include California in the list of states in which the species was known to occur. This species is cosmopolitan (Háva & Herrmann, 2021).
Dermestes (Dermestinus) marmoratus Say, 1823
Nomenclatural Authority: Beal (2003), Háva & Herrmann (2021)
Literature Records: San Clemente (Fall, 1897: 237), San Nicolas (Fall, 1897: 237), Santa Catalina (Fall, 1897: 237), Santa Rosa (Fall, 1897: 237)
Digitized Records: San Clemente (1 YPMC)
Range: Also known from mainland (Beal, 2003; Háva & Herrmann, 2021).
Dermestes (Dermestinus) rattus LeConte, 1854
Nomenclatural Authority: Beal (2003), Háva & Herrmann (2021)
Literature Records: Santa Rosa (Fall, 1897: 237)
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Beal, 2003; Beal & Seeno, 1977; Háva & Herrmann, 2021).
Notes. This species was recorded as Dermestes tristis by Fall (1897); the subspecies of D. rattus occurring on the islands is D. r. tristis Fall, 1897.
Dermestes (Dermestinus) talpinus Mannerheim, 1843
Nomenclatural Authority: Beal (2003), Háva & Herrmann (2021)
Literature Records: Santa Cruz (LeConte, 1876: 298; Fall, 1897: 237; Fall & Davis, 1934: 143)
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Beal, 2003; Háva & Herrmann, 2021).
Megatominae
Notes. Two tribes, seven genera, and 42 species of Dermestidae are known to occur in California (Háva & Herrmann, 2021; M. L. Gimmel, 2022, unpublished data).
Anthrenini
Notes. One genus and 10 species of Anthrenini have been recorded from California (Háva & Herrmann, 2021).
Anthrenus Geoffroy, 1762
Nomenclatural Authority: Kadej (2011), Háva & Herrmann (2021)
Notes. Ten species of Anthrenus have been reported from California, belonging to four subgenera, Anthrenops Reitter, 1881, Anthrenus (s.str.), Florilinus Mulsant & Rey, 1868, and Nathrenus Casey, 1900 (Beal, 2003; Háva & Herrmann, 2021). The North American species were revised by Beal (1998); Kadej (2011) described an additional species from California and provided an updated key to North American species.
Anthrenus (Anthrenus) lepidus LeConte, 1854
Nomenclatural Authority: Beal (1998), Beal (2003), Háva & Herrmann (2021)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Beal, 1998, 2003; Háva & Herrmann, 2021).
Anthrenus (Nathrenus) verbasci (Linnaeus, 1767)
Nomenclatural Authority: Beal (1998), Beal (2003), Háva & Herrmann (2021)
Literature Records: Santa Catalina (Cockerell, 1940: 286)
Digitized Records: Santa Catalina (1 iNat)
Range: Also known from mainland (Beal, 1998, 2003; Háva & Herrmann, 2021).
Notes. This is a cosmopolitan species (Beal, 2003).
Megatomini
Notes. Six genera and 32 species of Megatomini are known to occur in California (Háva & Herrmann, 2021; M. L. Gimmel, 2022, unpublished data).
Cryptorhopalum Guérin-Méneville, 1838
Nomenclatural Authority: Háva & Herrmann (2021)
Notes. Nine species of Cryptorhopalum have been recorded from California (Beal, 2003; Háva & Herrmann, 2021). These were mostly revised and keyed by Beal (1979), who later published a revision of the entire Nearctic fauna (Beal, 1985).
Cryptorhopalum apicale (Mannerheim, 1843)
Nomenclatural Authority: Beal (1985), Beal (2003), Háva & Herrmann (2021)
Literature Records: Santa Cruz (Beal, 1979: 16 [map]; Beal, 1985: 191 [map])
Digitized Records: Santa Cruz (18 SBMNH)
Range: Also known from mainland (Beal, 1979, 1985, 2003; Háva & Herrmann, 2021).
Cryptorhopalum triste LeConte, 1854
Nomenclatural Authority: Beal (1985), Beal (2003), Háva & Herrmann (2021)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Beal, 1979, 1985, 2003; Háva & Herrmann, 2021).
Megatoma Herbst, 1791
Nomenclatural Authority: Háva & Herrmann (2021)
Digitized Records (genus-only): Santa Rosa (6 SBMNH)
Notes. Eleven species of Megatoma have been reported from California, distributed among two subgenera, Megatoma (s.str.) and Pseudohadrotoma Kalík, 1957 (Háva & Herrmann, 2021). The species in North America were revised by Beal (1967).
Megatoma (Megatoma) variegata (Horn, 1875)
Nomenclatural Authority: Beal (2003), Háva & Herrmann (2021)
Literature Records: none
Digitized Records: Santa Cruz (10 SBMNH), Santa Rosa (11 SBMNH)
Range: Also known from mainland (Beal, 2003; Háva & Herrmann, 2021).
Trogoderma Dejean, 1821
Nomenclatural Authority: Beal (2003), Háva & Herrmann (2021)
Notes. Nine species of Trogoderma have been reported from California (Beal, 2003; Háva & Herrmann, 2021). The North American species were revised by Beal (1954).
Trogoderma sternale Jayne, 1882
Nomenclatural Authority: Beal (1954), Beal (2003), Háva & Herrmann (2021)
Literature Records: Santa Barbara (Miller & Miller, 1985: 125), Santa Catalina (Fall, 1897: 237; Fall, 1901: 93; Beal, 1954: 72 [map]; Miller & Miller, 1985: 125)
Digitized Records: Santa Barbara (1 SBMNH), Santa Catalina (4 SBMNH), Santa Cruz (75 SBMNH; 1 UCSB), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Beal, 1954, 2003; Háva & Herrmann, 2021).
Notes. The subspecies of T. sternale present on the islands is the nominate subspecies, T. s. sternale Jayne, 1882 (Beal, 1954).
Ptinidae
Notes. Nine subfamilies, 41 genera, and 145 species of Ptinidae are known to occur in California (M. L. Gimmel, 2022, unpublished data). The family Ptinidae in the modern sense corresponds with the combined former concepts of Anobiidae and Ptinidae. The species (excluding Ptininae) were cataloged for North America by White (1982). The classification we employ below follows Philips (2002), with the exception of the use of Ptinidae over Anobiidae.
Anobiinae
Notes. Seven tribes, 13 genera, and 31 species of Anobiinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Anobiini
Notes. Three genera and seven species of Anobiini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Hemicoelus LeConte, 1861
Nomenclatural Authority: White (1976a), White (1982)
Notes. Four species of Hemicoelus are known to occur in California, including three previously reported (White, 1982) and the new state record below. White (1976a) provided a key to North American species.
Hemicoelus nelsoni (Hatch, 1961)
Nomenclatural Authority: White (1976a), White (1982)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (White, 1982).
Notes. This species represents a new state record for California.
Colposternini
Notes. One species of Colposternini has been recorded from California (White, 1982).
Colposternus Fall, 1905
Nomenclatural Authority: White (1982)
Notes. One species of Colposternus has been recorded from California (White, 1982).
Colposternus tenuilineatus (Horn, 1894)
Nomenclatural Authority: White (1982)
Literature Records: Santa Catalina (Fall, 1897: 238; Fall, 1901: 132; Fall, 1905: 191)
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (White, 1982).
Notes. Fall (1897, 1901) recorded this species as Trypopitys tenuilineata; Fall (1905) subsequently transferred it to Colposternus.
Euceratocerini
Notes. Two or three genera and 10 or 11 species of Euceratocerini have been recorded from California (White, 1982; M. L. Gimmel, 2022, unpublished data).
Actenobius Fall, 1905
Nomenclatural Authority: White (1982)
Notes. One species of Actenobius has been recorded from California (White, 1982).
Actenobius pleuralis (Casey, 1898)
Nomenclatural Authority: White (1982)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (White, 1982).
Euceratocerus LeConte, 1874
Nomenclatural Authority: White (1982)
Notes. No species of Euceratocerus have been reported from California (White, 1982). The species were treated for North America by Hinson (2021).
Euceratocerus hornii LeConte, 1874
Nomenclatural Authority: White (1982)
Literature Records: Santa Catalina (Fall, 1897: 238)
Digitized Records: none
Range: Also known from mainland (White, 1982; Hinson, 2021).
Notes. This species is otherwise reported only from Texas, with all other species of the genus being from points farther east (White, 1982), casting extreme doubt on Fall’s (1897) record from Santa Catalina. Fall’s record may have referred to Santa Catalina Island specimens of another ptinid species with pectinate antennae, Ptilinus basalis LeConte, 1858, in the H.C. Fall collection at the Museum of Comparative Zoology, Harvard University (S. Miller, 2022, personal communication). This species was not mentioned in Fall (1897), so these specimens were probably (re)determined by H.C. Fall after that publication.
Hadrobregmini
Notes. Two genera and three species of Hadrobregmini have been recorded from California (White, 1982).
Priobium Motschulsky, 1845
Nomenclatural Authority: White (1982)
Notes. One species of Priobium has been recorded from California (White, 1982).
Priobium punctatum (LeConte, 1859)
Nomenclatural Authority: White (1982)
Literature Records: Santa Cruz (Cockerell, 1940: 286)
Digitized Records: Santa Catalina (1 iNat), Santa Cruz (7 SBMNH)
Range: Also known from mainland (White, 1982).
Notes. Cockerell (1940) reported this species as Trypopitys punctatus.
Stegobiini
Notes. Two genera and five species of Stegobiini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Oligomerus Redtenbacher, 1849
Nomenclatural Authority: White (1982)
Notes. Four species of Oligomerus are known to occur in California, including one putatively undescribed species (M. L. Gimmel, 2022, unpublished data). White (1976a) provided a key to species of Oligomerus.
Oligomerus delicatulus (Fall, 1920)
Nomenclatural Authority: White (1976a), White (1982)
Literature Records: none
Digitized Records: Anacapa (1 SBMNH)
Range: Also known from mainland (White, 1982).
Stegobium Motschulsky, 1860
Nomenclatural Authority: White (1982)
Notes. One species of Stegobium is known to occur in California (M. L. Gimmel, 2022, unpublished data).
Stegobium paniceum (Linnaeus, 1758)
Nomenclatural Authority: White (1982)
Literature Records: none
Digitized Records: Santa Rosa (4 SBMNH)
Range: Also known from mainland (White, 1982).
Notes. This pest species, known as the drugstore beetle, has been introduced around the world.
Dorcatominae
Notes. Three tribes, five genera, and seven species of Dorcatominae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Dorcatomini
Notes. Three genera and five species of Dorcatomini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Byrrhodes LeConte, 1878
Nomenclatural Authority: White (1982)
Notes. Two species of Byrrhodes have been recorded from California, including one putatively undescribed species (M. L. Gimmel, 2022, unpublished data). White (1973a) provided a key to North American species.
Byrrhodes ulkei (Fall, 1905)
Nomenclatural Authority: White (1973a), White (1982)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (White, 1982).
Notes. In the SBMNH collection, T.K. Philips identified a possible new species of Byrrhodes from specimens collected on the coastal side of the Santa Ynez Mountains in Santa Barbara County; the above record of B. ulkei possibly represents this species.
Petaliini
Notes. One species of Petaliini has been recorded from California (White, 1982).
Petalium LeConte, 1861
Nomenclatural Authority: Ford (1973), White (1982)
Notes. One species of Petalium has been recorded from California (White, 1982). Ford (1973) provided a key to North American species of this genus.
Petalium californicum Fall, 1905
Nomenclatural Authority: Ford (1973), White (1982)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Ford, 1973; White, 1982).
Ernobiinae
Notes. Three tribes, seven genera, and 27 species of Ernobiinae are known to occur in California (White, 1982; M. L. Gimmel, 2022, unpublished data).
Ernobiini
Notes. Three genera and 19 species of Ernobiini are known to occur in California (White, 1982; M. L. Gimmel, 2022, unpublished data).
Ernobius Thomson, 1859
Nomenclatural Authority: White (1982)
Notes. Seventeen species of Ernobius are known to occur in California, including one putatively undescribed species (M. L. Gimmel, 2022, unpublished data). The species of this genus were keyed for California by Ruckes (1957).
Ernobius debilis LeConte, 1865
Nomenclatural Authority: Ruckes (1957), White (1982)
Literature Records: Santa Cruz (LeConte, 1865: 225; Fall, 1897: 238; Fall, 1901: 131)
Digitized Records: Santa Cruz (10 SBMNH)
Range: Also known from mainland (Fall, 1901).
Notes. This species was considered endemic to Santa Cruz Island by LeConte (1865) and Fall (1897), but subsequently discovered on the mainland (Fall, 1901).
Ernobius punctulatus (LeConte, 1859)
Nomenclatural Authority: Ruckes (1957), White (1982)
Literature Records: Santa Cruz (Fall & Davis, 1934: 144)
Digitized Records: none
Range: Also known from mainland (White, 1982).
Ozognathini
Notes. Two genera and three species of Ozognathini have been recorded from California (White, 1982).
Ozognathus LeConte, 1861
Nomenclatural Authority: White (1982)
Notes. One species of Ozognathus has been reported from California (White, 1982).
Ozognathus cornutus (LeConte, 1859)
Nomenclatural Authority: White (1982)
Literature Records: none
Digitized Records: Santa Catalina (1 iNat), Santa Cruz (2 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (White, 1982).
Xarifa Fall, 1905
Nomenclatural Authority: White (1982)
Notes. Originally described as a genus endemic to the Channel Islands, one species, Xarifa lobata Fall, 1929, was subsequently described from mainland California (Carmel; Fall, 1929a). These remain the only two species known; they were keyed by White (1974).
Xarifa insularis Fall, 1905
Nomenclatural Authority: White (1982)
Literature Records: San Clemente (Fall, 1905: 138; Miller, 1985a: 20), Santa Catalina (Fall, 1905: 138; Miller, 1985a: 20; White, 1982: 3)
Digitized Records: Santa Catalina (3 SBMNH), Santa Cruz (1 LACM; 6 SBMNH), Santa Rosa (12 SBMNH)
Range: Endemic (Fall, 1905; Miller, 1985a).
Xestobiini
Notes. Two genera and five species of Xestobiini have been recorded from California (White, 1982).
Xestobium Motschulsky, 1845
Nomenclatural Authority: White (1982)
Notes. Three species of Xestobium have been reported from California (White, 1982). These species were partially keyed by White (1975), with another species added by White (1976b).
Xestobium marginicolle (LeConte, 1859)
Nomenclatural Authority: White (1982)
Literature Records: Santa Barbara (Miller & Miller, 1985: 123)
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (White, 1982).
Notes. Reported from Hemizonia (Asteraceae) on Santa Barbara Island by Miller & Miller (1985).
Eucradinae
Notes. Only one genus, belonging to the tribe Hedobiini, and three species of Eucradinae have been recorded from California (White, 1982).
Ptinomorphus Mulsant & Rey, 1868
Nomenclatural Authority: Zahradník & Háva (2014)
Notes. Three species of Ptinomorphus have been reported from California (White, 1982, as Hedobia Dejean, 1821).
Ptinomorphus granosus (LeConte, 1874)
Nomenclatural Authority: White (1982), Zahradník & Háva (2014)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (White, 1982).
Notes. This species was previously known as Hedobia granosa.
Mesocoelopodinae
Notes. Only one genus, belonging to the tribe Tricorynini, and 20 species of Mesocoelopodinae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Tricorynus Waterhouse, 1849
Nomenclatural Authority: White (1982)
Literature Records (genus-only): Santa Cruz (Naughton et al., 2014: 303)
Digitized Records (genus-only): Anacapa (2 SBMNH), San Clemente (5 SBMNH), San Nicolas (2 SBMNH), Santa Barbara (1 SBMNH), Santa Catalina (3 SBMNH), Santa Cruz (7 SBMNH)
Notes. Twenty species of Tricorynus have been recorded from California (M. L. Gimmel, 2022, unpublished data). This genus was revised for North America by White (1965).
Tricorynus nubilus (Fall, 1905)
Nomenclatural Authority: White (1965), White (1982)
Literature Records: Santa Catalina (White, 1965: 333)
Digitized Records: none
Range: Also known from mainland (White, 1965, 1982).
Tricorynus obsoletus (LeConte, 1865)
Nomenclatural Authority: White (1965), White (1982)
Literature Records: Santa Catalina (Fall, 1897: 238)
Digitized Records: none
Range: Also known from mainland (White, 1965).
Notes. Fall (1897) listed this species as Hemiptychus obsoletus with a question mark.
Ptilininae
Notes. Only one genus, belonging to the tribe Ptilinini, and four species of Ptilininae have been recorded from California (White, 1982).
Ptilinus Müller, 1764
Nomenclatural Authority: White (1982)
Notes. Four species of Ptilinum have been recorded from California (White, 1982). No modern key exists to separate these species.
Ptilinus basalis LeConte, 1858
Nomenclatural Authority: White (1982)
Literature Records: none
Digitized Records: Santa Cruz (3 SBMNH)
Range: Also known from mainland (White, 1982).
Notes. See Notes for Euceratocerus hornii above.
Ptininae: Ptinini
Notes. Four tribes, six genera, and 23 species of Ptininae, including three genera and 19 species of Ptinini, are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Ptinus Linnaeus, 1766
Nomenclatural Authority: Papp & Okumura (1959)
Notes. Fifteen species of Ptinus have been recorded from California, distributed among three subgenera, Gynopterus Mulsant & Rey, 1868, Ptinus (s.str.), and Tectoptinus Iablokoff-Khnzorian & Karapetyan, 1986 (Papp & Okumura, 1959; Zahradník & Háva, 2014). The key of Papp & Okumura (1959) can be used to separate these. Naughton et al. (2014: 303) recorded the genus only from Santa Cruz Island, but this has since been identified as P. agnatus (see below).
Ptinus (Gynopterus) fallax Fall, 1905
Nomenclatural Authority: Papp & Okumura (1959)
Literature Records: none
Digitized Records: Santa Catalina (6 SBMNH; 1 iNat)
Range: Also known from mainland (Papp & Okumura, 1959).
Ptinus (Ptinus) agnatus Fall, 1905
Nomenclatural Authority: Papp & Okumura (1959)
Literature Records: none
Digitized Records: Santa Cruz (3 SBMNH), Santa Rosa (13 SBMNH)
Range: Also known from mainland (Papp & Okumura, 1959).
Notes. The Santa Cruz Island specimen of “Ptinus” from Naughton et al. (2014) exists as a voucher (in SBMNH) and was identified to this species by MLG.
Xyletininae
Notes. Two tribes, six genera, and 29 species of Xyletininae have been recorded from California (White, 1982; M. L. Gimmel, 2022, unpublished data).
Lasiodermini
Notes. Two genera and five species of Lasiodermini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Lasioderma Stephens, 1835
Nomenclatural Authority: White (1982)
Notes. Two species of Lasioderma are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Lasioderma serricorne (Fabricius, 1792)
Nomenclatural Authority: White (1982)
Literature Records: none
Digitized Records: Santa Cruz (15 LACM)
Range: Also known from mainland (White, 1982).
Notes. This cosmopolitan pest is known as the cigarette beetle.
Xyletinini
Notes. Four genera and 24 species of Xyletinini have been recorded from California (White, 1982).
Euvrilletta Fall, 1905
Nomenclatural Authority: White (1982)
Notes. Seven species of Euvrilletta have been recorded from California (White, 1982). The key in White (1985) can be used to identify them.
Euvrilletta catalinae (Fall, 1905)
Nomenclatural Authority: White (1982)
Literature Records: Santa Catalina (Fall, 1897: 238; Fall, 1901: 131; Fall, 1905: 162; Miller, 1985a: 20; Caterino & Chandler, 2010: 187)
Digitized Records: none
Range: Endemic (Fall, 1905; Miller, 1985a; Caterino & Chandler, 2010).
Notes. Fall (1905) recorded this species as Oligomerodes catalinae Fall; the genus Oligomerodes Fall was subsequently synonymized with Euvrilletta Fall by White (1976a: 164), and the species was included in a key by White (1985: 191). Specimens referred to by Fall (1897: 238, 1901: 131) as “Oligomerus? new species” represent this species. Adults were collected from foliage of Heteromeles arbutifolia (Lindl.) M.Roem. (Rosaceae) (Fall, 1901; White, 1982).
Euvrilletta occidentalis (Fall, 1905)
Nomenclatural Authority: White (1982)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (White, 1982).
Vrilletta LeConte, 1874
Nomenclatural Authority: White (1982)
Notes. Ten species of Vrilletta have been recorded from California (White, 1982). White (1980) reviewed the genus and provided a key to species.
Vrilletta blaisdelli Fall, 1905
Nomenclatural Authority: White (1982)
Literature Records: Santa Catalina (Cockerell, 1940: 286)
Digitized Records: Santa Cruz (1 SBMNH), Santa Rosa (11 SBMNH)
Range: Also known from mainland (White, 1982).
Xyletinus Latreille, 1809
Nomenclatural Authority: White (1977), White (1982)
Notes. Six species of Xyletinus have been recorded from California (White, 1982). White (1977) provided a partial, updated key to these species, supplementing that of White (1973b).
Xyletinus undetermined species
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Notes. The single SBMNH specimen roughly keys to Xyletinus rotundicollis White, 1977 in White (1977). However, without adequate comparative material we hesitate to make a species-level identification.
CLEROIDEA
Byturidae
Notes. Two genera and two species are known from California (Springer & Goodrich, 1983). Springer & Goodrich (1983) provided a revision of the family for North America.
Xerasia Lewis, 1895
Nomenclatural Authority: Springer & Goodrich (1991)
Notes. Only one species of Xerasia is known from North America (Springer & Goodrich, 1991).
Xerasia grisescens (Jayne, 1882)
Nomenclatural Authority: Springer & Goodrich (1991)
Literature Records: Santa Catalina (Cockerell, 1940: 286; Springer & Goodrich, 1983: 190 [map only]), Santa Cruz (Springer & Goodrich, 1983: 190 [map only]; Naughton et al., 2014: 303)
Digitized Records: San Miguel (1 SBMNH), Santa Catalina (2 LACM; 11 SBMNH), Santa Cruz (2 SBMNH), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Springer & Goodrich, 1983).
Notes. This species was recorded by Cockerell (1940) as Byturus grisescens, and by Springer & Goodrich (1983) as Byturellus grisescens.
Cleridae
Notes. Four subfamilies, 18 genera, and about 88 species of Cleridae are known from California (M. L. Gimmel, 2022, unpublished data).
Clerinae
Notes. Five tribes, six genera and 43 species of Clerinae are known to occur in California (M. L. Gimmel, 2022, unpublished data). This subfamily was recently redefined and reclassified by Bartlett (2021).
Dieropsini
Notes. One genus and four species of Dieropsini are known to occur in California (Foster, 1976).
Trichodes Herbst, 1792
Nomenclatural Authority: Wolcott (1947)
Notes. Four species of Trichodes are known from California (Foster, 1976). These were keyed by Foster (1976).
Trichodes ornatus Say, 1823
Nomenclatural Authority: Foster (1976)
Literature Records: none
Digitized Records: Santa Cruz (7 SBMNH; 6 UCSB)
Range: Also known from mainland (Foster, 1976).
Notes. The subspecies of T. ornatus occurring in coastal California is T. o. douglasianus White, 1849 (Foster, 1976).
Hydnocerini
Notes. One genus and 15 species of Hydnocerini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Phyllobaenus Dejean, 1837
Nomenclatural Authority: Wolcott (1947)
Literature Records (genus-only): Santa Cruz (Naughton et al., 2014: 303)
Digitized Records (genus-only): Santa Catalina (1 SBMNH), Santa Cruz (2 SBMNH; 2 UCSB)
Notes. Fifteen species of Phyllobaenus are known to occur in California (M. L. Gimmel, 2022, unpublished data). The two SBMNH specimens from Santa Cruz are vouchers for the Naughton et al. (2014) study.
Phyllobaenus funebris (Chevrolat, 1874)
Nomenclatural Authority: Wolcott (1947)
Literature Records: none
Digitized Records: San Miguel (2 SBMNH), Santa Cruz (14 SBMNH)
Range: Also known from mainland (Wolcott, 1947).
Phyllobaenus scaber (LeConte, 1852)
Nomenclatural Authority: Wolcott (1947)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Wolcott, 1947).
Korynetinae
Notes. Six genera and 13 species of Korynetinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Loedelia Lucas, 1920
Nomenclatural Authority: Wolcott (1947)
Notes. One species of Loedelia is known from California (Wolcott, 1947).
Loedelia maculicollis (LeConte, 1874)
Nomenclatural Authority: Wolcott (1947)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH; 1 UCSB)
Range: Also known from mainland (Wolcott, 1947).
Necrobia Olivier, 1795
Nomenclatural Authority: Wolcott (1947)
Notes. Three species of Necrobia are known from California (M. L. Gimmel, 2022, unpublished data). Although these have long been thought to be introductions to North America from other parts of the world, evidence presented from tar pit material by Holden, Barclay & Angus (2018) casts doubt on this for the third known Californian species, Necrobia violacea (Linnaeus, 1758).
Necrobia ruficollis (Fabricius, 1775)
Nomenclatural Authority: Papp (1959)
Literature Records: San Clemente (Fall, 1897: 238; Fall, 1901: 130)
Digitized Records: Anacapa (1 SBMNH), Santa Cruz (5 LACM; 2 SBMNH)
Range: Also known from mainland (Wolcott, 1947).
Notes. Fall (1901) recorded this species as Corynetes ruficollis. This is a cosmopolitan species (Papp, 1959).
Necrobia rufipes (DeGeer, 1775)
Nomenclatural Authority: Papp (1959)
Literature Records: San Clemente (Fall, 1897: 238), Santa Catalina (Fall, 1897: 238), Santa Rosa (Fall, 1897: 238)
Digitized Records: Anacapa (9 LACM), San Clemente (22 LACM; 1 SBMNH), San Miguel (4 LACM; 3 SBMNH), San Nicolas (28 LACM; 7 SBMNH), Santa Catalina (1 LACM; 1 SBMNH), Santa Cruz (98 LACM; 8 SBMNH; 2 UCSB), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Fall, 1901; Wolcott, 1947). This is a cosmopolitan species (Papp, 1959).
Tillinae
Notes. Six genera and 32 species of Tillinae are known to occur in California (Burke, Leavengood & Zolnerowich, 2015; M. L. Gimmel, 2022, unpublished data).
Cymatodera Gray, 1832
Nomenclatural Authority: Burke, Leavengood & Zolnerowich (2015)
Digitized Records (genus-only): Santa Cruz (1 UCRC)
Notes. Twenty-five species of the genus Cymatodera have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Cymatodera angustata Spinola, 1844
Nomenclatural Authority: Burke, Leavengood & Zolnerowich (2015)
Literature Records: Santa Rosa (Fall, 1897: 238)
Digitized Records: none
Range: Also known from mainland (Rifkind, 2019).
Notes. This flightless species was not reported from the islands by Rifkind (2019), and Fall’s (1897) record may actually refer to C. caterinoi.
Cymatodera caterinoi Rifkind & Burke, 2019
Nomenclatural Authority: Rifkind (2019)
Literature Records: Anacapa (Rifkind, 2019: 556), Santa Cruz (Rifkind, 2019: 556), Santa Rosa (Rifkind, 2019: 556)
Digitized Records: Santa Cruz (5 SBMNH; 1 UCSB), Santa Rosa (1 CASC; 1 CSCA; 4 JNRC; 3 SBMNH)
Range: Endemic (Rifkind, 2019).
Notes. Flightless (Rifkind, 2019).
Cymatodera insularis Rifkind, 2019
Nomenclatural Authority: Rifkind (2019)
Literature Records: San Clemente (Rifkind, 2019: 553), Santa Catalina (Rifkind, 2019: 553)
Digitized Records: San Nicolas (1 SBMNH), Santa Catalina (2 SBMNH)
Range: Endemic (Rifkind, 2019).
Notes. Flightless (Rifkind, 2019).
Cymatodera ovipennis LeConte, 1859
Nomenclatural Authority: Burke, Leavengood & Zolnerowich (2015)
Literature Records: Santa Catalina (Fall, 1897: 238; Fall, 1901: 129)
Digitized Records: none
Range: Also known from mainland (Fall, 1901; Burke, Leavengood & Zolnerowich, 2015).
Melyridae
Notes. Two subfamilies, 29 genera, and 302 species of Melyridae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Dasytinae
Notes. Two tribes, 19 genera, and 211 species of Dasytinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Dasytini
Notes. Eight genera and 40 species of Dasytini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Dasytastes Casey, 1895
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Digitized Records (genus-only): Anacapa (3 SBMNH), San Clemente (3 SBMNH), Santa Barbara (35 SBMNH), Santa Cruz (24 SBMNH), Santa Rosa (2 SBMNH)
Notes. Like most genera of Dasytinae, Dasytastes is in desperate need of revision. Until this is completed, distributional status and endemicity of the species cannot be determined. Forms from San Clemente Island appear externally divergent.
Dasytastes catalinae (LeConte, 1866)
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records: Santa Catalina (LeConte, 1866c: 361; Casey, 1895: 583; Fall, 1901: 128; Miller, 1985a: 20; Caterino & Chandler, 2010: 187)
Digitized Records: Santa Catalina (12 SBMNH)
Range: Endemic (LeConte, 1866c; Casey, 1895; Fall, 1901; Miller, 1985a; Caterino & Chandler, 2010).
Notes. LeConte (1866c) recorded this species as Dasytes catalinae; it was transferred to the genus Dasytastes by Casey (1895). Fall (1901: 128) reported it to be “quite common on various flowers in July”.
Dasytastes insularis Fall, 1901
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records: Santa Catalina (Fall, 1897: 238; Fall, 1901: 251; Miller, 1985a: 20)
Digitized Records: none
Range: Endemic (Fall, 1897, 1901; Miller, 1985a).
Notes. Recorded as “Dasytes, sp. nov.” by Fall (1897).
“Dasytes” Paykull, 1799
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Digitized Records (genus-only): Santa Cruz (6 SBMNH)
Notes. The native North American species currently placed in Dasytes will soon receive a generic reassignment; true Dasytes only occurs in the Palearctic region (M. L. Gimmel, 2022, unpublished data). The North American fauna is in dire need of revision.
Dasytes clementae Fall, 1901
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records: San Clemente (Fall, 1897: 238; Fall, 1901: 251; Miller, 1985a: 20)
Digitized Records: none
Range: Endemic (Fall, 1897, 1901; Miller, 1985a).
Notes. Recorded as “Dasytes, sp. nov.” by Fall (1897).
Eschatocrepis LeConte, 1862
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Notes. This genus contains only one species in North America (M. L. Gimmel, 2022, unpublished data).
Eschatocrepis constrictus (LeConte, 1852)
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records: Santa Catalina (Fall, 1897: 238), Santa Cruz (Fall & Davis, 1934: 144)
Digitized Records: Anacapa (1 SBMNH), San Miguel (3 SBMNH), Santa Cruz (3 LACM; 35 SBMNH), Santa Rosa (5 SBMNH)
Range: Also known from mainland (Howell, 1985).
Listrini
Notes. Eleven genera and 171 species of Listrini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Listrus Motschulsky, 1860
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records (genus-only): San Clemente (Fall, 1897: 238), Santa Barbara (Miller & Miller, 1985: 126), Santa Rosa (Fall, 1897: 238)
Digitized Records (genus-only): Anacapa (1 SBMNH), San Clemente (74 LACM; 23 SBMNH), San Miguel (26 SBMNH), Santa Catalina (11 LACM; 14 SBMNH), Santa Cruz (6 LACM; 60 SBMNH), Santa Rosa (8 SBMNH)
Notes. Reported from Coreopsis (Asteraceae) on Santa Barbara Island by Miller & Miller (1985). This genus currently contains 56 species known to occur in California (M. L. Gimmel, 2022, unpublished data). It is in desperate need of revision.
Listrus anacapensis Blaisdell, 1924
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records: Anacapa (Blaisdell, 1924a: 21; Miller, 1985a: 20)
Digitized Records: none
Range: Endemic (Blaisdell, 1924a; Miller, 1985a).
Notes. This species, along with all other Listrus, was inexplicably moved to the genus Amecocerus by Pic (1937: 98), and was reported as Amecocerus anacapensis by Miller (1985a).
Listrus interruptus LeConte, 1866
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records: Santa Cruz (LeConte, 1866c: 357)
Digitized Records: none
Range: Also known from mainland (Casey, 1895).
Notes. Casey (1895: 547): “This species extends westward to the crests of the Sierras in California, but does not descend the western slope of the mountains.” The Santa Cruz Island record was thought to be incorrect by Fall (1901: 127).
Microasydates Gimmel & Mayor, 2022
Nomenclatural Authority: Gimmel & Mayor (2022)
Notes. This genus contains four species, all of which occur in California (Gimmel & Mayor, 2022). It was revised by Gimmel & Mayor (2022).
Microasydates punctipennis (LeConte, 1866)
Nomenclatural Authority: Gimmel & Mayor (2022)
Literature Records: Santa Catalina (LeConte, 1866c: 355; Casey, 1895: 532; Fall, 1897: 237; Fall, 1901: 126; Gimmel & Mayor, 2022: 555)
Digitized Records: Santa Catalina (1 SBMNH)
Range: Endemic (LeConte, 1866c; Casey, 1895; Fall, 1901; Gimmel & Mayor, 2022); reported from mainland (Blaisdell, 1930: 19), but based on a misidentification of Asydates kumeyaay Mayor & Gimmel, 2019 (Gimmel & Mayor, 2022: 555).
Notes. LeConte (1866c) and Fall (1897) recorded this species as Pristoscelis punctipennis; Casey (1895) and Fall (1901) recorded it as Trichochrous punctipennis.
Microasydates sanclemente Gimmel & Mayor, 2022
Nomenclatural Authority: Gimmel & Mayor (2022)
Literature Records: San Clemente (Gimmel & Mayor, 2022: 556)
Digitized Records: San Clemente (7 SBMNH)
Range: Endemic (Gimmel & Mayor, 2022).
Microasydates santabarbara Gimmel & Mayor, 2022
Nomenclatural Authority: Gimmel & Mayor (2022)
Literature Records: Anacapa (Gimmel & Mayor, 2022: 558), Santa Cruz (Gimmel & Mayor, 2022: 558), Santa Rosa (Gimmel & Mayor, 2022: 558)
Digitized Records: Anacapa (9 SBMNH), Santa Cruz (9 SBMNH; 15 LACM), Santa Rosa (3 LACM)
Range: Also known from mainland (Gimmel & Mayor, 2022).
Pseudasydates Blaisdell, 1938
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Notes. The genus Pseudasydates contains two currently described species in California, although there are additional species waiting to be described (M. L. Gimmel & A. Mayor, 2022, unpublished data).
Pseudasydates explanatus (Casey, 1895)
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records: none
Digitized Records: Santa Catalina (11 LACM)
Range: Also known from mainland (M. L. Gimmel & A. Mayor, 2022, unpublished data).
Notes. The specimens from Santa Catalina Island, collected by George P. Kanakoff (LACM) in 1941, are an unexpected outlier for this otherwise Mojave Desert-Central Valley genus. Effort should be made to recollect it on the island during early spring.
Trichochrous Motschulsky, 1860
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records (genus-only): San Miguel (Miller & Davis, 1986: 550), Santa Barbara (Miller & Miller, 1985: 126)
Notes. Miller & Miller (1985) reported 1–2 species of Trichochrous from Santa Barbara Island, including from Avena (Poaceae) and Frankenia (Frankeniaceae). Miller & Davis (1986) reported two undetermined species of this genus from San Miguel Island occurring on Malacothrix (Malvaceae).
Trichochrous brevicornis (LeConte, 1852)
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records: none
Digitized Records: Santa Cruz (16 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (M. L. Gimmel & A. Mayor, 2022, unpublished data).
Trichochrous calcaratus Fall, 1934
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records: Anacapa (Cockerell, 1940: 285; Miller, 1985a: 20), San Miguel (Cockerell, 1940: 285), Santa Cruz (Fall & Davis, 1934: 144; Cockerell, 1940: 285; Miller, 1985a: 20), Santa Rosa (Fall, 1897: 237; Fall, 1934: 143; Cockerell, 1940: 285; Miller, 1985a: 20)
Digitized Records: Anacapa (12 SBMNH), San Miguel (37 SBMNH), Santa Cruz (11 LACM; 57 SBMNH), Santa Rosa (1 LACM; 35 SBMNH)
Range: Endemic (Fall, 1934; Cockerell, 1940; Miller, 1985a).
Notes. Falls′ (1897) record from Santa Rosa Island was reported as Pristoscelis aenescens [= Trichochrous aenescens (LeConte, 1852)] and almost certainly represents T. calcaratus. LeConte (1866c: 355) recorded P. aenescens from “San Diego and the Islands off Santa Barbara”. Miller & Miller (1985: 132) reported that these records are in error. The populations on the four northern islands may each represent different endemic species (M. L. Gimmel, 2022, unpublished data).
Trichochrous pedalis (LeConte, 1866)
Nomenclatural Authority: M. L. Gimmel & A. Mayor, 2023, unpublished data
Literature Records: Santa Catalina (LeConte, 1866c: 355; Casey, 1895: 529; Fall, 1897: 237; Falls′ 1901: 126; Cockerell, 1940: 285; Miller, 1985a: 20)
Digitized Records: Santa Catalina (9 LACM; 3 SBMNH)
Range: Also known from mainland (M. L. Gimmel & A. Mayor, 2022, unpublished data).
Notes. LeConte (1866c) and Fall (1897) recorded this species as Pristoscelis pedalis; it was transferred to Trichochrous by Casey (1895). Records of this species from San Clemente by Fall (1897: 237), Cockerell (1940: 285), and Miller (1985a: 20) refer to the “undescribed species near pedalis” below. Seavey’s (1892: 262) record of Pristoscelis quadricollis [= Trichochrous quadricollis (LeConte, 1859)] from Santa Catalina is in error and almost certainly represents this species; he reported that it was collected from Heteromeles arbutifolia. Fall (1897: 235) also doubted the validity of this identification. All prior authors have considered T. pedalis to be endemic to Santa Catalina Island; however, individuals from a population occurring in the Palos Verdes Hills in mainland Los Angeles County have been collected which are not diagnosable from the island populations, neither externally nor using male genitalia, which are otherwise diagnostic within the genus (M. L. Gimmel & A. Mayor, 2022, unpublished data).
Trichochrous undescribed species 1 near brevicornis
Literature Records: none
Digitized Records: Anacapa (1 SBMNH), San Miguel (1 LACM) Santa Cruz (4 SBMNH), Santa Rosa (12 SBMNH; 9 LACM)
Range: Endemic (M. L. Gimmel & A. Mayor, 2022, unpublished data).
Notes. In our record set, the identifications of this new species are listed merely as “Trichochrous”.
Trichochrous undescribed species 2 near brevicornis
Literature Records: none
Digitized Records: San Nicolas (19 SBMNH; 1 LACM)
Range: Endemic (M. L. Gimmel & A. Mayor, 2022, unpublished data).
Notes. In our record set, the identifications of this new species are listed merely as “Trichochrous”.
Trichochrous undescribed species near pedalis
Literature Records: San Clemente (Fall, 1897: 237; Cockerell, 1940: 285; Miller, 1985a: 20), Santa Barbara (Fall, 1897: 237)
Digitized Records: San Clemente (35 SBMNH), San Nicolas (40 SBMNH; 58 LACM), Santa Barbara (18 LACM)
Range: Endemic (M. L. Gimmel & A. Mayor, 2022, unpublished data).
Notes. The San Clemente Island records were recorded as T. pedalis by Fall (1897), Cockerell (1940), and Miller (1985a). Fall’s (1897) record from Santa Barbara Island was of Pristoscelis aenescens [= Trichochrous aenescens (LeConte, 1852)] but almost certainly represents this species. In our record set, the identifications of this new species are listed merely as “Trichochrous”.
Malachiinae
Notes. Five tribes, 10 genera, and 91 species of Malachiinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Apalochrini
Notes. One genus and 21 species of Apalochrini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Collops Erichson, 1840
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Digitized Records (genus-only): San Nicolas (1 SBMNH)
Notes. Twenty-one species of Collops have been recorded from California (A. Mayor, 2022, unpublished catalog). The genus-only record from San Nicolas Island above refers to a larval specimen.
Collops cribrosus LeConte, 1852
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Literature Records: Santa Rosa (Fall, 1897: 237)
Digitized Records: San Miguel (1 SBMNH), Santa Cruz (32 SBMNH; 1 UCSB), Santa Rosa (11 SBMNH)
Range: Also known from mainland (A. Mayor, 2022, unpublished catalog).
Notes. Fall (1912) reported that this species lacks hind wings, but observation of specimens in SBMNH reveal that hind wings are present in at least most specimens of both sexes from both island and mainland populations. The wings extend about two-thirds the length of the elytra (M. L. Gimmel, 2021, personal observation).
Collops crusoe Fall, 1910 (Fig. 5)
Figure 5: Collops crusoe Fall, 1910 (Melyridae).
Painting of endemic California Channel Islands species. Painting by Lucie Gimmel.
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Literature Records: San Nicolas (Fall, 1910: 140; Fall, 1912: 269; Cockerell, 1940: 285; Miller, 1985a: 20), Santa Cruz (Miller, 1985a: 20), Santa Rosa (Miller, 1985a: 20)
Digitized Records: San Miguel (2 LACM), San Nicolas (60 LACM; 15 SBMNH; 3 iNat), Santa Cruz (2 LACM), Santa Rosa (1 LACM)
Range: Endemic (Fall, 1910, 1912; Cockerell, 1940; Miller, 1985a).
Notes. Fall (1912) reported that this species lacks hind wings. However, specimens in SBMNH appear to have poorly developed hind wings in both sexes, extending about half the length of the elytra (M. L. Gimmel, 2021, personal observation). Almost certainly they are flightless.
Collops vittatus (Say, 1823)
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Literature Records: none
Digitized Records: Santa Catalina (3 SBMNH)
Range: Also known from mainland (A. Mayor, 2022, unpublished catalog).
Notes. Island specimens of both sexes of this species present in SBMNH are fully winged (M. L. Gimmel, 2021, personal observation).
Attalini
Notes. Three genera and 38 species of Attalini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Attalus Erichson, 1840
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Notes. Seventeen described species of Attalus are known from California (A. Mayor, 2022, unpublished catalog). The two species below may prove to belong to the genus Attalusinus Leng, 1918 (tribe Troglopini) upon further study. The latter genus contains one described species in California (A. Mayor, 2022, unpublished catalog).
Attalus transmarinus Fall, 1898
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Literature Records: San Clemente (Fall, 1897: 237; Fall, 1901: 125; Miller, 1985a: 20)
Digitized Records: none
Range: Endemic (Fall, 1897, 1901; Miller, 1985a).
Notes. Fall (1897) originally recorded this species as Attalus subfasciatus Fall, 1897; this was, however, discovered to be a homonym and replaced with A. transmarinus by Fall (1898b: 267). Champion (1914: 65) subsequently and unnecessarily proposed Attalus falli Champion, 1914 as a replacement name.
Attalus undescribed species
Nomenclatural Authority: A. Mayor (2021, personal communication)
Literature Records: none
Digitized Records: San Clemente (1 iNat)
Range: Endemic (A. Mayor, 2021, personal communication).
Notes. This record is based on the specimen of Attalus on iNaturalist here (www.inaturalist.org/observations/71409011), which has been confirmed to belong to an undescribed species (A. Mayor, 2021, personal communication).
Endeodes LeConte, 1859
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Notes. The genus Endeodes contains three species recorded from California (A. Mayor, 2022, unpublished catalog), all of which are flightless and beach-dwelling, and all of which are represented on the Channel Islands.
Endeodes basalis (LeConte, 1852)
Nomenclatural Authority: Moore & Legner (1975)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: San Clemente (6 SBMNH), San Miguel (15 SBMNH), San Nicolas (14 SBMNH), Santa Cruz (14 SBMNH), Santa Rosa (9 SBMNH)
Range: Also known from mainland (Moore & Legner, 1975).
Notes. This species was reported by Fall (1897) as Endeodes abdominalis (LeConte, 1852), which is now considered a junior synonym of E. basalis (see Moore & Legner, 1975). Some SBMNH specimens from San Miguel and Santa Rosa are nearly all-black (M. L. Gimmel, 2021, personal observation).
Endeodes collaris (LeConte, 1852)
Nomenclatural Authority: Moore & Legner (1975)
Literature Records: Santa Catalina (Fall, 1901: 124), Santa Rosa (Fall, 1897: 237)
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Fall, 1901; Moore & Legner, 1975).
Notes. Fall (1897: 240) noted an undetermined Endeodes from Santa Rosa that was based on one almost entirely black specimen, which may be a color variety of E. collaris.
Endeodes insularis Blackwelder, 1932
Nomenclatural Authority: Moore & Legner (1975)
Literature Records: San Miguel (Blackwelder, 1932: 134; Moore, 1954: 198; Moore & Legner, 1975: 80), Santa Catalina (Fall, 1897: 237)
Digitized Records: San Nicolas (1 SBMNH), Santa Rosa (5 SBMNH)
Range: Also known from mainland (Moore, 1954).
Notes. The Santa Catalina record from Fall (1897) is based on a specimen cited as an undetermined Endeodes taken during July and possessing very minute elytra; this almost certainly refers to E. insularis. This species was originally thought to be endemic to the islands until it was collected on the mainland in Gaviota State Park, Santa Barbara County by Moore (1954).
Ebaeini
Notes. Three genera and five species of Ebaeini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Charopus Erichson, 1840
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Notes. The genus Charopus contains two described species known from California (A. Mayor, 2022, unpublished catalog), plus the undescribed species below.
Charopus undescribed species
Nomenclatural Authority: A. Mayor (2021, personal communication)
Literature Records: Santa Catalina (Fall, 1917: 78)
Digitized Records: Santa Cruz (3 SBMNH)
Range: Also known from mainland (A. Mayor, 2021, personal communication).
Notes. Recorded by Fall (1917) as Microlipus longicollis (now Charopus longicollis Motschulsky, 1860). However, according to A. Mayor (2021, personal communication), the island populations represent an undescribed species of Charopus that also occurs on the mainland.
Malachiini
Notes. Two genera and 26 species of Malachiini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
“Malachius” Fabricius, 1775
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Notes. Twenty-one species of Malachius have been reported from California (A. Mayor, 2022, unpublished catalog). All native North American species currently placed in this genus will soon be reassigned to Hapalorhinus LeConte, 1859.
“Malachius” undetermined species
Literature Records: Santa Rosa (Fall, 1897: 237)
Digitized Records: none
Notes. The genus Malachius has not been recorded from the Channel Islands since Fall’s (1897) record of “Malachius, sp. nov.?” from Santa Rosa (A. Mayor, 2021, personal communication). Fall’s original specimens, if still extant, should be reexamined.
Microlipus LeConte, 1852
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Notes. Five species of the genus Microlipus have been reported from California (A. Mayor, 2022, unpublished catalog).
Microlipus laticeps LeConte, 1852
Nomenclatural Authority: A. Mayor (2022, unpublished catalog)
Literature Records: none
Digitized Records: San Miguel (1 SBMNH), Santa Catalina (1 SBMNH), Santa Cruz (6 SBMNH; 1 UCSB), Santa Rosa (13 SBMNH)
Range: Also known from mainland (A. Mayor, 2022, unpublished catalog).
Trogossitidae
Notes. The family Trogossitidae, as delimited by Gimmel et al. (2019), contains two subfamilies, seven genera, and 26 species in California (Barron, 1971; M. L. Gimmel, 2022, unpublished data). These species were keyed by Barron (1971).
Trogossitinae
Notes. Six genera and 24 species of Trogossitinae are known to occur in California (Barron, 1971; M. L. Gimmel, 2022, unpublished data).
Temnoscheila Westwood, 1830
Nomenclatural Authority: Kolibáč (2013)
Notes. The genus Temnoscheila, often misspelled Temnochila, contains six species in California (Barron, 1971).
Temnoscheila chlorodia (Mannerheim, 1843)
Nomenclatural Authority: Barron (1971), Kolibáč (2013)
Literature Records: Santa Cruz (Barron, 1971: 84)
Digitized Records: Santa Cruz (1 LACM; 1 SBMNH; 6 UCSB)
Range: Also known from mainland (Barron, 1971).
Notes. Barron (1971) recorded this species as Temnochila chlorodia.
Tenebroides Piller & Mitterpacher, 1783
Nomenclatural Authority: Kolibáč (2013)
Notes. The genus Tenebroides contains six species in California (M. L. Gimmel, 2022, unpublished data).
Tenebroides crassicornis (Horn, 1862)
Nomenclatural Authority: Barron (1971), Kolibáč (2013)
Literature Records: Santa Cruz (Barron, 1971: 97)
Digitized Records: none
Range: Also known from mainland (Barron, 1971).
Tenebroides occidentalis Fall, 1910
Nomenclatural Authority: Barron (1971), Kolibáč (2013)
Literature Records: none
Digitized Records: Santa Cruz (9 SBMNH; 2 UCSB)
Range: Also known from mainland (Barron, 1971).
TENEBRIONOIDEA
Anthicidae
Notes. Three subfamilies, 20 genera, and 99 species of Anthicidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Anthicinae
Notes. Twelve genera and 53 species of Anthicinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Amblyderus LaFerté-Sénectère, 1849
Nomenclatural Authority: Chandler (2002)
Notes. Two species of Amblyderus have been recorded from California (Chandler, 1999). The species of this genus were treated for North America by Chandler (1999).
Amblyderus obesus Casey, 1895
Nomenclatural Authority: Chandler (1999)
Literature Records: San Nicolas (Chandler, 1999: 282)
Digitized Records: San Miguel (1 SBMNH), San Nicolas (2 SBMNH)
Range: Also known from mainland (Chandler, 1999).
Amblyderus parviceps Casey, 1895
Nomenclatural Authority: Chandler (1999)
Literature Records: none
Digitized Records: Santa Cruz (5 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Chandler, 1999).
Anthicus Paykull, 1798
Nomenclatural Authority: Chandler (2002)
Notes. Twenty-one species of Anthicus are known to occur in California (M. L. Gimmel, 2022, unpublished data). The species were treated for North America by Werner (1964).
Anthicus cribratus LeConte, 1851
Nomenclatural Authority: Werner (1964)
Literature Records: Santa Cruz (Werner, 1964: 215)
Digitized Records: Santa Cruz (11 SBMNH)
Range: Also known from mainland (Werner, 1964).
Anthicus maritimus LeConte, 1851
Nomenclatural Authority: Werner (1964)
Literature Records: none
Digitized Records: San Nicolas (9 SBMNH)
Range: Also known from mainland (Werner, 1964).
Anthicus nanus LeConte, 1851
Nomenclatural Authority: Werner (1964)
Literature Records: none
Digitized Records: Santa Catalina (3 SBMNH), Santa Cruz (10 SBMNH)
Range: Also known from mainland (Werner, 1964).
Anthicus punctulatus LeConte, 1851
Nomenclatural Authority: Werner (1964)
Literature Records: none
Digitized Records: San Clemente (3 SBMNH), Santa Catalina (9 SBMNH), Santa Cruz (2 SBMNH)
Range: Also known from mainland (Werner, 1964).
Anthicus rufulus LeConte, 1851
Nomenclatural Authority: Werner (1964)
Literature Records: none
Digitized Records: Santa Catalina (5 SBMNH)
Range: Also known from mainland (Werner, 1964).
Cyclodinus Mulsant & Rey, 1866
Nomenclatural Authority: Chandler (2002)
Notes. Five species of Cyclodinus have been recorded from California (Chandler, 2005). The genus was revised for the New World by Chandler (2005).
Cyclodinus annectens (LeConte, 1851)
Nomenclatural Authority: Chandler (2005)
Literature Records: San Clemente (Fall, 1897: 239; Fall, 1901: 181), Santa Catalina (Fall, 1897: 239; Fall, 1901: 181; Chandler, 2005: 7)
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (Fall, 1901; Chandler, 2005).
Notes. Fall (1897, 1901) recorded this species as Anthicus californicus LaFerté-Senéctère, 1849. However, this name actually applies to an eastern North American species now placed in the genus Cyclodinus Mulsant & Rey, 1866 (see Chandler, 2005). The Channel Island records of this species almost certainly pertain to C. annectens.
Ischyropalpus LaFerté-Sénectère, 1849
Nomenclatural Authority: Chandler (2002)
Notes. Five species of Ischyropalpus are known to occur in California (M. L. Gimmel, 2022, unpublished data). The species of this genus were revised for North America by Werner (1973).
Ischyropalpus nitidulus (LeConte, 1851)
Nomenclatural Authority: Werner (1973)
Literature Records: San Clemente (Fall, 1897: 239), Santa Catalina (Fall, 1897: 239; Werner, 1973: 1060), Santa Cruz (Werner, 1973: 1060)
Digitized Records: Anacapa (1 LACM), Santa Catalina (4 LACM), Santa Cruz (20 SBMNH)
Range: Also known from mainland (Werner, 1973).
Notes. Fall (1897) originally reported this species as “Anthicus sp.”, but later redetermined the same specimens in the Museum of Comparative Zoology, Harvard University to Ischyropalpus sturmi (LaFerté-Sénectère, 1849) (S. Miller, 2022, personal communication), California specimens of which are now considered to be I. nitidulus.
Omonadus Mulsant & Rey, 1866
Nomenclatural Authority: Chandler (2002)
Notes. Two adventive species of Omonadus have been recorded from California (Werner, 1964). The species were revised for North America by Werner (1964; as part of Anthicus).
Omonadus floralis (Linnaeus, 1758)
Nomenclatural Authority: Werner (1964), Chandler (2002)
Literature Records: Santa Catalina (Cockerell, 1940: 285)
Digitized Records: none
Range: Also known from mainland (Cockerell, 1940; Werner, 1964).
Notes. This species is introduced from the Old World (Chandler, 2002). Cockerell (1940) recorded it as Hemantus floralis, and Werner (1964) treated it as Anthicus floralis.
Notoxinae
Notes. Two genera and 24 species of Notoxinae have been recorded from California (Chandler, 1983; M. L. Gimmel, 2022, unpublished data).
Notoxus Geoffroy, 1762
Nomenclatural Authority: Chandler (2002)
Digitized Records (genus-only): Santa Catalina (8 LACM), Santa Rosa (1 LACM; 5 SBMNH)
Notes. Twenty-three species of Notoxus have been recorded from California (Chandler, 1983). The North American species were revised by Chandler (1983).
Notoxus desertus Casey, 1895
Nomenclatural Authority: Chandler (1983)
Literature Records: Santa Catalina (Fall, 1897: 238)
Digitized Records: Santa Catalina (8 SBMNH), Santa Cruz (11 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Chandler, 1983).
Notes. This species was recorded as Notoxus constrictus Casey, 1895 by Fall (1897), which is now treated as a junior synonym of N. desertus (see Chandler, 1983: 354).
Notoxus sparsus LeConte, 1859
Nomenclatural Authority: Chandler (1983)
Literature Records: Santa Cruz (Chandler, 1983: 395)
Digitized Records: none
Range: Also known from mainland (Chandler, 1983).
Ciidae
Notes. Two subfamilies, nine genera, and 27 species of Ciidae have been recorded from California (Lawrence, 1982). The North American species were monographed by Lawrence (1971), treated for California by Lawrence (1974), and cataloged for North America by Lawrence (1982). Lopes-Andrade et al. (2016) made additional taxonomic changes affecting the California species.
Ciinae: Ciini
Notes. Two tribes, eight genera, and 26 species of Ciinae, seven genera and 25 species belonging to Ciini, have been recorded from California (Lawrence, 1982).
Ceracis Mellié, 1848
Nomenclatural Authority: Lawrence (1982)
Notes. Two species of Ceracis have been recorded from California (Lawrence, 1982).
Ceracis californicus (Casey, 1884)
Nomenclatural Authority: Lawrence (1971), Lawrence (1974)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Lawrence, 1971, 1974).
Notes. This widespread species has not been reported from the islands in the literature, but is well-known from coastal California and the western half of North America (Lawrence, 1971, 1974).
Cis Latreille, 1796
Nomenclatural Authority: Lawrence (1982)
Notes. Seventeen species of Cis have been recorded from California (Lawrence, 1982).
Cis undetermined species
Literature Records: Santa Catalina (Fall, 1897: 238)
Digitized Records: none
Notes. Fall (1897) recorded “Cis sp.” from Santa Catalina. There are 17 species of this genus reported from California (M. L. Gimmel, 2022, unpublished data), but we know of no additional specimens or literature records from the Channel Islands.
Hadreule Thomson, 1859
Nomenclatural Authority: Lopes-Andrade et al. (2016)
Notes. One species of Hadreule has been recorded from California (Lawrence, 1982). In many publications, including those of Lawrence (1971, 1974, 1982), the genus has been misspelled and misattributed as Hadraule Thomson, 1863, but the proper spelling and attribution are Hadreule Thomson, 1859 (see Lopes-Andrade et al., 2016: 359).
Hadreule blaisdelli (Casey, 1900)
Nomenclatural Authority: Lawrence (1971), Lawrence (1974)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Lawrence, 1971, 1974).
Notes. This widespread species and occasional herbarium pest has not been reported from the islands in the literature, but it is well documented from California and the rest of North America (Lawrence, 1971, 1974).
Orthocis Casey, 1898
Nomenclatural Authority: Lawrence (1982)
Notes. One species of Orthocis has been recorded from California (Lawrence, 1982).
Orthocis punctatus (Mellié, 1848)
Nomenclatural Authority: Lawrence (1971), Lawrence (1974)
Literature Records: none
Digitized Records: Santa Catalina (2 SBMNH), Santa Rosa (7 SBMNH)
Range: Also known from mainland (Lawrence, 1971, 1974).
Notes. This widespread species has not been reported from the islands in the literature, but the town of Santa Barbara was cited as a known locality by Lawrence (1971: 486, 1974: 19).
Sulcacis Dury, 1917
Nomenclatural Authority: Lawrence (1982)
Notes. One species of Sulcacis has been recorded from California (Lawrence, 1982).
Sulcacis curtulus (Casey, 1898)
Nomenclatural Authority: Lawrence (1971), Lawrence (1974)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Lawrence, 1971, 1974).
Notes. This widespread species has not been reported from the islands in the literature, but it is well documented from coastal California and across North America (Lawrence, 1971, 1974).
Meloidae
Notes. Two subfamilies, 18 genera, and 122 species of Meloidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Meloinae
Notes. Twelve genera and 85 species of Meloinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Cordylospasta Horn, 1875
Nomenclatural Authority: Pinto (1972)
Notes. Two species of Cordylospasta have been recorded from California (Pinto, 1972). The genus was revised by Pinto (1972).
Cordylospasta opaca (Horn, 1868)
Nomenclatural Authority: Pinto (1972)
Literature Records: Santa Cruz (Pinto, 1972: 1170)
Digitized Records: none
Range: Also known from mainland (Pinto, 1972).
Epicauta Dejean, 1834
Nomenclatural Authority: Pinto (1991)
Notes. Pinto (1991) provided keys to all North and Central American species of this large genus, of which 25 species have been recorded from California in two subgenera, Epicauta (s.str.) and Macrobasis LeConte, 1862 (M. L. Gimmel, 2022, unpublished data).
Epicauta (Epicauta) puncticollis Mannerheim, 1843
Nomenclatural Authority: Pinto (1991)
Literature Records: Santa Rosa (Ballmer, 1980: 79 [map])
Digitized Records: Santa Rosa (21 LACM; 2 SBMNH)
Range: Also known from mainland (Ballmer, 1980; Pinto, 1991).
Lytta Fabricius, 1775
Nomenclatural Authority: Selander (1960)
Notes. Thirty species of Lytta have been recorded from California, belonging to the subgenera Adicolytta Selander, 1960, Paralytta Selander, 1960, and Poreospasta Horn, 1868 (Selander, 1960). The species of this genus were monographed by Selander (1960).
Lytta (Poreospasta) stygica (LeConte, 1851)
Nomenclatural Authority: Selander (1960)
Literature Records: none
Digitized Records: San Clemente (1 SBMNH)
Range: Also known from mainland (Selander, 1960).
Meloe Linnaeus, 1758
Nomenclatural Authority: Pinto & Selander (1970)
Literature Records (genus-only): Santa Catalina (Fall, 1897: 239)
Notes. Eleven species of Meloe have been recorded from California, belonging to the subgenera Meloe (s.str.) and Treiodous Dugès, 1869 (M. L. Gimmel, 2022, unpublished data). The species of this genus were monographed for the New World by Pinto & Selander (1970). Although listed separately as “Meloe sp.”, Fall’s (1897) record probably refers to M. barbarus.
Meloe (Meloe) strigulosus Mannerheim, 1852
Nomenclatural Authority: Pinto & Selander (1970)
Literature Records: San Miguel (Cockerell, 1940: 285; Pinto & Selander, 1970: 159)
Digitized Records: San Miguel (2 SBMNH)
Range: Also known from mainland (Pinto & Selander, 1970).
Notes. Triungulin larvae of this species have been reported as phoretic both on flower-visiting bee hosts and other taxa such as flies to travel from flower to flower (Pinto et al., 2020). It is worth noting that the single known island that this species inhabits is the only island where M. barbarus is not known to occur.
Meloe (Treiodous) barbarus LeConte, 1861
Nomenclatural Authority: Pinto & Selander (1970)
Literature Records: San Clemente (Van Dyke, 1928: 445; Pinto & Selander, 1970: 120; Huether & Huether, 2015: 162; Miller & Miller, 1985: 128), San Nicolas (Miller & Miller, 1985: 128), Santa Barbara (LeConte, 1861: 354; Van Dyke, 1928: 445; Fall, 1897: 239; Fall, 1901: 183; Cockerell, 1940: 285; Pinto & Selander, 1970: 120; Miller & Miller, 1985: 128), Santa Catalina (Pinto & Selander, 1970: 120; Miller & Miller, 1985: 128), Santa Cruz (Miller & Miller, 1985: 128)
Digitized Records: San Clemente (3 LACM), San Nicolas (1 SBMNH), Santa Catalina (2 LACM), Santa Cruz (1 UCRC), Santa Rosa (1 UCRC)
Range: Also known from mainland (Cockerell, 1940; Pinto & Selander, 1970).
Notes. This species, originally described from Santa Barbara Island and thought to be endemic, was shown to be widespread in the Pacific coast states by Pinto & Selander (1970).
Mordellidae
Notes. Two tribes, four genera, and twenty-six species of Mordellidae are known to occur in California (Bright, 1986; M. L. Gimmel, 2022, unpublished data). The family was revised for North America by Liljeblad (1945) and cataloged for North America by Bright (1986). However, generic concepts around Mordellistena have changed significantly since those publications.
Mordellini
Notes. One genus and three species of Mordellini have been recorded from California (Bright, 1986).
Mordella Linnaeus, 1758
Nomenclatural Authority: Bright (1986)
Notes. Three species of Mordella have been recorded from California (Bright, 1986).
Mordella albosuturalis Liljeblad, 1922
Nomenclatural Authority: Liljeblad (1945)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Liljeblad, 1945).
Mordella hubbsi Liljeblad, 1922
Nomenclatural Authority: Liljeblad (1945)
Literature Records: Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: Santa Cruz (2 LACM; 4 SBMNH)
Range: Also known from mainland (Liljeblad, 1945).
Mordellistenini
Notes. Three genera and 23 species of Mordellistenini are known to occur in California (Bright, 1986).
Mordellina Schilsky, 1908
Nomenclatural Authority: Lisberg (2003)
Notes. Six species now placed in Mordellina have been recorded from California (Bright, 1986; Lisberg, 2003).
Mordellina undetermined species
Literature Records: none
Digitized Records: San Clemente (10 SBMNH), San Miguel (6 LACM), San Nicolas (9 SBMNH), Santa Barbara (3 SBMNH), Santa Catalina (1 SBMNH), Santa Cruz (4 SBMNH), Santa Rosa (5 SBMNH)
Notes. Characters used to separate putative species of Mordellina from Mordellistena in the SBMNH collection were those outlined by Lisberg (2003), who noted that additional species require transfer from Mordellistena to Mordellina. Because of this issue and the lack of a modern revision, no attempt was made to determine these specimens to species.
Mordellistena Costa, 1854
Nomenclatural Authority: Bright (1986)
Notes. Sixteen species currently placed in Mordellistena are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Mordellistena undetermined species
Literature Records: Santa Barbara (Miller & Miller, 1985: 128), Santa Catalina (Fall, 1897: 238), Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: Anacapa (6 SBMNH), San Clemente (1 SBMNH), San Nicolas (1 SBMNH), Santa Barbara (2 SBMNH), Santa Catalina (7 SBMNH), Santa Cruz (3 SBMNH), Santa Rosa (2 SBMNH)
Notes. Fall (1897) listed two separate undetermined species of Mordellistena from Santa Catalina; Miller & Miller (1985) indicated that the Santa Barbara Island species belongs to a group including Mordellistena nubila (LeConte, 1858), Mordellistena ruficeps LeConte, 1862, and Mordellistena subfucus Liljeblad, 1945. MLG notes that there are at least four morphospecies of Mordellistena represented among island material in SBMNH, including an undescribed apterous species based on four specimens from San Clemente (1), San Nicolas (1), and Santa Barbara (2) islands.
Mycetophagidae, NEW FAMILY RECORD
Notes. Two subfamilies, four genera, and seven species of Mycetophagidae have been recorded from California (Parsons, 1975). The North American fauna was revised by Parsons (1975).
Mycetophaginae
Notes. Three genera and six species of Mycetophaginae have been recorded from California (Parsons, 1975).
Litargus Erichson, 1846
Nomenclatural Authority: Young (2002)
Notes. One species of Litargus has been recorded from California (Parsons, 1975).
Litargus balteatus LeConte, 1856
Nomenclatural Authority: Parsons (1975)
Literature Records: none
Digitized Records: Santa Cruz (4 SBMNH), Santa Rosa (6 SBMNH)
Range: Also known from mainland (Parsons, 1975).
Notes. This is a cosmopolitan species (Parsons, 1975), but is presumably native to North America.
Mycetophagus Hellwig, 1792
Nomenclatural Authority: Young (2002)
Notes. Four species of Mycetophagus have been recorded from California, belonging to three subgenera, Gratusus Casey, 1900, Mycetophagus (s.str.), and Parilendus Casey, 1900 (Parsons, 1975).
Mycetophagus (Gratusus) pluriguttatus LeConte, 1856
Nomenclatural Authority: Parsons (1975)
Literature Records: none
Digitized Records: Santa Cruz (4 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Parsons, 1975).
Typhaea Curtis, 1830
Nomenclatural Authority: Young (2002)
Notes. One species of Typhaea has been recorded from California (Parsons, 1975).
Typhaea stercorea (Linnaeus, 1758)
Nomenclatural Authority: Parsons (1975)
Literature Records: none
Digitized Records: Santa Catalina (1 iNat)
Range: Also known from mainland (Parsons, 1975).
Notes. This is a cosmopolitan species (Parsons, 1975), presumably Palearctic in origin.
Mycteridae
Notes. Three subfamilies, three genera, and six species of Mycteridae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Eurypinae
Notes. One genus and species of Eurypinae has been recorded from California (Pollock & Majka, 2012).
Lacconotus LeConte, 1862
Nomenclatural Authority: Pollock & Majka (2012)
Notes. One species of Lacconotus has been recorded from California, belonging to the subgenus Alcconotus Pollock & Majka, 2012 (Pollock & Majka, 2012). The North American species were reviewed by Pollock & Majka (2012).
Lacconotus (Alcconotus) pinicola Horn, 1879
Nomenclatural Authority: Pollock & Majka (2012)
Literature Records: Santa Catalina (Pollock & Majka, 2012: 21), Santa Cruz (Pollock & Majka, 2012: 22; Naughton et al., 2014: 304)
Digitized Records: Santa Catalina (2 SBMNH), Santa Cruz (5 SBMNH)
Range: Also known from mainland (Pollock & Majka, 2012).
Oedemeridae, NEW FAMILY RECORD
Notes. Two subfamilies, 13 genera, and 33 species of Oedemeridae are known to occur in California (M. L. Gimmel, 2022, unpublished data). Most of the North American fauna of Oedemeridae was treated by Arnett (1951).
Oedemerinae
Notes. Eleven genera and 31 species of Oedemerinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Asclerini
Notes. Eight genera and 24 species of Asclerini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Copidita LeConte, 1866
Nomenclatural Authority: Arnett (1951)
Notes. One species of Copidita has been recorded from California (Arnett, 1951).
Copidita quadrimaculata (Motschulsky, 1853)
Nomenclatural Authority: Arnett (1951)
Literature Records: none
Digitized Records: San Clemente (2 SBMNH), San Miguel (2 LACM; 16 SBMNH), San Nicolas (3 LACM; 1 SBMNH), Santa Catalina (4 LACM; 1 SBMNH), Santa Cruz (33 LACM; 3 SBMNH), Santa Rosa (1 LACM; 8 SBMNH)
Range: Also known from mainland (Arnett, 1951).
Notes. This is a widespread beach-dwelling species on the Pacific coast of North America (Arnett, 1951).
Nacerdini
Notes. Two genera and five species of Nacerdini have been recorded from California (Arnett, 1951).
Nacerdes Dejean, 1834
Nomenclatural Authority: Arnett (1951)
Notes. One species of Nacerdes has been recorded from California (Arnett, 1951).
Nacerdes melanura (Linnaeus, 1758)
Nomenclatural Authority: Arnett (1951)
Literature Records: none
Digitized Records: Santa Catalina (1 LACM)
Range: Also known from mainland (Arnett, 1951).
Notes. Known as the wharf borer, this species is adventive in North America, and probably occurs on coasts worldwide (Arnett, 1951).
Xanthochroa Schmidt, 1846
Nomenclatural Authority: Arnett (1951)
Notes. Four species of Xanthochroa have been recorded from California (Arnett, 1951).
Xanthochroa marina Horn, 1896
Nomenclatural Authority: Arnett (1951)
Literature Records: none
Digitized Records: Santa Cruz (8 SBMNH)
Range: Also known from mainland (Arnett, 1951).
Notes. This species, described from Marin County, is known from California and Oregon (Arnett, 1951).
Pyrochroidae, NEW FAMILY RECORD
Notes. Four subfamilies, five genera, and 35 species of Pyrochroidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Pedilinae
Notes. One genus and 25 species of Pedilinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Pedilus Fischer von Waldheim, 1820
Nomenclatural Authority: Bouchard et al. (2011)
Digitized Records (genus-only): Santa Cruz (1 SBMNH; 1 UCSB)
Notes. Twenty-five species of Pedilus are known to occur in California (M. L. Gimmel, 2022, unpublished data). Most of the California species of this genus were treated by Abdullah (1964, 1966, 1969).
Pedilus bardii (Horn, 1874)
Nomenclatural Authority: Abdullah (1966)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Abdullah, 1966).
Salpingidae, NEW FAMILY RECORD
Notes. This family is currently divided into seven subfamilies worldwide and its composition has changed dramatically and often over the last century. Five subfamilies, six genera, and 14 species are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Salpinginae
Notes. Two genera and two or three species of Salpinginae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Rhinosimus Latreille, 1802
Nomenclatural Authority: Bouchard et al. (2011)
Notes. One or two species of this genus have been reported from California (M. L. Gimmel, 2022, unpublished data). The North American species were treated by Blair (1932), but the genus still needs revision. Rhinosimus has a Holarctic distribution.
Rhinosimus undetermined species
Literature Records: none
Digitized Records: Santa Cruz (4 SBMNH)
Notes. Rhinosimus is known from coastal habitats from Alaska south, with the Santa Cruz records representing the southernmost extent of its range known to us.
Scraptiidae
Notes. This family is divided into two subfamilies, Anaspidinae and Scraptiinae, both occurring in California; the former was historically included in Mordellidae (Liljeblad, 1945), while the latter was historically included within Melandryidae (Pollock, 2002). Five genera and 19 species of Scraptiidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Anaspidinae
Notes. Four genera and 17 species of Anaspidinae have been recorded from California (M. L. Gimmel, 2022, unpublished data). These were treated for North America by Liljeblad (1945).
Anaspis Geoffroy, 1762
Nomenclatural Authority: Pollock (2002)
Digitized Records (genus-only): Santa Cruz (14 SBMNH)
Notes. Seven species of Anaspis have been recorded from California (Liljeblad, 1945).
Anaspis atrata Champion, 1891
Nomenclatural Authority: Liljeblad (1945)
Literature Records: none
Digitized Records: Santa Cruz (2 LACM; 6 SBMNH)
Range: Also known from mainland (Liljeblad, 1945).
Anaspis collaris LeConte, 1851
Nomenclatural Authority: Liljeblad (1945)
Literature Records: Santa Catalina (Fall, 1897: 238; Liljeblad, 1945: 216)
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (Liljeblad, 1945).
Pentaria Mulsant, 1856
Nomenclatural Authority: Pollock (2002)
Notes. Five species of Pentaria have been recorded from California (Liljeblad, 1945).
Pentaria trifasciata (Melsheimer, 1845)
Nomenclatural Authority: Liljeblad (1945)
Literature Records: Santa Catalina (Fall, 1897: 238)
Digitized Records: San Nicolas (1 SBMNH), Santa Catalina (7 SBMNH), Santa Cruz (5 LACM; 6 SBMNH)
Range: Also known from mainland (Liljeblad, 1945).
Notes. Liljeblad (1945) treated this as an extremely variable and widespread species, including Pentaria trifasciata nubila (LeConte, 1859) as a variety representing a color morph. Fall (1897) reported this species as Pentaria nubila. We have seen specimens from Santa Cruz Island corresponding to both color morphs in the sense of Liljeblad (1945).
Tenebrionidae
Notes. This family has a recent and reliable catalog for North America by Bousquet et al. (2018) which serves as a good starting point for taxonomy and finding identification references. The constituent groups that comprise the Channel Island diversity vary in accessibility of authoritative treatments and identification resources. Many of the tribes and genera require a thorough revision before the species reported from the islands can be truly verified. Eight subfamilies, 37 tribes, 110 genera, 525 species of Tenebrionidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Alleculinae
Notes. Six genera and 33 species of Alleculinae, all belonging to the tribe Alleculini, have been recorded from California (Bousquet et al., 2018). “Alleculinae larvae” were reported from San Clemente and Santa Catalina by Straughan & Hadley (1980: 392).
Hymenorus Mulsant, 1852
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records (genus-only): Santa Cruz (Naughton et al., 2014: 304)
Notes. This genus was last revised by Fall (1931) and requires significant work, at least for species in western North America. There are 19 species reported from California (Bousquet et al., 2018).
Hymenorus infuscatus Casey, 1891
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Fall, 1897: 238; Fall, 1901: 176; Fall, 1931: 186)
Digitized Records: none
Range: Also known from mainland (Fall, 1931).
Notes. The southern California species are difficult to identify. We have not seen any Hymenorus specimens from the Channel Islands, but trust the records given by Fall who completed the last revision of this genus.
Isomira Mulsant, 1856
Nomenclatural Authority: Bousquet et al. (2018)
Digitized Records (genus-only): San Clemente (16 SBMNH), San Nicolas (2 SBMNH), Santa Catalina (9 SBMNH), Santa Cruz (21 SBMNH), Santa Rosa (11 SBMNH)
Notes. This genus was revised in the dissertation of Marshall (1964) which was subsequently published in numerous parts except for a final key and treatments of the western species. Species are difficult to identify without comparative material and dissected males (M. A. Johnston, 2022, personal observation). There are five species recorded from California (Bousquet et al., 2018).
Isomira comstocki Papp, 1956
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Cruz (Marshall, 1964:145)
Digitized Records: Santa Barbara (1 LACM), Santa Cruz (4 SBMNH)
Range: Also known from mainland (Marshall, 1964).
Notes. This widespread and variable species is primarily identified by dissected male genitalia.
Isomira damnata Marshall, 1970
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Santa Catalina (1 LACM)
Range: Also known from mainland (Marshall, 1964).
Notes. The single Channel Island specimen of this species is a male with extruded genitalia that was reliably determined by J.M. Campbell.
Isomira luscitiosa Casey, 1891
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Santa Cruz (5 LACM), Santa Rosa (2 LACM)
Range: Also known from mainland (Marshall, 1964).
Notes. The specimens representing this species were reliably determined by J.M. Campbell.
Isomira variabilis (Horn, 1875)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Clemente (Fall, 1897: 238; Fall, 1901: 176)
Digitized Records: none
Range: Also known from mainland (Marshall, 1964).
Notes. Marshall (1964) recognized a species group that most workers before him had lumped into a single concept of I. variabilis, in which he included I. damnata and I. luscitiosa. It is possible that Fall (1897, 1901) was actually referring to one of the latter two species. However, I. variabilis is widespread and known from the coastal habitats of southern California and may well be the correct identification.
Mycetochara Guérin-Méneville, 1827
Nomenclatural Authority: Bousquet et al. (2018)
Digitized Records (genus-only): Santa Catalina (4 SBMNH), Santa Cruz (7 SBMNH)
Notes. Campbell (1978) revised this genus and synonymized many of the previously described species along with providing a reliable key. Three species are recorded from California (Bousquet et al., 2018).
Mycetochara pubipennis LeConte, 1878
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Campbell, 1978: 936)
Digitized Records: none
Range: Also known from mainland (Campbell, 1978).
Notes. Campbell (1978) only saw male specimens from the island and noted that the eyes are slightly smaller than those on the mainland, a character which is traditionally used as part of species diagnoses.
Blaptinae
Notes. Three tribes, 18 genera, and 122 species of Blaptinae are known to occur in California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data).
Amphidorini
Notes. Six genera and 89 species of Amphidorini are known to occur in California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data). This tribe has seen recent treatments for many constituent species groups but has lacked a thorough revision of the genera and subgenera. The nomenclaturally unavailable dissertation of Johnston (2018) provides keys, diagnoses, and new concepts of genera and subgenera. Since this study has not been published in compliance with the ICZN, the taxonomy follows Bousquet et al. (2018) where most species are included in the large genus Eleodes.
Eleodes Eschscholtz, 1829
Nomenclatural Authority: Bousquet et al., 2018
Digitized Records (genus-only): Anacapa (1 LACM), San Clemente (19 LACM; 1 UCMC), San Miguel (38 LACM; 3 SDNHM), San Nicolas (27 LACM), Santa Catalina (1 OSUC; 54 LACM), Santa Cruz (25 LACM; 1 UCSB; 1 YPMC; 1 iNat), Santa Rosa (31 LACM)
Notes. The nomenclaturally unavailable dissertation by Johnston (2018) breaks this genus into multiple genera and it is expected that the nomenclature reported here will soon be outdated. Digitized records identified as Amphidora Eschscholtz, 1829, a current subgenus of Eleodes, are included in the genus-only records above. This large genus of flightless arid-adapted species is restricted to western North America and has its center of diversity in California with 67 species recorded from the state. A possible fossil of Eleodes was reported by Lipps (1964) from a deposit on West Anacapa Island, though it is likely to be a modern contaminant that made its way into the sandy deposit.
Eleodes (Amphidora) littoralis (Eschscholtz, 1829)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Fall, 1897: 238), Santa Cruz (Fall & Davis, 1934: 144), Santa Rosa (Fall, 1897: 238)
Digitized Records: Anacapa (14 LACM; 1 SBMNH), San Clemente (5 LACM), San Miguel (3 SBMNH), Santa Catalina (2 OSUC; 24 LACM; 26 SBMNH), Santa Cruz (5 SBMNH), Santa Rosa (9 SBMNH)
Range: Also known from mainland (Blaisdell, 1909).
Notes. This small, hirsute species is common in leaf litter along the coastal mountain ranges of California.
Eleodes (Amphidora) nigropilosa (LeConte, 1851)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Straughan & Hadley, 1980: 392)
Digitized Records: Santa Catalina (2 LACM; 1 MAJC; 2 OSUC; 6 SBMNH), Santa Cruz (3 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Blaisdell, 1909; Triplehorn, 1996).
Notes. This species is common in coastal habitats in California and Baja California.
Eleodes (Blapylis) clavicornis Eschscholtz, 1829
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Anacapa (3 LACM)
Range: Also known from mainland (Somerby, 1972).
Notes. This relatively small species is known from coastal sand dunes.
Eleodes (Blapylis) inculta LeConte, 1861
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Anacapa (Somerby, 1972: 179; Miller, 1985a: 21), San Miguel (Blaisdell, 1918: 384; Spilman, 1962: 57; Tanner, 1961: 73; Somerby, 1972: 179; Miller, 1985a: 21), Santa Barbara (LeConte, 1861: 352; Miller, 1985a: 21; Miller & Miller, 1985: 128), Santa Cruz (Blaisdell, 1918: 384; Fall & Davis, 1934: Tanner, 1961: 73; 144; Somerby, 1972: 179; Miller, 1985a: 21; Naughton et al., 2014: 304), Santa Rosa (Blaisdell, 1909: 331; Blaisdell, 1918: 384; Blaisdell, 1939: 52; Spilman, 1962: 57; Somerby, 1972: 179; Miller, 1985a: 21)
Digitized Records: Anacapa (1 BYUC; 10 LACM; 1 SBMNH), San Miguel (1 BYUC; 3 OSUC; 131 LACM; 2 MAJC; 76 SBMNH), Santa Catalina (2 OSUC; 1 LACM), Santa Cruz (1 BYUC; 2 OSUC; 19 LACM; 10 MAJC; 1 SBMNH; 1 iNat), Santa Rosa (20 LACM; 6 MAJC; 16 SBMNH)
Range: Endemic (LeConte, 1861; Blaisdell, 1909; Spilman, 1962; Somerby, 1972).
Notes. Blaisdell (1909) incorrectly documented the type locality as Santa Rosa Island, while LeConte (1861) stated that it was Santa Barbara Island. Somerby (1972) and Miller & Miller (1985) suggested that the type may have been mislabeled and was actually from Santa Cruz Island. The subspecies E. inculta affinis Blaisdell, 1918 was reported from Santa Cruz and San Miguel islands by Blaisdell (1918) and later synonymized by Miller (1985a) following the nomenclaturally unavailable thesis of Somerby (1972). A single record of the species Eleodes cordata Eschscholtz, 1829 from Santa Cruz Island in the BYUC is here considered to almost certainly represent E. inculta. Eleodes cordata is otherwise not known from southern California.
Eleodes (Blapylis) scabripennis LeConte, 1859
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Barbara (Fall, 1897: 238; Fall, 1901: 168), Santa Rosa (Fall, 1897: 238; Fall, 1901: 168)
Digitized Records: none
Range: Also known from mainland (Somerby, 1972).
Notes. This species was described from Fort Tejon, Kern County, California. The female holotype has a comparatively small pronotum but is quite similar to Eleodes (Blapylis) consobrina LeConte, 1851 (LeConte, 1861; Blaisdell, 1909). It may be that E. scabripennis is simply a synonym of E. consobrina, but it does superficially resemble E. inculta quite closely. The above records are almost certainly from Fall examining LeConte’s material, and applying this name to E. inculta. However, until the genus is revised, these records cannot be fully discounted.
Eleodes (Blapylis) subvestita (Blaisdell, 1939)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Nicolas (Blaisdell, 1939: 55; Cockerell, 1939: 317; Cockerell, 1940: 284; Steele, 1979: 30; Spilman, 1962: 57; Somerby, 1972: 180; Miller, 1985a: 21)
Digitized Records: San Nicolas (26 LACM; 1 SBMNH)
Range: Endemic (Blaisdell, 1939; Spilman, 1962; Somerby, 1972).
Notes. The original description was apparently made using a combination of this actual Channel Island endemic Eleodes and a genitalic dissection belonging to a specimen of another subfamily, and thus was placed into a new monotypic genus and subfamily, before the error was discovered and the species placed correctly into the genus Eleodes subgenus Blapylis Horn, 1870 (Spilman, 1962). Accordingly, Blaisdell (1939), Cockerell (1939, 1940), and Steele (1979) reported the original combination of Eleodopsis subvestita Blaisdell, 1939.
Eleodes (Cratidus) osculans (LeConte, 1851)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Cruz (LeConte, 1876: 299; Fall, 1897: 238; Fall & Davis, 1934: 144), Santa Rosa (Fall, 1897: 238)
Digitized Records: San Miguel (2 SBMNH), Santa Catalina (5 SBMNH; 2 iNat), Santa Cruz (1 LACM), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Blaisdell, 1909; Triplehorn, 1996).
Notes. This somewhat charismatic beetle is abundant in southern California. LeConte (1876), Fall (1897), and Fall & Davis (1934) recorded this species as Cratidus osculans.
Eleodes (Eleodes) acuticauda LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Anacapa (Blaisdell, 1921: 219; Cockerell, 1940: 284; Miller, 1985a: 21), San Clemente (Blaisdell, 1909: 283; Blaisdell, 1921: 219; Doyen, 1974: 87; Miller, 1985a: 21), San Miguel (Blaisdell, 1921: 219; Cockerell, 1940: 284; Miller, 1985a: 21), San Nicolas (Blaisdell, 1921: 219; Cockerell, 1940: 284; Tanner, 1961: 73; Triplehorn, Thomas & Smith, 2015: 161), Santa Barbara (Blaisdell, 1921: 219; Cockerell, 1940: 284; Miller, 1985a: 21; Miller & Miller, 1985: 128), Santa Cruz (Miller, 1985a: 21), Santa Rosa (Miller, 1985a: 21)
Digitized Records: Anacapa (3 LACM; 3 SBMNH), San Clemente (5 LACM; 6 MAJC; 5 SBMNH; 13 SDNHM), San Miguel (10 LACM; 19 SBMNH), San Nicolas (36 LACM; 24 SBMNH), Santa Barbara (16 LACM), Santa Catalina (5 SBMNH), Santa Cruz (7 LACM; 3 MAJC; 2 SBMNH), Santa Rosa (1 LACM; 17 SBMNH)
Range: Also known from mainland (Triplehorn, Thomas & Smith, 2015).
Notes. The recent revision by Triplehorn, Thomas & Smith (2015) only mentioned records from San Nicolas Island, but did not give specimen locality data other than types for any of the taxa in the revision; the accompanying distribution map shows a record for San Nicolas and no other islands. This species is abundant in southern California and is difficult to separate from E. dentipes in the vicinity of the city of Santa Barbara; see the remarks under that species. The name Eleodes laticollis LeConte, 1851 was first synonymized with E. acuticauda by Horn (1870) but subsequently used as a subspecies in the island records of Blaisdell (1909) and Doyen (1974). The subspecies E. laticollis apprima Blaisdell, 1921 was erected for the Channel Islands populations and used by Blaisdell (1921), Cockerell (1940), Tanner (1961), and Miller (1985a) before it was synonymized with E. acuticauda by Triplehorn (1996).
Eleodes (Eleodes) dentipes Eschscholtz, 1829
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Anacapa (Triplehorn, Thomas & Smith, 2015: 165 [map]), San Clemente (Fall, 1897: 238), San Nicolas (Fall, 1897: 238), Santa Cruz (Fall & Davis, 1934: 144), Santa Rosa (Fall, 1897: 238)
Digitized Records: none
Range: Also known from mainland (Triplehorn, Thomas & Smith, 2015).
Notes. The recent revision of Triplehorn, Thomas & Smith (2015) gave no specimen locality data for this or any other treated taxa. However, their accompanying distribution map has a mark on Anacapa Island. This species and E. acuticauda are quite similar and can be difficult to distinguish in the region of Santa Barbara County where the two species distributions meet. These two species and island populations along with the mainland populations need to be carefully considered. The two species are distinguished by E. acuticauda having a much more transverse and strongly rounded pronotum than E. dentipes which reliably separates most species from the northern Central Valley and Bay Area of California from those around San Diego and Los Angeles. These literature records presumably overlap with the digitized records of E. acuticauda, but the species are retained as separate since the last revision (Triplehorn, Thomas & Smith, 2015) indicated that both species are known from the Channel Islands.
Eleodes (Melaneleodes) carbonaria (Say, 1824)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Fall, 1897: 238; Fall, 1901: 167; Blaisdell, 1909: 75; Cockerell, 1940: 284; Tanner, 1961: 69)
Digitized Records: San Clemente (1 LACM), Santa Catalina (1 LACM; 8 SBMNH), Santa Rosa (3 LACM)
Range: Also known from mainland (Triplehorn & Thomas, 2012).
Notes. This species has been well documented from Santa Catalina island under several different species names. Originally referred to Eleodes quadricollis Eschscholtz, 1829 by Fall (1897, 1901) (only known from around San Francisco as currently circumscribed), Blaisdell (1909) and Tanner (1961) recognized the island population as Eleodes omissa forma catalinae Blaisdell, 1909, which was later included within the subspecies concept of Eleodes carbonaria omissa LeConte, 1858 of Triplehorn & Thomas (2012), which is the only subspecies distributed throughout southern California and Baja California. Cockerell (1940: 284) listed this species twice under the names E. omissa catalinae and E. omissa pygmaea Blaisdell, 1909, both of which are now considered synonyms of E. carbonaria omissa (Triplehorn & Thomas, 2012).
Eleodes (Steneleodes) gigantea Mannerheim, 1843
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: San Miguel (43 SBMNH; 4 SDNHM), Santa Rosa (6 LACM; 3 SBMNH)
Range: Also known from mainland (Blaisdell, 1909).
Notes. This species, abundant in the coastal ranges, is known from California and Baja California but has never been reported from the Channel Islands in the literature.
Opatrini
Notes. Eleven genera and 31 species of Opatrini are known to occur in California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data).
Blapstinus Dejean, 1821
Nomenclatural Authority: Davis (1970), Bousquet et al. (2018)
Literature Records (genus-only): Santa Cruz (Fall & Davis, 1934: 144)
Digitized Records (genus-only): San Clemente (1 SBMNH), Santa Catalina (13 LACM), Santa Rosa (9 LACM; 2 SBMNH)
Notes. Fall & Davis (1934) stated that their record was not Blapstinus rufipes (= B. discolor) or B. brevicollis. The nomenclaturally unavailable dissertation of Davis (1970, 1976) provides keys and treatments for the genus but many specimens in collections are dubiously identified, particularly if done before 1970 (M. A. Johnston, 2022, personal observation). Fifteen species have been recorded from California (Bousquet et al., 2018).
Blapstinus angustus LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: San Clemente (1 SDNHM)
Range: Also known from mainland (Bousquet et al., 2018).
Notes. This is the type species of the genus Mecysmus Horn, 1870, where it has been included since the genus description until recently, when Lumen et al. (2019b) synonymized it with Blapstinus. The species is known from throughout southern California and western Arizona.
Blapstinus brevicollis LeConte, 1851
Nomenclatural Authority: Davis (1970), Bousquet et al. (2018)
Literature Records: Santa Cruz (Davis, 1970: 138), Santa Rosa (Fall, 1897: 238; in doubt, Davis, 1970: 138)
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (4 SBMNH), Santa Rosa (9 SBMNH)
Range: Also known from mainland (Davis, 1970).
Notes. This species is common in southern California and Arizona and has been collected from driftwood (Davis, 1970: 136). The thorough yet nomenclaturally unavailable dissertation of Davis (1970) clearly indicated two island records on the distribution map (see his figure 143), and in the material examined lists both “Santa Cruz” and “Santa Rosa Island” as localities from “Los Angeles County, California” which are interpreted as the two island records listed above.
Blapstinus discolor Horn, 1870
Nomenclatural Authority: Davis (1970), Bousquet et al. (2018)
Literature Records: Santa Catalina (Fall, 1897: 238; Baker, 1905: 59; Davis, 1970: 302)
Digitized Records: San Miguel (1 SBMNH), Santa Cruz (10 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Davis, 1970).
Notes. This flightless and somewhat variable species is widely distributed throughout the western United States and is a common pest of crops along the southern California coast. Davis (1970) reported this species from “Avalon” (Santa Catalina Island) in the material examined but did not give a distinct island marker on the distribution map (his figure 163). Fall (1897) and Baker (1905) recorded this species as Blapstinus rufipes Casey, 1890 which was synonymized with B. discolor by Davis (1982).
Conibius LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus of flightless beetles has two species recorded from California (Bousquet et al., 2018).
Conibius seriatus LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: San Clemente (1 SBMNH), Santa Catalina (2 SBMNH), Santa Cruz (2 SBMNH)
Range: Also known from mainland (Bousquet et al., 2018).
Notes. This species can be commonly found under stones or crawling on the ground at night.
Tonibius Casey, 1895
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus is monotypic with its single species known from California (Bousquet et al., 2018).
Tonibius sulcatus (LeConte, 1851)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Clemente (Fall, 1897: 238; Fall, 1901: 172)
Digitized Records: San Clemente (1 LACM; 3 SBMNH; 8 SDNHM)
Range: Also known from mainland (Bousquet et al., 2018).
Notes. This species is common in arid and coastal habitats in California and Baja California. Fall (1897, 1901) recorded this species as Notibius sulcatus, its original combination.
Ulus Horn, 1870
Nomenclatural Authority: Lumen et al. (2019a), Bousquet et al. (2018)
Notes. Two species of Ulus are known from California (Lumen et al., 2019a).
Ulus crassus (LeConte, 1851)
Nomenclatural Authority: Lumen et al. (2019a)
Literature Records: Santa Cruz (Fall, 1934: 144)
Digitized Records: none
Range: Also known from mainland (Lumen et al., 2019a).
Notes. This species is fairly common throughout southern California but the recent revision of Lumen et al. (2019a) gave no Channel Island records and none have been found in museums. It seems unlikely that Fall (1934) would have confused this genus with anything else, but this record certainly requires verification.
Diaperinae
Notes. Five tribes, 13 genera, and 29 species of Diaperinae have been recorded from California (Bousquet et al., 2018).
Diaperini
Notes. Eight genera and 15 species of Diaperini have been recorded from California (Bousquet et al., 2018).
Platydema Laporte & Brullé, 1831
Nomenclatural Authority: Bousquet et al. (2018)
Notes. Four species of Platydema are recorded from California (Bousquet et al., 2018).
Platydema oregonensis LeConte, 1857
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Santa Cruz (4 SBMNH)
Range: Also known from mainland (Bousquet et al., 2018).
Notes. This species is widespread throughout the Pacific coastal region and has been recorded as Platydema oregonense since the time of its description until Bousquet et al. (2018) corrected the gender of the genus.
Hypophlaeini
Notes. One genus and six species of Hypophlaeini have been recorded from California (Bousquet et al., 2018).
Corticeus Piller & Mitterpacher, 1783
Nomenclatural Authority: Bousquet et al. (2018)
Notes. Six species of Corticeus have been recorded from California (Bousquet et al., 2018). The species were reviewed for North America by Triplehorn (1990).
Corticeus opaculus (LeConte, 1878)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Cruz (Triplehorn, 1990: 294)
Digitized Records: Santa Cruz (1 OSUC)
Range: Also known from mainland (Triplehorn, 1990).
Notes. This subcortical species is not uncommon in coastal California habitats.
Phaleriini
Notes. Two genera and six species of Phaleriini have been recorded from California (Bousquet et al., 2018).
Phaleria Latreille, 1802
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus inhabits coastal dunes around the world and can be readily found under beach wrack and dead fish. The New World components were revised by Triplehorn & Watrous (1979). A single species is known from California (Bousquet et al., 2018).
Phaleria rotundata LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Anacapa (Triplehorn & Watrous, 1979: 284), San Nicolas (Triplehorn & Watrous, 1979: 286), Santa Catalina (Triplehorn & Watrous, 1979: 286), Santa Cruz (Triplehorn & Watrous, 1979: 286)
Digitized Records: Anacapa (2 LACM), San Clemente (6 SBMNH), San Nicolas (1 LACM; 3 SBMNH), Santa Catalina (3 SBMNH), Santa Cruz (14 SBMNH), Santa Rosa (8 SBMNH)
Range: Also known from mainland (Triplehorn & Watrous, 1979).
Notes. This species has a somewhat restricted range, from northern Baja California through San Francisco, and is the only species of this genus known from the United States Pacific coastline.
Pimeliinae
Notes. Eleven tribes, 41 genera, and 248 species of Pimeliinae are known to occur in California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data).
Anepsiini
Notes. Five genera and nine species of Anepsiini have been recorded from California (Bousquet et al., 2018).
Batuliodes Casey, 1907
Nomenclatural Authority: Doyen (1987)
Notes. This genus, and tribe, was thoroughly revised by Doyen (1987) who provided reliable keys to genera and species. Four species of Batuliodes are known from California (Doyen, 1987).
Batuliodes rotundicollis (LeConte, 1851)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: San Clemente (1 SBMNH)
Range: Also known from mainland (Doyen, 1987).
Notes. This small and relatively infrequently collected species was not recorded from the Channel Islands in the revision by Doyen (1987).
Cnemeplatiini
Notes. Two genera and eight species of Cnemeplatiini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Alaudes Horn, 1870
Nomenclatural Authority: Aalbu, Caterino & Smith (2018)
Notes. Individuals of Alaudes are the smallest of all known Tenebrionidae from the Channel Islands; they were revised recently by Aalbu, Caterino & Smith (2018). Six species of Alaudes are recorded from California (Aalbu, Caterino & Smith, 2018).
Alaudes singularis Horn, 1870
Nomenclatural Authority: Aalbu, Caterino & Smith (2018)
Literature Records: San Clemente (Aalbu, Caterino & Smith, 2018: 265), San Nicolas (Aalbu, Caterino & Smith, 2018: 265)
Digitized Records: San Clemente (2 SBMNH)
Range: Also known from mainland (Aalbu, Caterino & Smith, 2018).
Lepidocnemeplatia Bousquet & Bouchard, 2018
Nomenclatural Authority: Bousquet et al. (2018)
Notes. Only a single species of Lepidocnemeplatia is known from California (Bousquet et al., 2018).
Lepidocnemeplatia sericea (Horn, 1870)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Santa Cruz (10 SBMNH)
Range: Also known from mainland (Bousquet et al., 2018).
Notes. This species is widespread in the arid regions of western North America and is particularly prevalent in habitats with sandy substrate.
Coniontini
Notes. Three genera and 63 species of Coniontini have been recorded from California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data).
Coelus Eschscholtz, 1829
Nomenclatural Authority: Doyen (1976)
Digitized Records (genus-only): San Clemente (2 SBMNH), Santa Catalina (1 TAMU), Santa Cruz (1 TAMU)
Notes. Doyen (1976) revised Coelus and provided a reliable key. Chatzimanolis, Norris & Caterino (2010) explored the historical biogeography and phylogenetic relationships of Channel Island and mainland coastal populations of this genus. Four species are known from California, three of which occur on the islands (Doyen, 1976).
Coelus ciliatus Eschscholtz, 1829
Nomenclatural Authority: Doyen (1976), Bousquet et al. (2018)
Literature Records: Anacapa (Doyen, 1976: 616)
Digitized Records: San Nicolas (1 CASC), Santa Cruz (1 CASC), Santa Rosa (3 OSUC)
Range: Also known from mainland (Doyen, 1976).
Notes. This species is abundant and widely distributed along the mainland Pacific Coast.
Coelus globosus LeConte, 1851
Nomenclatural Authority: Doyen (1976), Bousquet et al. (2018)
Literature Records: Anacapa (Doyen, 1976: 618), San Miguel (Doyen, 1976: 618), San Nicolas (Doyen, 1976: 618; Chatzimanolis, Norris & Caterino, 2010: 787), Santa Barbara (Doyen, 1976: 618), Santa Catalina (Doyen, 1976: 618), Santa Cruz (Doyen, 1976: 618; Chatzimanolis, Norris & Caterino, 2010: 787), Santa Rosa (Doyen, 1976: 618)
Digitized Records: Anacapa (7 LACM), San Miguel (1 LACM; 4 SBMNH), San Nicolas (104 LACM; 1 SBMNH), Santa Cruz (2 MAJC; 10 SBMNH; 4 UCMC), Santa Rosa (6 LACM; 3 SBMNH)
Range: Also known from mainland (Doyen, 1976).
Notes. This coastal dune-inhabiting species is known from most of California’s shoreline including all Channel Islands except San Clemente (Doyen, 1976: 618).
Coelus pacificus Fall, 1897
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Anacapa (Doyen, 1976: 623; Miller, 1985a: 21), San Clemente (Fall, 1897: 238; Fall, 1901: 166; Casey, 1908: 158; Blaisdell, 1919: 321; Doyen, 1974: 87; Doyen, 1976: 623; Miller, 1985a: 21; Chatzimanolis, Norris & Caterino, 2010: 787), San Miguel (Blaisdell, 1919: 321; Doyen, 1976: 623; Miller, 1985a: 21; Chatzimanolis, Norris & Caterino, 2010: 787), San Nicolas (Fall, 1897: 238; Casey, 1908: 158; Blaisdell, 1919: 321; Doyen, 1976: 623; Miller, 1985a: 21; Chatzimanolis, Norris & Caterino, 2010: 787), Santa Barbara (Fall, 1901: 166), Santa Catalina (Miller, 1985a: 21; Chatzimanolis, Norris & Caterino, 2010: 787), Santa Cruz (Blaisdell, 1919: 321; Fall & Davis, 1934: 144; Doyen, 1976: 623; Miller, 1985a: 21; Chatzimanolis, Norris & Caterino, 2010: 787), Santa Rosa (Fall, 1897: 238; Fall, 1901: 166; Blaisdell, 1919: 321; Doyen, 1976: 623; Miller, 1985a: 21; Chatzimanolis, Norris & Caterino, 2010: 787)
Digitized Records: San Clemente (292 LACM; 16 SBMNH; 5 SDNHM), San Miguel (39 CASC; 264 LACM; 144 SBMNH), San Nicolas (7 CASC; 103 LACM; 33 SBMNH; 3 iNat), Santa Barbara (5 LACM), Santa Catalina (3 LACM; 14 SBMNH), Santa Cruz (1 OSUC; 59 CASC; 10 LACM; 4 MAJC; 27 SBMNH), Santa Rosa (1 OSUC; 5 CASC; 69 LACM; 18 SBMNH; 1 USNM; 1 iNat)
Range: Endemic (Fall, 1897; Doyen, 1976).
Notes. A larva and pupa from San Clemente Island were studied by Doyen (1976: 611–612), who also synonymized Coelus remotus Fall, 1897 with C. pacificus. Unfortunately he merely stated that this species is distributed on the “California Channel Islands” (Doyen, 1976: 619) and beyond this only gave type specimen island records and several in the appendix. This species is recorded from all eight Channel Islands. The type locality of C. pacificus is San Nicolas Island (Fall, 1897). The name C. remotus, whose type locality is San Clemente Island, was used for the San Clemente population by Fall (1897, 1901), Casey (1908), Blaisdell (1919), and Doyen (1974).
Coniontis Eschscholtz, 1829
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records (genus-only): Santa Cruz (Fall, 1934: 144; Naughton et al., 2014: 304)
Digitized Records (genus-only): San Clemente (1 SDNHM; 4 iNat), San Nicolas (1 SBMNH; 1 iNat), Santa Catalina (15 SBMNH; 2 iNat), Santa Cruz (3 CSUC; 4 SBMNH), Santa Rosa (12 SBMNH; 2 SDNHM)
Notes. Coniontis is a problematic genus which requires a comprehensive revision, though one may start with the synonymy established by Doyen (1977), which included no identification resources, in conjunction with voluminous species descriptions provided by Casey (1908), or instead attempt to use the outdated treatment by Horn (1870). It seems clear that there is at least one endemic species (C. lata LeConte, 1866) and likely a second (C. santarosae Blaisdell, 1921) but the literature records and identified museum specimen records should otherwise all be considered dubious until the genus is revised. One Santa Cruz Island specimen labeled “Coniontis musculus ?” in the H.C. Fall collection in the Museum of Comparative Zoology, Harvard University was determined by H.C. Fall (S. Miller, 2022, personal communication) and it alludes to C. muscula Blaisdell, 1918; this questionable species record is probably the origin of the genus-only citation above. Forty-nine currently valid species are recorded from California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data).
Coniontis elliptica Casey, 1884
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Fall, 1897: 238; Fall, 1901: 165; Baker, 1905: 59; Casey, 1908: 88; Doyen, 1977: 2), Santa Rosa (Fall, 1897: 238)
Digitized Records: none
Range: Also known from mainland (Casey, 1908).
Notes. Casey erected the subspecies C. elliptica catalinae Casey, 1918 from Santa Catalina Island, which was later synonymized by Doyen (1977).
Coniontis lamentabilis Blaisdell, 1924
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Cockerell, 1940: 284)
Digitized Records: none
Range: Also known from mainland (Blaisdell, 1924b).
Notes. Cockerell (1940: 284) listed this species from Santa Catalina based on the authority of Fall, who apparently concluded that the previous literature records of C. subpubescens belonged to this species. Without a synthetic revision or subsequent work explaining these conclusions, we have kept the records of these species distinct from each other.
Coniontis lata LeConte, 1866
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Anacapa (Blaisdell, 1921: 211; Miller, 1985a: 20), San Clemente (LeConte, 1866a: 113; Horn, 1870: 298; Casey, 1890: 377; Fall, 1897: 238; Fall, 1901: 165; Casey, 1908: 78; Casey, 1908: 80; Cockerell, 1940: 283; Blaisdell, 1921: 211; Doyen, 1974: 87; Doyen, 1977: 3; Miller, 1985a: 20), San Miguel (Blaisdell, 1921: 211; Cockerell, 1940: 283; Miller, 1985a: 20), San Nicolas (Blaisdell, 1921: 211), Santa Barbara (Fall, 1897: 238; Fall, 1901: 165; Blaisdell, 1921: 211; Miller, 1985a: 20; Miller & Miller, 1985: 128), Santa Cruz (Casey, 1890: 377; Fall, 1897: 238; Fall, 1901: 165; Casey, 1908: 79; Fall & Davis, 1934: 144; Blaisdell, 1921: 211; Doyen, 1977: 3; Miller, 1985a: 20), Santa Rosa (Fall, 1897: 238; Fall, 1901: 165; Blaisdell, 1921: 211; Miller, 1985a: 20)
Digitized Records: Anacapa (15 ASUHIC; 30 CASC; 5 LACM; 2 SBMNH), San Clemente (27 CASC; 2 LACM; 1 MAJC), San Miguel (7 CASC; 34 LACM), San Nicolas (6 CASC; 2 LACM; 1 SBMNH), Santa Barbara (1 OSUC; 14 CASC; 50 LACM; 6 SBMNH), Santa Cruz (8 OSUC; 17 CASC; 3 LACM)
Range: Endemic (LeConte, 1866a; Fall, 1897; Casey, 1908; Doyen, 1977).
Notes. Originally described from San Clemente (LeConte, 1866a) and subsequently found on Santa Cruz and described as a subspecies, this taxon was given its own genus, Coniontides Casey, 1908 by Casey (1908), who recognized four species which were subsequently subsumed back under the present species: Blaisdell (1921, who reported this taxon simply as “Coniontides” across all the islands) synonymized Coniontides clementinus Casey, 1908 (described from San Clemente); Doyen (1977) synonymized Coniontides finitimus Casey, 1908 (uncertain type locality, likely Santa Rosa [Casey, 1908: 80]) and Coniontis lata var. insularis Casey, 1890 (described from Santa Cruz); Coniontides was synonymized with Coniontis by Doyen (1972). Fall (1901) primarily recognized Coniontis lata but used C. lata var insularis for the Santa Cruz population. Fall & Davis (1934) used Coniontides insularis for the Santa Cruz and Santa Rosa populations. Cockerell (1940) variously used the names Coniontides clementinus and Coniontides.
Coniontis microsticta Casey, 1908
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Santa Cruz (5 CASC)
Range: Also known from mainland (Casey, 1908).
Notes. The type and only given locality of this species is Alameda County, California (Casey, 1908), which makes these determinations dubious but impossible to discount until further revisionary works are undertaken.
Coniontis nemoralis Eschscholtz, 1829
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Santa Cruz (5 CASC)
Range: Also known from mainland (Casey, 1908; Bousquet et al., 2018)
Notes. This species was described and reported from the vicinity of San Francisco and is currently separated into two putative subspecies from California and Oregon (Bousquet et al., 2018). The determinations of these specimens are dubious but impossible to discount until further revisionary works are undertaken.
Coniontis santarosae Blaisdell, 1921
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Miguel (Blaisdell, 1921: 210; Cockerell, 1940: 284; Miller, 1985a: 20), Santa Cruz (Cockerell, 1940: 284), Santa Rosa (Blaisdell, 1921: 210; Cockerell, 1940: 284; Miller, 1985a: 20)
Digitized Records: San Miguel (2 CASC), Santa Cruz (3 CASC), Santa Rosa (75 CASC)
Range: Endemic (Blaisdell, 1921).
Notes. Blaisdell (1921) reported that this species clearly belonged to a different species group than C. lata as defined by Casey (1908). Though these two taxa are likely distinct from each other and are both putative island endemics, their status in relation to the mainland species remains to be critically examined.
Coniontis subpubescens LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Fall, 1897: 238), Santa Cruz (LeConte, 1876: 299; Fall, 1897: 238; Fall & Davis, 1934: 144)
Digitized Records: none
Range: Also known from mainland (Bousquet et al., 2018).
Notes. Cockerell (1940: 284) stated on the authority of Fall that the previous reports of this species should in fact refer to C. lamentabilis; this identification correction has not been seen in any other published works or museum records.
Coniontis viatica Eschscholtz, 1829
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Cruz (LeConte, 1876: 299; Fall, 1897: 238; Fall & Davis, 1934: 144)
Digitized Records: none
Range: Also known from mainland.
Eusattus LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus was thoroughly revised by Doyen (1984) who provided reliable keys and distribution information. Ten species are reported from California (Bousquet et al., 2018).
Eusattus difficilis LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Clemente (Doyen, 1984: 97).
Digitized Records: none
Range: Also known from mainland (Doyen, 1984).
Notes. This island record is taken from a distinct marker on a range map, though no further specimen data are given for this or any other distribution points on the map within the main text. This species is broadly distributed on coastal and mainland southern California.
Eusattus politus Horn, 1883
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Miguel (Blaisdell, 1921: 215; Cockerell, 1940: 283; Doyen, 1984: 93; Miller, 1985a: 21), Santa Cruz (Doyen, 1984: 93), Santa Rosa (Fall, 1897: 238; Fall, 1901: 166; Blaisdell, 1921: 215; Doyen, 1984: 93; Miller, 1985a: 21)
Digitized Records: San Miguel (6 LACM; 20 SBMNH), Santa Rosa (1 LACM; 5 SBMNH)
Range: Endemic (Doyen, 1984).
Notes. This species was described from “Santa Barbara California” (Horn, 1883: 304) but no other specimens have ever been reported, definitively or putatively, from the mainland (Doyen, 1984). Eusattus vanduzeei Blaisdell, 1921 (type locality: Prince Island off San Miguel Island) was synonymized by Doyen (1984). Two subspecies were recognized in the last revision by Doyen (1984) which can be separated by the size of the punctures on the head and pronotal disc. Eusattus politus politus Horn, 1883 is known from San Miguel, Prince, and Santa Rosa islands, while E. politus cruzensis Doyen, 1984 is known from Santa Cruz Island. The record from Santa Barbara Island by Fall (1901: 166) is problematic and here discounted. It likely resulted from Horn’s original type locality, but the species was not reported from Santa Barbara Island by either Doyen (1984) or Miller & Miller (1985). The populations from San Miguel and Santa Rosa islands were reported as E. vanduzeei by Blaisdell (1921) and Cockerell (1940).
Eusattus robustus LeConte, 1866
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Clemente (LeConte, 1866a: 112; Horn, 1870: 293; Fall, 1897: 238; Fall, 1901: 166; Casey, 1908: 59; Doyen, 1974: 87; Tschinkel & Doyen, 1976: 331; Doyen, 1977: 6; Doyen, 1984: 95; Miller, 1985a: 21), San Nicolas (Doyen, 1984: 95)
Digitized Records: San Clemente (1 BYUC; 1 OSUC; 36 LACM; 9 MAJC; 14 SBMNH; 2 SDNHM; 2 iNat), San Miguel (1 LACM), San Nicolas (13 LACM; 1 SBMNH), Santa Barbara (1 LACM), Santa Rosa (1 LACM)
Range: Endemic (LeConte, 1866a; Fall, 1897; Doyen, 1977).
Notes. Casey (1908) considered this species to form a distinct genus, Nesostes Casey, 1908, and recognized a subspecies, E. robustus postremus Casey, 1908, from a single specimen also from San Clemente Island. Triplehorn (1968) synonymized Nesostes under Eusattus, and Doyen (1977) synonymized the subspecies. Doyen (1984: 95) commented on what he believed were mislabeled specimens in the LACM from Santa Barbara, San Miguel, Anacapa, and Santa Rosa islands and reasoned that because the species was abundant on the other islands then singleton records must be faulty data. We include specimen records from LACM that likely overlap with those discounted by Doyen above. The specimens were collected by different people at different times so would all have to represent unique mislabeling events. This could be the case but we prefer to not discount these records at this time since we see no evidence to support them being erroneous other than the assumption that the species should be common throughout its range and across time.
Edrotini
Notes. Fifteen genera and 78 species of Edrotini are known to occur in California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data). This tribe experienced a manyfold increase in described species from Casey (1907), and groups that have not been revised subsequently have proved intractable for reliable identifications since. Crypadius was revised by Thomas (1985). Metoponium and Hylocrinus are in need of a revision consisting primarily of synonymy (M. A. Johnston, K. Kanda, R. L. Aalbu, C. C. Wirth, 2023, unpublished data).
Cryptadius LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus inhabits coastal dunes and is distributed from the Channel Islands region south along the Baja California peninsula and coasts of the Gulf of California. The genus was revised by Thomas (1985), who provided distributions and a reliable key to species. One species is known from California (Bousquet et al., 2018).
Cryptadius inflatus LeConte, 1852
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Cruz (Straughan & Hadley, 1980: 392; Thomas, 1985: 197)
Digitized Records: Santa Cruz (1 MAJC; 14 SBMNH), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Thomas, 1985).
Notes. All Cryptadius known from the United States, including the Channel Islands, belong to the nominate subspecies, C. inflatus inflatus LeConte, 1852.
Hylocrinus Casey, 1907
Nomenclatural Authority: Bousquet et al. (2018), M. A. Johnston, K. Kanda, R. L. Aalbu, C. C. Wirth, 2023, unpublished data
Notes. This genus is in great need of revision and seemingly many synonymies for the United States fauna, which was last treated by Casey (1907). Seven putative species are known from California (Bousquet et al., 2018)
Hylocrinus longulus (LeConte, 1851)
Nomenclatural Authority: Bousquet et al. (2018), M. A. Johnston, K. Kanda, R. L. Aalbu, C. C. Wirth, 2023, unpublished data
Literature Records: none
Digitized Records: San Nicolas (1 SBMNH)
Range: Also known from mainland (Casey, 1907; M. A. Johnston, 2022, personal data).
Notes. This is one of two species recognized by M. A. Johnston, K. Kanda, R. L. Aalbu, C. C. Wirth, 2023, unpublished data that are fairly widespread throughout southern California. The single known specimen from San Nicolas is from a reliable collecting event and shows no observable difference from the mainland population.
Metoponium Casey, 1907
Nomenclatural Authority: Bousquet et al. (2018)
Digitized Records (genus-only): San Clemente (1 LACM; 2 SBMNH), Santa Catalina (10 LACM; 23 SBMNH)
Notes. This genus, like the others in this tribe, requires extensive revision. It is highly probable that it only represents a small handful of valid species and is likely itself a synonym of the genus Eurymetopon Eschscholtz, 1829 (M. A. Johnston, 2022, personal observation). Literature and digitized records to the level of species are dubious until a proper revision can be undertaken. Twenty nominal species are currently known from California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data)
Metoponium convexicolle (LeConte, 1851)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Fall, 1897: 238; Baker, 1905: 59)
Digitized Records: none
Range: Also known from mainland (LeConte, 1851; Bousquet et al., 2018).
Notes. The literature records for this species were under the name Eurymetopon convexicolle, its original combination.
Metoponium insulare Casey, 1908
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Casey, 1907: 308; Miller, 1985a: 21)
Digitized Records: Santa Catalina (1 USNM)
Range: Endemic (Casey, 1907).
Notes. The validity of this species is dubious (M. A. Johnston, 2022, personal observation).
Telabis Casey, 1890
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus is in great need of revision and many synonymies (M. A. Johnston, 2022, personal observation). Six currently valid species are recorded from California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data).
Telabis serratus (LeConte, 1866)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Santa Catalina (1 LACM)
Range: Also known from mainland (Horn, 1870).
Notes. This species is widespread across southern California and is common in sandy habitats.
Nyctoporini
Notes. One genus and five species of Nyctoporini have been recorded from California (Bousquet et al., 2018).
Nyctoporis Eschscholtz, 1829
Nomenclatural Authority: Bousquet et al. (2018)
Digitized Records (genus-only): Anacapa (6 LACM), San Miguel (3 LACM), Santa Catalina (5 LACM), Santa Cruz (5 LACM)
Notes. The catalog of Bousquet et al. (2018) included a number of synonymies for this genus which at present comprises four valid species, but there is no key or modern treatment to identify these taxa. This genus is restricted to California, with all five of its species known from the state (Bousquet et al., 2018).
Nyctoporis carinata LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Miguel (Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Catalina (Fall, 1897: 238; Baker, 1905: 59; Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Cruz (Fall & Davis, 1934: 144; Polihronakis & Caterino, 2010a: 426; Caterino, Chatzimanolis & Richmond, 2015: 278), Santa Rosa (Caterino, Chatzimanolis & Richmond, 2015: 278)
Digitized Records: Anacapa (2 SBMNH), San Miguel (1 LACM; 7 SBMNH), Santa Catalina (1 BYUC; 13 SBMNH; 1 iNat), Santa Cruz (8 LACM; 12 SBMNH; 5 TAMU), Santa Rosa (16 SBMNH)
Range: Also known from mainland (Caterino, Chatzimanolis & Richmond, 2015).
Notes. Polihronakis & Caterino (2010a) showed that this species has a high degree of genetic distance in mitochondrial loci between mainland populations, which seem to be the same species as the populations on the islands. Caterino, Chatzimanolis & Richmond (2015) provided a detailed molecular phylogeographic analysis of this and several other beetle species for the California Channel Islands. It seems clear that the island populations and those of southern mainland California are the same species, but the species boundaries become more uncertain in the northern half of the state (M. A. Johnston, 2022, personal observation).
Stenochiinae
Notes. Four genera and 15 species of Stenochiinae, all belonging to the tribe Cnodalonini, have been recorded from California (Bousquet et al., 2018).
Cibdelis Mannerheim, 1843
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus is only known from California and needs a modern revision. All five currently valid species are recorded from California (Bousquet et al., 2018).
Cibdelis bachei LeConte, 1861
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Clemente (Horn, 1870: 341; Fall, 1901: 170), Santa Barbara (LeConte, 1861: 353; Fall, 1897: 238; Fall, 1901: 170; Miller, 1985a: 20; Miller & Miller, 1985: 128), Santa Catalina (Fall, 1901: 170; Miller, 1985a: 20), Santa Cruz (Miller, 1985a: 20)
Digitized Records: Santa Catalina (9 LACM; 2 MAJC; 5 SBMNH), Santa Cruz (4 MAJC; 18 SBMNH), Santa Rosa (8 SBMNH)
Range: Endemic (LeConte, 1861).
Notes. Miller & Miller (1985) suggested that the published type locality of Santa Barbara Island may be due to mislabeling and that the type might actually be from Santa Cruz Island. No other records are known from Santa Barbara Island, so this seems plausible.
Coelocnemis Mannerheim, 1843
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus was revised by Doyen (1973), who provided excellent keys and species treatments. Specimens of this genus are often misidentified or placed within unsorted Eleodes specimens in collections. Three specimens from Santa Cruz Island in the BYUC are determined as Coelocnemis dilaticollis Mannerheim, 1843 (= Coelocnemis californica Mannerheim, 1843) which are deemed likely misidentified or mis-georeferenced, but they may represent a new island record for this genus. Another unverified specimen from Santa Catalina Island determined by H.C. Fall as C. dilaticollis exists in the H.C. Fall collection at the Museum of Comparative Zoology, Harvard (S. Miller, 2022, personal communication). Six species are recorded from California (Bousquet et al., 2018).
Coelocnemis magna LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Doyen, 1973: 90)
Digitized Records: Santa Catalina (9 LACM; 1 SBMNH)
Range: Also known from mainland (Doyen, 1973).
Notes. This species is relatively abundant in coastal habitats of southern California and can most frequently be found under bark and on dead logs.
Tenebrioninae
Notes. Thirteen tribes, 25 genera, and 72 species of Tenebrioninae are known to occur in California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data).
Apocryphini
Notes. One genus and three species of Apocryphini have been recorded from California (Bousquet et al., 2018).
Apocrypha Eschscholtz, 1831
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus putatively has species in both North America and South America. There are three species in North America, all of which are recorded from and restricted to California (Doyen & Kitayama, 1980).
Apocrypha anthicoides Eschscholtz, 1831
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Anacapa (5 SBMNH), Santa Cruz (3 SBMNH)
Range: Also known from mainland (Doyen & Kitayama, 1980).
Notes. This species is widespread along coastal California, but was not recorded from the islands in the last review of the genus (Doyen & Kitayama, 1980).
Eulabini
Notes. Three genera and eight species of Eulabini have been recorded from California (Bousquet et al., 2018).
Apsena LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus was revised by Blaisdell (1932), who provided keys and species treatments. Determinations made since then seem fairly dubious and there is likely some synonymy that needs to be made within this group (M. A. Johnston, 2022, personal observation). Six species are reported from California (Bousquet et al., 2018).
Apsena barbarae Blaisdell, 1932
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Blaisdell, 1932: 63), Santa Cruz (Fall & Davis, 1934: 144)
Digitized Records: none
Range: Also known from mainland (Blaisdell, 1932).
Notes. Blaisdell (1932) described this species in the “pubescens-group” with a type locality of Santa Barbara. Subsequently identified material may have been determined as Apsena pubescens (LeConte, 1851), listed below. Berry (1970: 304), in a nomenclaturally unavailable dissertation, suggested that this species should be a junior synonym of A. pubescens.
Apsena grossa (LeConte, 1866)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Anacapa (Berry, 1970: 299), San Clemente (LeConte, 1866a: 118, Horn, 1870: 324; Fall, 1897: 238; Fall, 1901: 168; Blaisdell, 1932: 75; Berry, 1970: 299; Doyen, 1974: 87; Miller, 1985a: 20), San Nicolas (Fall, 1901: 169; Blaisdell, 1932: 75; Berry, 1970: 299; Miller, 1985a: 20), Santa Barbara (Fall, 1901: 169; Blaisdell, 1932: 75; Berry, 1970: 299; Miller, 1985a: 20; Miller & Miller, 1985: 128), Santa Catalina (Blaisdell, 1932: 75; Berry, 1970: 299; Miller, 1985a: 20), Santa Rosa (Berry, 1970: 299)
Digitized Records: Anacapa (8 LACM), San Clemente (74 LACM; 2 MAJC; 15 SBMNH; 13 SDNHM), San Nicolas (103 LACM; 16 SBMNH), Santa Barbara (130 LACM; 2 MAJC; 17 SBMNH), Santa Catalina (13 LACM), Santa Rosa (4 LACM)
Range: Endemic (LeConte, 1866a; Fall, 1897; Blaisdell, 1932; Berry, 1970).
Notes. This is the largest species of the genus and is immediately recognizable by its rotund form. Blaisdell (1932) included this species in the “pubescens-group”. It has only been recorded from the Channel Islands and was listed as its original combination Eulabis grossa by LeConte (1866a), Horn (1870), and Fall (1897, 1901).
Apsena pubescens (LeConte, 1851)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Santa Catalina (Fall, 1897: 238; Baker, 1905: 57; Blaisdell, 1932: 55; Berry, 1970: 308)
Digitized Records: San Clemente (1 LACM), Santa Catalina (1 LACM; 13 SBMNH), Santa Cruz (3 LACM; 14 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Blaisdell, 1932; Berry, 1970).
Notes. Blaisdell (1932) largely separated this species, namesake of the “pubescens-group” and type of the genus, from A. barbarae by means of more attenuate males and difference in setal length. All island records of these two taxa will need to be critically examined as part of a revision of this genus.
Apsena rufipes (Eschscholtz, 1829)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Santa Cruz (1 CSUC; 1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Blaisdell, 1932; Berry, 1970).
Notes. Blaisdell (1932) placed this species in its own species group, the “rufipes-group”, but provided no island records. The single specimen from CSUC is determined as “Eulabis rufipes”.
Epantius LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This is a monotypic genus with its single species known from California (Bousquet et al., 2018).
Epantius obscurus LeConte, 1851
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: Anacapa (Blaisdell, 1943: 238; Berry, 1970: 294), San Nicolas (Blaisdell, 1932: 94; Blaisdell, 1943: 238), Santa Catalina (Berry, 1970: 294), Santa Cruz (Blaisdell, 1932: 94; Blaisdell, 1943: 238; Berry, 1970: 294), Santa Rosa (Fall, 1897: 238; Blaisdell, 1943: 238)
Digitized Records: Anacapa (41 LACM), San Clemente (2 LACM; 4 SBMNH), San Miguel (2 LACM; 46 SBMNH; 1 SDNHM), San Nicolas (95 LACM; 7 SBMNH), Santa Catalina (4 SBMNH), Santa Cruz (98 LACM; 33 SBMNH), Santa Rosa (1 LACM; 5 SBMNH)
Range: Also known from mainland (Blaisdell, 1932).
Notes. This species is common along the California shoreline and can be found under wrack and dune vegetation near and above the high-tide line. Fall (1897) recorded this species as Eulabis obscura.
Helopini
Notes. Two genera and 25 species of Helopini are known to occur in California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data).
Helops Fabricius, 1775
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records (genus-only): Santa Barbara (Miller & Miller, 1985: 128), Santa Catalina (Fall, 1897: 238)
Digitized Records (genus-only): San Clemente (6 SDNHM)
Notes. This genus, as currently circumscribed, is worldwide and the New World species are in great need of revision, all likely not being congeneric with the European type species. The Helops fauna of the western United States is particularly in need of revision, and both literature and digitized records should be reviewed. The most reliable key is that of Horn (1870). Miller & Miller (1985) referred to a species of Helops other than H. bachei LeConte, 1861 from Santa Barbara Island on the authority of T.J. Spilman. There are 23 species known to occur in California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data).
Helops bachei LeConte, 1861
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Clemente (Doyen, 1974: 87), Santa Barbara (LeConte, 1861: 353; Horn, 1870: 396; Fall, 1897: 238; Fall, 1901: 175; Miller & Miller, 1985: 128)
Digitized Records: Anacapa (1 LACM), San Clemente (9 LACM; 1 SBMNH), San Miguel (5 SBMNH), San Nicolas (13 LACM; 9 SBMNH), Santa Barbara (3 LACM), Santa Catalina (3 LACM; 15 SBMNH), Santa Cruz (1 CSUC; 1 MAJC; 3 SBMNH), Santa Rosa (1 LACM; 20 SBMNH)
Range: Endemic (LeConte, 1861; Fall, 1901); also known from mainland (Horn, 1870; Fall, 1901).
Notes. Fall (1901: 175) reported this species from the mainland but perhaps the “true form” is only found on the “Santa Barbara Islands”. This species is very similar to the mainland species Helops rugicollis LeConte, 1866, but most noticeably differs by having rounded tubercles on the elytra. This complex and all the island records need to be closely examined. Whether this is an island endemic species is hard to know and is debatable according to the literature.
Helops blaisdelli Casey, 1891
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: San Nicolas (Cockerell, 1940: 284)
Digitized Records: none
Range: Also known from mainland (Casey, 1891).
Notes. This identification by Cockerell (1940: 284) is somewhat dubious. Casey (1891) described this species from San Diego and likened it to H. bachei. It is possible this taxon corresponds to the mainland populations of the latter species reported by Horn (1870), or it could be a valid species and, the Cockerell (1940) record notwithstanding, validates H. bachei as a true island endemic with a mainland sister species. Until this genus is revised, this record cannot be fully discounted for the Channel Islands.
Helops rugicollis LeConte, 1866
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Santa Catalina (2 SBMNH)
Range: Also known from mainland (Horn, 1870).
Notes. See discussion on two species above; this taxon is dubious for the islands but is in an unrevised species complex with the other two species recorded from the Channel Islands. The true identity and number of species on the Channel Islands requires revision.
Triboliini
Notes. Five genera and 13 species of Triboliini are known to occur in California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data).
Tribolium MacLeay, 1825
Nomenclatural Authority: Bousquet et al. (2018)
Notes. This genus contains several cosmopolitan species that are strongly synanthropic. Six species are recorded from California (Bousquet et al., 2018; M. L. Gimmel, 2022, unpublished data)
Tribolium castaneum (Herbst, 1797)
Nomenclatural Authority: Bousquet et al. (2018)
Literature Records: none
Digitized Records: Santa Catalina (1 BYUC)
Range: Also known from mainland (Bousquet et al., 2018).
Notes. This species, commonly referred to as the red flour beetle, is synanthropic and found throughout California wherever people live.
Zopheridae
Notes. Two subfamilies, 18 genera, and 42 species of Zopheridae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Colydiinae
Notes. Three tribes, 12 genera, and 28 species of Colydiinae have been recorded from California (M. L. Gimmel, 2022, unpublished data). The North American species of the subfamily were treated by Stephan (1989); Ivie et al. (2016) provided an overview and key to the New World genera, as well as a checklist of New World species.
Rhagoderini
Notes. One genus and three species of Rhagoderini have been recorded from California (Stephan, 1989; Krinsky, 2015).
Rhagodera Mannerheim, 1843
Nomenclatural Authority: Stephan (1989)
Notes. Three species of Rhagodera have been recorded from California (Stephan, 1989; Krinsky, 2015).
Rhagodera costaefragmenta Krinsky, 2015
Nomenclatural Authority: Ivie et al. (2016)
Literature Records: San Clemente (Krinsky, 2015: 294)
Digitized Records: none
Range: Endemic (Krinsky, 2015).
Notes. This species was described from five specimens collected on San Clemente Island which are reportedly deposited in the YPMC (Krinsky, 2015).
Rhagodera interrupta Stephan, 1989
Nomenclatural Authority: Ivie et al. (2016)
Literature Records: San Nicolas (Krinsky, 2015: 293)
Digitized Records: San Nicolas (6 SBMNH)
Range: Unknown.
Notes. This species was described from five specimens located in the Ulke collection at the Carnegie Museum of Natural History which were only labeled as “California” (Stephan, 1989). No other localities have been recorded in print or in digitized specimen data besides the island record given here.
Rhagodera tuberculata (Mannerheim, 1843)
Nomenclatural Authority: Ivie et al. (2016)
Literature Records: San Clemente (Miller & Miller, 1985: 127), Santa Barbara (Miller & Miller, 1985: 127; Krinsky, 2015: 293), Santa Cruz (Krinsky, 2015: 293)
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Stephan, 1989).
Notes. This is one of the more widespread and commonly collected species of this genus, with most of its known specimens originating from Los Angeles County, California (Stephan, 1989).
Synchitini
Notes. Nine genera and 22 species of Synchitini have been recorded from California (Stephan, 1989; M. L. Gimmel, 2022, unpublished data).
Lasconotus Erichson, 1845
Nomenclatural Authority: Stephan (1989)
Notes. Twelve species of Lasconotus have been recorded from California (Stephan, 1989).
Lasconotus linearis Crotch, 1874
Nomenclatural Authority: Ivie et al. (2016)
Literature Records: none
Digitized Records: Santa Cruz (3 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Crotch, 1874).
Megataphrus Casey, 1890
Nomenclatural Authority: Stephan (1989)
Notes. One species of Megataphrus has been recorded from California (Stephan, 1989).
Megataphrus tenuicornis Casey, 1890
Nomenclatural Authority: Ivie et al. (2016)
Literature Records: none
Digitized Records: Santa Rosa (16 SBMNH)
Range: Also known from mainland (Stephan, 1989).
Synchita Hellwig, 1792
Nomenclatural Authority: Stephan (1989)
Notes. One species of Synchita has been recorded from California (Stephan, 1989).
Synchita lecontei Ivie et al., 2016
Nomenclatural Authority: Ivie et al. (2016)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Stephan, 1989).
Notes. This species was known as Microsicus variegatus (LeConte, 1858) in Stephan (1989).
Zopherinae: Zopherini
Notes. Four tribes, six genera, and 14 species of Zopherinae, of which two genera and seven species belong to Zopherini, have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Phloeodes LeConte, 1862
Nomenclatural Authority: Foley & Ivie (2008)
Notes. Two species of Phloeodes have been recorded from California (Foley & Ivie, 2008). The species of this genus were revised by Foley & Ivie (2008).
Phloeodes diabolicus (LeConte, 1851)
Nomenclatural Authority: Foley & Ivie (2008)
Literature Records: Santa Cruz (LeConte, 1876: 299; Fall, 1897: 238; Fall & Davis, 1934: 144)
Digitized Records: none
Range: Also known from mainland (García-París, Coca-Abia & Parra-Olea, 2006; Foley & Ivie, 2008; Polihronakis & Caterino, 2010b).
Notes. No island records were given by García-París, Coca-Abia & Parra-Olea (2006), Foley & Ivie (2008), or Polihronakis & Caterino (2010b). The literature records of this species are curious. This is a large and charismatic species which is readily separable from its congener Phloeodes plicatus (LeConte, 1859). We presume that LeConte (1876) and those that followed merely recorded the wrong species name, but these authors were familiar with these two species so it may yet prove to be a valid record.
Phloeodes plicatus (LeConte, 1859)
Nomenclatural Authority: Foley & Ivie (2008)
Literature Records: Santa Catalina (Polihronakis & Caterino, 2010b: 3), Santa Cruz (García-París, Coca-Abia & Parra-Olea, 2006: 228; Foley & Ivie, 2008: 46)
Digitized Records: Santa Catalina (5 SBMNH), Santa Cruz (2 OSUC; 7 SBMNH)
Range: Also known from mainland (García-París, Coca-Abia & Parra-Olea, 2006; Foley & Ivie, 2008; Polihronakis & Caterino, 2010b).
Notes. We have examined specimens of this species and are confident of its occurrence on the islands, but see comments under P. diabolicus.
COCCINELLOIDEA
Akalyptoischiidae
Notes. One genus and 17 species of Akalyptoischiidae have been recorded from California (Hartley, Andrews & McHugh, 2008).
Akalyptoischion Andrews, 1976
Nomenclatural Authority: Hartley, Andrews & McHugh (2008)
Notes. Seventeen species of Akalyptoischion have been recorded from California (Hartley, Andrews & McHugh, 2008). This genus was revised by Hartley, Andrews & McHugh (2008).
Akalyptoischion heterotrichos Hartley, Andrews & McHugh, 2008
Nomenclatural Authority: Hartley, Andrews & McHugh (2008)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (Hartley, Andrews & McHugh, 2008).
Akalyptoischion hormathos Andrews, 1976
Nomenclatural Authority: Hartley, Andrews & McHugh (2008)
Literature Records: Santa Barbara (Andrews, 1976a: 9; Miller & Miller, 1985: 127; Hartley, Andrews & McHugh, 2008: 37)
Digitized Records: San Clemente (19 SBMNH), Santa Catalina (14 SBMNH), Santa Cruz (15 SBMNH), Santa Rosa (30 SBMNH)
Range: Also known from mainland (Andrews, 1976a; Hartley, Andrews & McHugh, 2008).
Cerylonidae, NEW FAMILY RECORD
Notes. Three genera and four species of Cerylonidae have been recorded from California (Lawrence & Stephan, 1975).
Cerylon Latreille, 1802
Nomenclatural Authority: (Lawrence & Stephan, 1975).
Notes. The genus Cerylon contains two species recorded from California (Lawrence & Stephan, 1975). These were keyed by Lawrence & Stephan (1975).
Cerylon unicolor (Ziegler, 1845)
Nomenclatural Authority: Lawrence & Stephan (1975)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Lawrence & Stephan, 1975).
Coccinellidae
Notes. The known North American species of Coccinellidae were fully treated by Gordon (1985), with additions to the introduced fauna by Gordon & Vandenberg (1991). Two subfamilies, 42 genera, and 175 species are known from California (M. L. Gimmel, 2022, unpublished data).
Coccinellinae
Notes. Eight tribes, 38 genera, and 164 species of Coccinellinae have been recorded from California (Gordon, 1985; M. L. Gimmel, 2022, unpublished data).
Cephaloscymnini
Notes. One genus and species of Cephaloscymnini has been recorded from California (Gordon, 1985).
Cephaloscymnus Crotch, 1873
Nomenclatural Authority: Gordon (1985)
Notes. One species of Cephaloscymnus is recorded from California (Gordon, 1985).
Cephaloscymnus occidentalis Horn, 1895
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Fall, 1897: 237; Fall, 1901: 87)
Digitized Records: none
Range: Also known from mainland (Fall, 1901; Gordon, 1985).
Chilocorini
Notes. Five genera and 17 species of Chilocorini have been recorded from California (Gordon, 1985).
Axion Mulsant, 1850
Nomenclatural Authority: Gordon (1985)
Notes. One species of Axion is recorded from California (Gordon, 1985).
Axion plagiatum (Olivier, 1808)
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Santa Cruz (5 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Chilocorus Leach, 1815
Nomenclatural Authority: Gordon (1985)
Notes. Five species of Chilocorus have been recorded from California, two of which are adventive (Gordon, 1985). These were keyed by Gordon (1985).
Chilocorus undetermined species
Literature Records: Santa Catalina (Seavey, 1892: 263; Fall, 1897: 237)
Digitized Records: Santa Cruz (1 UCSB)
Notes. The records from Seavey (1892) and Fall (1897) were reported as Chilocorus bivulnerus Mulsant, 1850, a current junior synonym of Chilocorus stigma (Say, 1835), a species that does not occur in California. Early California records of C. bivulnerus, therefore, refer to either Chilocorus fraternus LeConte, 1860 or Chilocorus orbus Casey, 1899, two species which can only be reliably distinguished by examination of male genitalia (Gordon, 1985).
Coccidulini
Notes. Nine genera and 51 species of Coccidulini have been recorded from California (Gordon, 1985; M. L. Gimmel, 2022, unpublished data).
Nephus Mulsant, 1846
Nomenclatural Authority: Gordon (1985)
Digitized Records (genus-only): Anacapa (1 LACM), Santa Barbara (1 LACM), Santa Catalina (1 LACM)
Notes. Six species of Nephus in three subgenera (Scymnobius Casey, 1899; Sidis Mulsant, 1850; Turboscymnus Gordon, 1976) have been reported from California (Gordon, 1985).
Nephus (Scymnobius) guttulatus (LeConte, 1852)
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Fall, 1897: 237; Fall, 1901: 86)
Digitized Records: Anacapa (4 SBMNH), Santa Barbara (1 SBMNH), Santa Cruz (3 SBMNH)
Range: Also known from mainland (Fall, 1901; Gordon, 1985).
Notes. Fall (1897, 1901) recorded this species as Scymnus guttulatus.
Nephus (Scymnobius) sordidus (Horn, 1895)
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Anacapa (1 SBMNH), San Nicolas (9 SBMNH), Santa Catalina (2 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Nephus (Sidis) binaevatus (Mulsant, 1850)
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Cockerell, 1940: 286; Gordon, 1985: 293 [map])
Digitized Records: Santa Catalina (1 iNat)
Range: Also known from mainland (Gordon, 1985).
Notes. This species was recorded as Scymnus binaevatus by Cockerell (1940). It was introduced to California from South Africa in 1921 for mealybug control (Gordon, 1985).
Rhyzobius Stephens, 1829
Nomenclatural Authority: Gordon (1985)
Notes. Two introduced species of Rhyzobius are recorded from California (Gordon, 1985).
Rhyzobius forestieri (Mulsant, 1853)
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Pope, 1981: 26), Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Notes. This species was introduced to North America (Gordon, 1985).
Rhyzobius lophanthae (Blaisdell, 1892)
Nomenclatural Authority: Gordon (1985)
Literature Records: San Clemente (Fall, 1897: 237; Cockerell, 1940: 286)
Digitized Records: San Nicolas (1 SBMNH), Santa Cruz (2 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Notes. Fall (1897) reported this species as “Rhizobius lophanthae”, and Cockerell (1940) reported it as “Lindorus lophantae”. This species was introduced to North America (Gordon, 1985).
Scymnus Kugelann, 1794
Nomenclatural Authority: Gordon (1985)
Digitized Records (genus-only): Santa Barbara (1 SBMNH)
Notes. Thirty-six species of Scymnus have been reported from California, all but four belonging to the subgenus Pullus Mulsant, 1846 and the remainder to Scymnus (Scymnus) (Gordon, 1985). These were keyed out by Gordon (1976) and Gordon (1985). The above Santa Barbara Island record is based on a single female specimen of Pullus that keys out to couplet 29 of Gordon’s (1985) key to Pullus of “Region IV”, which relies on male genitalia to separate the species.
Scymnus (Pullus) ardelio Horn, 1895
Nomenclatural Authority: Gordon (1985)
Literature Records: San Clemente (Fall, 1897: 237), Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (Gordon, 1985).
Scymnus (Pullus) cervicalis Mulsant, 1850
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Fall, 1897: 237; Fall, 1901: 86)
Digitized Records: Santa Cruz (3 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Fall, 1901; Gordon, 1985).
Scymnus (Pullus) coniferarum Crotch, 1874
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Scymnus (Pullus) falli Gordon, 1976
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Barbara (Gordon, 1985: 199; Miller, 1985a: 20; Miller & Miller, 1985: 127), Santa Cruz (Gordon, 1976: 140; 1985: 199 [map]; Miller, 1985a: 20; Miller & Miller, 1985: 127; Naughton et al., 2014: 303), Santa Rosa (Miller, 1985a: 20; Miller & Miller, 1985: 127)
Digitized Records: San Miguel (1 SBMNH), Santa Cruz (8 SBMNH), Santa Rosa (3 SBMNH)
Range: Endemic (Gordon, 1976; Gordon, 1985; Miller, 1985a; Miller & Miller, 1985; Naughton et al., 2014).
Scymnus (Pullus) jacobianus Casey, 1899
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: San Clemente (2 SBMNH), San Miguel (2 SBMNH), Santa Barbara (6 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Scymnus (Pullus) loewii Mulsant, 1850
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Cruz (Fall & Davis, 1934: 143; Gordon, 1976: 124)
Digitized Records: San Clemente (2 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Gordon, 1976, 1985).
Notes. Fall & Davis (1934) recorded this species as Scymnus cinctus LeConte, 1852, which is now a junior synonym of S. loewii (see Gordon, 1985).
Scymnus (Pullus) marginicollis Mannerheim, 1843
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Horn, 1895: 105; Fall, 1897: 237), Santa Cruz (Gordon, 1976: 128)
Digitized Records: Anacapa (1 SBMNH), San Nicolas (5 SBMNH), Santa Catalina (4 SBMNH), Santa Cruz (10 LACM; 1 SBMNH), Santa Rosa (2 LACM; 1 SBMNH)
Range: Also known from mainland (Gordon, 1976, 1985).
Scymnus (Pullus) pallens LeConte, 1852
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Cockerell, 1940: 286), Santa Cruz (Gordon, 1976: 86; Naughton et al., 2014: 303)
Digitized Records: Santa Cruz (4 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Gordon, 1976, 1985).
Scymnus (Scymnus) difficilis Casey, 1899
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: San Miguel (10 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Scymnus (Scymnus) fenderi Malkin, 1943
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Santa Rosa (3 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Scymnus (Scymnus) nebulosus LeConte, 1852
Nomenclatural Authority: Gordon (1985)
Literature Records: San Miguel (Miller & Davis, 1986: 550), Santa Catalina (Fall, 1897: 237; Fall, 1901: 86; Cockerell, 1940: 286)
Digitized Records: Santa Catalina (1 OSUC; 5 SBMNH; 1 iNat), Santa Cruz (28 LACM; 10 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Fall, 1901; Gordon, 1985).
Stethorus Weise, 1885
Nomenclatural Authority: Gordon (1985)
Notes. One species of Stethorus has been reported from California (Gordon, 1985).
Stethorus punctum (LeConte, 1852)
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Santa Catalina (3 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Notes. The subspecies of S. punctum occurring in California is S. p. picipes Casey, 1899 (Gordon, 1985).
Zagloba Casey, 1899
Nomenclatural Authority: Gordon (1985)
Notes. One species of Zagloba has been reported from California (Gordon, 1985).
Zagloba ornata (Horn, 1895)
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Horn, 1895: 112; Fall, 1897: 237; Fall, 1901: 87)
Digitized Records: San Miguel (2 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Fall, 1901; Gordon, 1985).
Notes. Horn (1895) and Fall (1897, 1901) recorded this species as Cephaloscymnus ornatus.
Coccinellini
Notes. Sixteen genera and 41 species of Coccinellini have been recorded from California (Gordon, 1985; M. L. Gimmel, 2022, unpublished data).
Coccinella Linnaeus, 1758
Nomenclatural Authority: Gordon (1985)
Digitized Records (genus-only): San Clemente (4 LACM), San Miguel (12 LACM), San Nicolas (29 LACM), Santa Catalina (1 LACM), Santa Cruz (2 LACM)
Notes. Eleven species of Coccinella have been recorded from California (Gordon, 1985; Gordon & Vandenberg, 1991). This genus was revised for North America by Brown (1962), but an updated key was provided by Gordon (1985). Fall (1901: 84) reported the taxon “Coccinella transversoguttata var. transversalis” from “the islands”. Dobzhansky (1931: 16), however, noted that “This form [C. transversoguttata] is apparently lacking in southern California”. We did not attempt to include this record below.
Coccinella californica Mannerheim, 1843
Nomenclatural Authority: Gordon (1985)
Literature Records: Anacapa (Miller & Miller, 1985: 126), San Clemente (Fall, 1897: 237; Dobzhansky, 1931: 13; Miller & Miller, 1985: 126), San Nicolas (Fall, 1897: 237; Dobzhansky, 1931: 13; Miller & Miller, 1985: 126), Santa Barbara (Miller & Miller, 1985: 126), Santa Catalina (Fall, 1897: 237; Miller & Miller, 1985: 126), Santa Cruz (Dobzhansky, 1931: 13; Miller & Miller, 1985: 126), Santa Rosa (Fall, 1897: 237; Dobzhansky, 1931: 13; Miller & Miller, 1985: 126)
Digitized Records: Anacapa (3 LACM; 7 SBMNH), San Clemente (2 LACM; 1 SBMNH; 25 iNat), San Miguel (2 LACM; 1 SBMNH), San Nicolas (11 LACM; 2 SBMNH; 9 iNat), Santa Barbara (18 LACM; 1 SBMNH), Santa Catalina (8 LACM; 8 iNat), Santa Cruz (2 LACM; 7 SBMNH; 3 UCSB; 4 iNat), Santa Rosa (3 LACM; 14 SBMNH; 5 iNat)
Range: Also known from mainland (Gordon, 1985).
Coccinella johnsoni Casey, 1908
Nomenclatural Authority: Gordon (1985)
Literature Records: San Clemente (Dobzhansky, 1931: 14; Brown, 1962: 794; Miller & Miller, 1985: 127), San Nicolas (Dobzhansky, 1931: 14; Brown, 1962: 794; Miller & Miller, 1985: 127), Santa Barbara (Miller & Miller, 1985: 127)
Digitized Records: San Clemente (8 SBMNH), San Nicolas (10 SBMNH), Santa Barbara (3 LACM)
Range: Also known from mainland (Dobzhansky, 1931; Brown, 1962; Gordon, 1985).
Coccinella novemnotata Herbst, 1793
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: San Miguel (1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Coccinella septempunctata (Linnaeus, 1758)
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Anacapa (1 iNat), San Clemente (2 SBMNH; 13 iNat), San Miguel (2 SBMNH), San Nicolas (1 SBMNH; 3 iNat), Santa Barbara (1 iNat), Santa Catalina (4 SBMNH; 3 iNat), Santa Cruz (4 SBMNH; 7 iNat), Santa Rosa (4 iNat)
Range: Also known from mainland (Gordon, 1985).
Notes. This species was introduced to North America for control of aphids (Gordon, 1985).
Cycloneda Crotch, 1871
Nomenclatural Authority: Gordon (1985)
Digitized Records (genus-only): Santa Cruz (1 iNat)
Notes. Two species of Cycloneda have been recorded from California (Gordon, 1985).
Cycloneda polita Casey, 1899
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Fall, 1897: 237), Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: Santa Cruz (3 LACM; 5 SBMNH; 5 UCSB), Santa Rosa (1 LACM)
Range: Also known from mainland (Gordon, 1985).
Notes. Fall (1897) recorded this species as “Cycloneda oculata Fabricius”.
Cycloneda sanguinea (Linnaeus, 1763)
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Seavey, 1892: 263; Fall, 1897: 237)
Digitized Records: Anacapa (1 SBMNH), Santa Catalina (1 LACM; 5 SBMNH; 2 iNat), Santa Cruz (12 SBMNH; 2 UCSB; 6 iNat)
Range: Also known from mainland (Gordon, 1985).
Notes. Seavey (1892) recorded this species as Coccinella sanguinea. The subspecies occurring in California is the nominate subspecies, C. s. sanguinea (Linnaeus, 1763).
Hippodamia Dejean, 1837
Nomenclatural Authority: Gordon (1985)
Notes. Ten species of Hippodamia have been reported from California (Gordon, 1985).
Hippodamia convergens Guérin-Méneville, 1842
Nomenclatural Authority: Gordon (1985)
Literature Records: Anacapa (Miller & Miller, 1985: 127), San Miguel (Miller & Miller, 1985: 127), Santa Barbara (Miller & Miller, 1985: 127), Santa Catalina (Seavey, 1892: 263; Fall, 1897: 237; Cockerell, 1940: 286), Santa Cruz (Miller & Miller, 1985: 127), Santa Rosa (Miller & Miller, 1985: 127)
Digitized Records: Anacapa (8 SBMNH), San Clemente (1 SBMNH; 6 iNat), San Miguel (1 SBMNH), San Nicolas (1 LACM; 1 SBMNH; 1 iNat), Santa Barbara (1 SBMNH), Santa Catalina (1 SBMNH; 3 iNat), Santa Cruz (27 LACM; 15 SBMNH; 3 UCSB; 2 iNat), Santa Rosa (1 LACM; 5 SBMNH)
Range: Also known from mainland (Chapin, 1946; Gordon, 1985).
Notes. The map in Chapin (1946: plate 21) shows the presence of this species on at least two islands, which were not mentioned. Cockerell (1940) reported this species as “Hippodamia obsoleta LeConte” (= Hippodamia convergens var. obsoleta Crotch, 1873), a current synonym of H. convergens (see Gordon, 1985).
Hippodamia quinquesignata (Kirby, 1837)
Nomenclatural Authority: Gordon (1985)
Literature Records: San Clemente (Fall, 1897: 237), San Nicolas (Cockerell, 1940: 286), Santa Catalina (Seavey, 1892: 263; Fall, 1897: 237), Santa Cruz (Fall & Davis, 1934: 143), Santa Rosa (Fall, 1897: 237)
Digitized Records: Anacapa (5 SBMNH), San Clemente (10 SBMNH; 1 iNat), San Miguel (55 LACM; 16 SBMNH), San Nicolas (12 LACM; 7 SBMNH), Santa Catalina (3 LACM; 4 SBMNH), Santa Cruz (12 LACM; 15 SBMNH; 18 iNat), Santa Rosa (29 LACM; 37 SBMNH; 1 iNat)
Range: Also known from mainland (Gordon, 1985).
Notes. Seavey (1892), Fall (1897), and Fall & Davis (1934) recorded this species as Hippodamia ambigua LeConte, 1852. Cockerell (1940) recorded it as “H. quinquesignata Kirby, variety”. Chapin (1946: 16) recorded this taxon as “Abundant in western California and the Channel Islands” as the subspecies H. q. punctulata LeConte, 1852, now considered a synonym of H. q. ambigua LeConte, 1852, the only subspecies occurring in coastal California (see Gordon, 1985). Chapin’s (1946: plate 20) map shows its presence on multiple islands. Fall (1897: 239) noted the Santa Rosa specimens might well be Hippodamia convergens. Based on the virtual absence of elytral dark maculation (except for scutellar spot), all Hippodamia quinquesignata (Kirby, 1837) represented on the Channel Islands belong to H. q. ambigua. Members of this subspecies with the white convergent lines present on the pronotum can be difficult to distinguish from immaculate members of H. convergens without examination of male genitalia (Gordon, 1985: 727).
Olla Casey, 1899
Nomenclatural Authority: Gordon (1985)
Notes. One species of Olla has been recorded from California (Gordon, 1985).
Olla v-nigrum (Mulsant, 1866)
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Anacapa (1 LACM), Santa Catalina (1 LACM)
Range: Also known from mainland (Gordon, 1985).
Paranaemia Casey, 1899
Nomenclatural Authority: Gordon (1985)
Notes. One species of Paranaemia has been recorded from California (Gordon, 1985).
Paranaemia vittigera (Mannerheim, 1843)
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Cruz (LeConte, 1876: 298; Fall, 1897: 237; Fall & Davis, 1934: 143)
Digitized Records: none
Range: Also known from mainland (Gordon, 1985).
Notes. This species was recorded as Hippodamia vittigera by LeConte (1876) and Fall (1897), and as Ceratomegilla vittigera by Fall & Davis (1934).
Psyllobora Dejean, 1836
Nomenclatural Authority: Gordon (1985)
Digitized Records (genus-only): Santa Cruz (10 EMEC; 17 LACM; 9 UCSB)
Notes. Three species of Psyllobora have been recorded from California (Gordon, 1985).
Psyllobora renifer Casey, 1899
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Santa Cruz (1 UCSB)
Range: Also known from mainland (Gordon, 1985).
Psyllobora vigintimaculata (Say, 1824)
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Seavey, 1892: 263; Fall, 1897: 237), Santa Cruz (Fall & Davis, 1934: 143; Naughton et al., 2014: 303)
Digitized Records: San Miguel (6 SBMNH), San Nicolas (4 SBMNH), Santa Catalina (12 SBMNH; 1 iNat), Santa Cruz (21 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Notes. This species was recorded as Psyllobora taedata LeConte, 1860 by Seavey (1892) and Fall (1897), and as Psyllobora 20-maculata var. taedata by Fall & Davis (1934). Seavey (1892) reported it from Artemisia californica.
Diomini
Notes. One genus and three species of Diomini have been recorded from California (Gordon, 1985).
Diomus Mulsant, 1850
Nomenclatural Authority: Gordon (1985)
Notes. Three species of Diomus have been recorded from California (Gordon, 1985).
Diomus debilis (LeConte, 1852)
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Anacapa (3 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Hyperaspidini
Notes. Four genera and 47 species of Hyperaspidini have been recorded from California (Gordon, 1985).
Hyperaspidius Crotch, 1873
Nomenclatural Authority: Gordon (1985)
Digitized Records (genus-only): San Miguel (1 SBMNH), San Nicolas (1 LACM; 1 SBMNH)
Notes. Eight species of Hyperaspidius have been recorded from California (Gordon, 1985).
Hyperaspidius comparatus Casey, 1899
Nomenclatural Authority: Gordon (1985)
Literature Records: San Miguel (Cockerell, 1940: 286; Gordon, 1985: 361)
Digitized Records: none
Range: Also known from mainland (Gordon, 1985).
Notes. Cockerell’s (1940) San Miguel Island record was referred to Hyperaspidius vittigerus (LeConte, 1852), which was indicated as previously bearing the name Hyperaspidius trimaculatus (Linnaeus, 1767). However, Hyperaspidius vittigerus was not shown to occur west of the Rocky Mountains by Gordon (1985), so Cockerell’s (1940) record probably refers to H. comparatus.
Hyperaspis Redtenbacher, 1844
Nomenclatural Authority: Gordon (1985)
Literature Records (genus-only): Santa Cruz (Naughton et al., 2014: 303)
Digitized Records (genus-only): San Clemente (1 SBMNH), Santa Rosa (4 SBMNH)
Notes. Thirty-six species of Hyperaspis have been recorded from California (Gordon, 1985). The Naughton et al. (2014) record above is based on a record of “Hyperaspis sp.” in addition to the three taxa listed below.
Hyperaspis lateralis Mulsant, 1850
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Catalina (Seavey, 1892: 262; Fall, 1897: 237), Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: Santa Catalina (1 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Notes. This species was reported from Artemisia californica by Seavey (1892).
Hyperaspis species near annexa LeConte, 1852
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: Santa Cruz (3 SBMNH)
Range: Unknown.
Notes. Naughton et al. (2014) recorded this species as “Hyperaspis nr. annexa”. MLG observed two of these vouchers and one additional specimen from SBMNH, and they do appear quite similar to mainland H. annexa, but with much less yellow and more extensive black coloration. These specimens require more detailed study.
Hyperaspis taeniata LeConte, 1852
Nomenclatural Authority: Gordon (1985)
Literature Records: Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: Santa Cruz (3 LACM; 1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Microweiseinae
Notes. Three tribes, five genera, and 11 species of Microweiseinae are known to occur in California (M. L. Gimmel, 2022, unpublished data). Escalona & Ślipiński (2011) provided a generic revision and reclassification of this subfamily.
Carinodulini
Notes. One genus and species of Carinodulini is known to occur in California (Escalona & Ślipiński, 2011).
Carinodulinka Ślipiński & Tomaszewska, 2002
Nomenclatural Authority: Escalona & Ślipiński (2011)
Notes. No described species of Carinodulinka have yet been recorded from California (see below).
Carinodulinka undescribed species near baja Ślipiński & Tomaszewska, 2002
Nomenclatural Authority: Escalona & Ślipiński (2011)
Literature Records: none
Digitized Records: San Clemente (1 SBMNH)
Range: Also known from mainland (Escalona & Ślipiński, 2011).
Notes. According to Escalona & Ślipiński (2011: 13), California specimens of the genus Carinodulinka Ślipiński & Tomaszewska, 2002 are an unnamed species.
Microweiseini
Notes. Three genera and eight species of Microweiseini are known to occur in California (Gordon, 1985; M. L. Gimmel, 2022, unpublished data).
Coccidophilus Brèthes, 1905
Nomenclatural Authority: Escalona & Ślipiński (2011)
Notes. One species of Coccidophilus has been recorded from California (Gordon, 1985).
Coccidophilus atronitens (Casey, 1899)
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Santa Cruz (16 SBMNH), Santa Rosa (6 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Microweisea Cockerell, 1903
Nomenclatural Authority: Escalona & Ślipiński (2011)
Notes. Certain species in this genus were until recently known as Gnathoweisea Gordon, 1970, a genus synonymized with Microweisea by Escalona & Ślipiński (2011).
Microweisea undetermined species
Literature Records: none
Digitized Records: Santa Catalina (1 iNat)
Nipus Casey, 1899
Nomenclatural Authority: Escalona & Ślipiński (2011)
Notes. Three species of the genus Nipus are known from California (M. L. Gimmel, 2022, unpublished data).
Nipus niger Casey, 1899
Nomenclatural Authority: Gordon (1985)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Serangiini
Notes. One genus and two species of Serangiini have been recorded from California (Gordon, 1985).
Delphastus Casey, 1899
Nomenclatural Authority: Escalona & Ślipiński (2011)
Notes. Two species of Delphastus have been recorded from California (Gordon, 1985).
Delphastus catalinae (Horn, 1895)
Nomenclatural Authority: Gordon (1985)
Literature Records: San Clemente (Fall, 1897: 237), Santa Catalina (Horn, 1895: 83; Fall, 1897: 237; Casey, 1899: 112; Fall, 1901: 85; Gordon, 1970: 367; Gordon, 1985: 64), Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: Santa Catalina (1 LACM; 1 SBMNH), Santa Cruz (3 SBMNH)
Range: Also known from mainland (Gordon, 1985).
Notes. Horn (1895) and Fall (1897, 1901) recorded this species as Cryptognatha catalinae. The species was originally (Horn, 1895; Casey, 1899) considered endemic to Santa Catalina Island. Fall (1897: 239) doubted its taxonomic validity.
Corylophidae
Notes. This family contains five tribes, six genera, and 14 species known from California (M. L. Gimmel, 2022, unpublished data), all of which belong to the subfamily Corylophinae. The classification used here follows Robertson et al. (2013). This family is very poorly understood in North America, and modern keys to species do not exist for most genera.
Aenigmaticini
Notes. One genus and species of Aenigmaticini occurs in California (Pakaluk, 1985).
Aenigmaticum Matthews, 1888
Nomenclatural Authority: Robertson et al. (2013)
Notes. One species of Aenigmaticum occurs in California (Pakaluk, 1985). This genus was reviewed by Pakaluk (1985).
Aenigmaticum californicum Casey, 1889
Nomenclatural Authority: Pakaluk (1985)
Literature Records: Anacapa (Miller & Miller, 1985: 126), Santa Barbara (Miller & Miller, 1985: 126)
Digitized Records: Anacapa (7 SBMNH), San Miguel (3 SBMNH), San Nicolas (8 SBMNH), Santa Barbara (2 SBMNH)
Range: Also known from mainland (Pakaluk, 1985).
Notes. Reported from Erophyllum, Hemizonia (both Asteraceae) and Frankenia (Frankeniaceae) on Santa Barbara Island by Miller & Miller (1985).
Orthoperini
Notes. One genus and four species of Orthoperini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Orthoperus Stephens, 1829
Nomenclatural Authority: Bowestead & Leschen (2002), Robertson et al. (2013)
Notes. Four species of Orthoperus have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Orthoperus undetermined species
Literature Records: none
Digitized Records: Santa Cruz (9 SBMNH)
Sericoderini
Notes. One genus and three species of Sericoderini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Sericoderus Stephens, 1829
Nomenclatural Authority: Bowestead & Leschen (2002), Robertson et al. (2013)
Notes. Three species of Sericoderus have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Sericoderus undetermined species
Literature Records: none
Digitized Records: San Nicolas (3 SBMNH), Santa Cruz (3 SBMNH), Santa Rosa (1 SBMNH)
Endomychidae
Notes. Six subfamilies, eight genera, and 13 species of Endomychidae have been recorded from California (Shockley, Tomaszewska & McHugh, 2009).
Lycoperdininae
Notes. Two genera and four species of Lycoperdininae have been recorded from California (Shockley, Tomaszewska & McHugh, 2009).
Aphorista Gorham, 1873
Nomenclatural Authority: Shockley, Tomaszewska & McHugh (2009)
Notes. Two species of Aphorista have been recorded from California (Shockley, Tomaszewska & McHugh, 2009).
Aphorista morosa (LeConte, 1859)
Nomenclatural Authority: Shockley, Tomaszewska & McHugh (2009)
Literature Records: Santa Rosa (Fall, 1897: 237)
Digitized Records: Santa Catalina (2 LACM), Santa Cruz (2 SBMNH; 1 UCSB), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Shockley, Tomaszewska & McHugh, 2009).
Latridiidae
Notes. Two subfamilies, 13 genera, and 61 species of Latridiidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Corticariinae
Notes. Five genera and 31 species of Corticariinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Corticaria Marsham, 1802
Nomenclatural Authority: Rücker (2021)
Notes. Ten species of Corticaria have been reported from California (M. L. Gimmel, 2022, unpublished data).
Corticaria undetermined species
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: Santa Rosa (2 SBMNH)
Notes. Fall’s (1897) above record refers to the genus only.
Corticarina Reitter, 1881
Nomenclatural Authority: Rücker (2021)
Literature Records (genus-only): Santa Cruz (Naughton et al., 2014: 303)
Digitized Records (genus-only): Anacapa (5 SBMNH), San Clemente (5 SBMNH), San Miguel (25 SBMNH), San Nicolas (7 SBMNH), Santa Barbara (15 SBMNH), Santa Catalina (1 SBMNH), Santa Cruz (51 SBMNH), Santa Rosa (14 SBMNH)
Notes. The record from Naughton et al. (2014) refers to the genus only, and probably represents one of the four species below. Nine species of Corticarina have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Corticarina cavicollis (Mannerheim, 1844)
Nomenclatural Authority: Rücker (2021)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Rücker, 2021).
Corticarina herbivagans (LeConte, 1855)
Nomenclatural Authority: Rücker (2021)
Literature Records: San Miguel (Miller & Miller, 1985: 127), Santa Barbara (Miller & Miller, 1985: 127)
Digitized Records: Santa Barbara (1 SBMNH)
Range: Also known from mainland (Rücker, 2021).
Corticarina milleri Andrews, 1992
Nomenclatural Authority: Andrews (1992), Rücker (2021)
Literature Records: San Miguel (Andrews, 1992: 278), San Nicolas (Andrews, 1992: 278), Santa Barbara (Andrews, 1992: 277), Santa Rosa (Andrews, 1992: 278)
Digitized Records: Anacapa (1 SBMNH), Santa Cruz (1 SBMNH)
Range: Endemic (Andrews, 1992).
Corticarina minuta (Fabricius, 1792)
Nomenclatural Authority: Rücker (2021)
Literature Records: none
Digitized Records: Anacapa (1 SBMNH), Santa Cruz (4 SBMNH)
Range: Also known from mainland (Rücker, 2021).
Fuchsina Fall, 1899
Nomenclatural Authority: Rücker (2021)
Notes. Two described species of Fuchsina have been reported from California (Andrews, 1976c). These were revised by Andrews (1976c), but an apparently undescribed species occurs on the Channel Islands.
Fuchsina undescribed species
Literature Records: Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: San Clemente (15 SBMNH), Santa Catalina (47 SBMNH), Santa Cruz (29 SBMNH), Santa Rosa (16 SBMNH)
Range: Endemic (M. L. Gimmel, 2021, personal observation).
Notes. The record from Naughton et al. (2014) refers to the genus only. The SBMNH specimens from the Channel Islands are morphologically different from either of the two described species of Fuchsina as circumscribed by Andrews (1976c). The Channel Island specimens are similar to Fuchsina occulta Fall, 1899 in the lack of eye facets and antenna with 10 antennomeres, but have a shorter, broader pronotum and shorter elytra (M. L. Gimmel, 2021, personal observation).
Melanophthalma Motschulsky, 1866
Nomenclatural Authority: Rücker (2021)
Literature Records (genus-only): Santa Cruz (Naughton et al., 2014: 303)
Digitized Records (genus-only): Anacapa (1 SBMNH), San Clemente (2 SBMNH), San Nicolas (2 SBMNH), Santa Catalina (9 SBMNH), Santa Cruz (5 SBMNH), Santa Rosa (1 SBMNH)
Notes. Naughton et al. (2014) mistakenly reported this genus as “Melanophthalmus”. Eight species of Melanophthalma have been reported from California in two subgenera, Cortilena Motschulsky, 1867 and Melanophthalma (s.str.) (M. L. Gimmel, 2022, unpublished data).
Melanophthalma (Cortilena) casta Fall, 1899
Nomenclatural Authority: Rücker (2021)
Literature Records: San Nicolas (Miller & Miller, 1985: 127), Santa Barbara (Miller & Miller, 1985: 127)
Digitized Records: Santa Barbara (1 SBMNH)
Range: Also known from mainland (Fall, 1899).
Notes. Recorded by Miller & Miller (1985) as Cortilena casta.
Melanophthalma (Melanophthalma) americana (Mannerheim, 1844)
Nomenclatural Authority: Rücker (2021)
Literature Records: San Clemente (Fall, 1897: 237), San Miguel (Cockerell, 1940: 286), Santa Catalina (Fall, 1897: 237)
Digitized Records: Santa Cruz (3 SBMNH)
Range: Also known from mainland (Rücker, 2021).
Notes. Reported by Fall (1897) as Corticaria distinguenda Comolli, 1837, but specimens in North American identified as this species are presently known as M. americana.
Melanophthalma (Melanophthalma) insularis Fall, 1899
Nomenclatural Authority: Rücker (2021)
Literature Records: San Clemente (Fall, 1899: 174; Fall, 1901: 102; Miller, 1985a: 20)
Digitized Records: none
Range: Endemic (Fall, 1899, 1901; Miller, 1985a).
Notes. The status of this purportedly endemic taxon has not been reviewed since Fall’s (1899) original description.
Latridiinae
Notes. Eight genera and 30 species of Latridiinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Cartodere Thomson, 1859
Nomenclatural Authority: Rücker (2021)
Notes. Four species of Cartodere in two subgenera, Aridius Motschulsky, 1866 and Cartodere (s.str.), occur in California (M. L. Gimmel, 2022, unpublished data).
Cartodere (Aridius) australica (Belon, 1887)
Nomenclatural Authority: Rücker (2021)
Literature Records: none
Digitized Records: Santa Cruz (25 SBMNH)
Range: Also known from mainland (Rücker, 2021).
Notes. This species is presumably adventive in California.
Dienerella Reitter, 1911
Nomenclatural Authority: Rücker (2021)
Notes. Three species of Dienerella are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Dienerella undetermined species
Literature Records: Santa Catalina (Caterino & Chandler, 2010: 191)
Digitized Records: Santa Catalina (1 SBMNH)
Notes. More morphospecies exist in the SBMNH collection than there are named species known to occur in California (M. L. Gimmel, 2021, personal observation). Consequently, we have not attempted to identify the single Channel Islands specimen known to us.
Enicmus Thomson, 1859
Nomenclatural Authority: Rücker (2021)
Notes. Six species of Enicmus have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Enicmus aterrimus Motschulsky, 1866
Nomenclatural Authority: Rücker (2021)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Rücker, 2021).
Metophthalmus Motschulsky, 1850
Nomenclatural Authority: Rücker (2021)
Literature Records (genus-only): Santa Catalina (Caterino & Chandler, 2010: 191), Santa Cruz (Naughton et al., 2014: 303)
Notes. This genus was revised by Andrews (1976b). Six species of this genus have been reported from California (Andrews, 1976b) belonging to two subgenera, Metatypus Belon, 1897 and Metophthalmus (s.str.), three of which are reported from the Channel Islands below. Caterino & Chandler (2010: 191) were first to mention the presence of this genus on Santa Catalina. Five of the 12 specimens of Metophthalmus reported from Santa Cruz by Naughton et al. (2014) were not determined to species; vouchers for these are not located in SBMNH (M. L. Gimmel, 2021, personal observation). Almost certainly the latter represent one or more of the three species recorded below, though two additional species, Metophthalmus kanei Andrews, 1976 and Metophthalmus septemstriatus Hatch, 1962, may occur on the Channel Islands.
Metophthalmus (Metatypus) haigi Andrews, 1976
Nomenclatural Authority: Rücker (2021)
Literature Records: Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: San Clemente (3 SBMNH), Santa Catalina (3 SBMNH), Santa Cruz (7 SBMNH)
Range: Also known from mainland (Andrews, 1976b; Rücker, 2021).
Metophthalmus (Metatypus) rudis Fall, 1899
Nomenclatural Authority: Rücker (2021)
Literature Records: Santa Cruz (Andrews, 1976b: 53; Naughton et al., 2014: 303)
Digitized Records: San Clemente (26 SBMNH), Santa Catalina (42 SBMNH), Santa Cruz (14 SBMNH), Santa Rosa (24 SBMNH)
Range: Also known from mainland (Andrews, 1976b; Rücker, 2021).
Metophthalmus (Metatypus) trux Fall, 1899
Nomenclatural Authority: Rücker (2021)
Literature Records: Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: San Clemente (5 SBMNH), Santa Catalina (3 SBMNH), Santa Cruz (12 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Andrews, 1976b; Rücker, 2021).
Revelieria Perris, 1869
Nomenclatural Authority: Rücker (2021)
Notes. One species of Revelieria has been reported from California (Rücker, 2021).
Revelieria californica Fall, 1899
Nomenclatural Authority: Rücker (2021)
Literature Records: none
Digitized Records: Santa Cruz (8 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Fall, 1899).
Stephostethus LeConte, 1878
Nomenclatural Authority: Rücker (2021)
Notes. Four species of Stephostethus have been reported from California (M. L. Gimmel, 2022, unpublished data).
Stephostethus armatulus (Fall, 1899)
Nomenclatural Authority: Rücker (2021)
Literature Records: Santa Catalina (Fall, 1899: 118)
Digitized Records: none
Range: Also known from mainland (Fall, 1899; Rücker, 2021).
Notes. Fall (1899) recorded this species as Lathridius armatulus.
Stephostethus costicollis (LeConte, 1855)
Nomenclatural Authority: Rücker (2021)
Literature Records: Santa Catalina (Fall, 1897: 237; Cockerell, 1940: 286)
Digitized Records: San Clemente (1 SBMNH), Santa Catalina (1 SBMNH)
Range: Also known from mainland (Rücker, 2021).
Notes. This species was recorded as Coninomus fulvipennis Mannerheim by Fall (1897), and corrected to Latridius costicollis by Cockerell (1940). It has since been placed in the genus Stephostethus.
Stephostethus liratus (LeConte, 1863)
Nomenclatural Authority: Rücker (2021)
Literature Records: none
Digitized Records: Santa Rosa (2 SBMNH)
Range: Also known from mainland (Rücker, 2021).
EROTYLOIDEA
Erotylidae
Notes. Three subfamilies, six genera, and 11 species of Erotylidae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Cryptophilinae
Notes. One genus and species of Cryptophilinae has been recorded from California (M. L. Gimmel, 2022, unpublished data).
Cryptophilus Reitter, 1874
Nomenclatural Authority: Gimmel, Leschen & Esser (2019)
Notes. One species of the genus Cryptophilus has been recorded from California (M. L. Gimmel, 2022, unpublished data).
Cryptophilus angustus (Rosenhauer, 1856)
Nomenclatural Authority: Esser (2017)
Literature Records: none
Digitized Records: Santa Cruz (3 SBMNH)
Range: Also known from mainland (Esser, 2017).
Notes. This species was long known as Cryptophilus integer (Heer, 1841), but the type of that species was discovered to belong to Cryptophagidae. Cryptophilus angustus is the proper name for this species, which was introduced from the Palearctic realm (Esser, 2017; Gimmel, Leschen & Esser, 2019).
Erotylinae
Notes. Four genera and eight species of Erotylinae have been recorded from California (Boyle, 1956; M. L. Gimmel, 2022, unpublished data).
Dacne Latreille, 1796
Nomenclatural Authority: Boyle (1956)
Notes. Four species of Dacne have been recorded from California (Boyle, 1956).
Dacne californica (Horn, 1870)
Nomenclatural Authority: Boyle (1956)
Literature Records: Santa Catalina (Cockerell, 1940: 286; Boyle, 1956: 142), Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: San Clemente (3 SBMNH), Santa Catalina (5 SBMNH), Santa Cruz (6 SBMNH), Santa Rosa (13 SBMNH)
Range: Also known from mainland (Boyle, 1956).
Notes. The island vouchers of this species housed in SBMNH are certainly Dacne (s.str.), which has just one described species in California (D. californica). However, they are morphologically different from mainland exemplars of that species, being narrower and more setose, and the prosternal lines are differently shaped. These may prove to be a distinct, undescribed species.
NITIDULOIDEA
Kateretidae
Notes. Four genera and six species of Kateretidae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Amartus LeConte, 1861
Nomenclatural Authority: Cline & Audisio (2010)
Notes. Two species of Amartus have been reported from California (Savage & Seeno, 1981). Savage & Seeno (1981) reviewed the genus Amartus for North America.
Amartus tinctus (Mannerheim, 1843)
Nomenclatural Authority: Savage & Seeno (1981)
Literature Records: San Clemente (Savage & Seeno, 1981: 80), San Miguel (Cockerell, 1940: 286), Santa Rosa (Savage & Seeno, 1981: 80)
Digitized Records: San Clemente (6 LACM), Santa Rosa (9 SBMNH)
Range: Also known from mainland (Savage & Seeno, 1981).
Heterhelus Jacquelin du Val, 1858
Nomenclatural Authority: Cline & Audisio (2010)
Notes. One species of Heterhelus has been reported from California (Parsons, 1943).
Heterhelus sericans (LeConte, 1869)
Nomenclatural Authority: Habeck (2002)
Literature Records: Santa Catalina (Fall, 1897: 237)
Digitized Records: none
Range: Also known from mainland (Fall, 1901; Habeck, 2002).
Notes. Fall (1897, 1901) recorded this species as Cercus sericans, and Fall (1901: 98) listed it as occurring “throughout Southern California and the adjacent islands”.
Monotomidae
Notes. Two subfamilies, eight genera, and 19 species of Monotomidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Monotominae
Notes. Seven genera and 14 species of Monotominae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Hesperobaenus LeConte, 1861
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Notes. The genus Hesperobaenus contains two species occurring in California (Bousquet, 2002b). The genus was revised for North America by Bousquet (2002b).
Hesperobaenus abbreviatus (Motschulsky, 1845)
Nomenclatural Authority: Bousquet (2002b)
Literature Records: Santa Cruz (Bousquet, 2002b: 210)
Digitized Records: Anacapa (1 SBMNH), Santa Cruz (9 SBMNH), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Bousquet, 2002b).
Macreurops Casey, 1916
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Notes. One species of Macreurops is known from California (Bousquet, 2002a).
Macreurops longicollis (Horn, 1879)
Nomenclatural Authority: Bousquet (2002a)
Literature Records: none
Digitized Records: Santa Cruz (15 SBMNH)
Range: Also known from mainland (Bousquet, 2002a).
Phyconomus LeConte, 1861
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Notes. One species of Phyconomus is known from California (Bousquet, 2002a).
Phyconomus marinus (LeConte, 1858)
Nomenclatural Authority: Bousquet (2002a)
Literature Records: none
Digitized Records: San Miguel (10 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Bousquet, 2002a).
Nitidulidae
Notes. Eight subfamilies, 22 genera, and 63 species of Nitidulidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Carpophilinae
Notes. Four genera and 17 species of Carpophilinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Carpophilus Stephens, 1829
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Digitized Records (genus-only): Santa Cruz (1 LACM; 3 SBMNH), Santa Rosa (4 LACM; 5 SBMNH)
Notes. Eleven species of Carpophilus (sensu Powell et al., 2020) are known from California (M. L. Gimmel, 2022, unpublished data).
Carpophilus (Ecnomorphus) discoideus LeConte, 1858
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Parsons, 1943).
Carpophilus (Ecnomorphus) ligneus Murray, 1864
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Literature Records: none
Digitized Records: Anacapa (1 SBMNH), San Miguel (1 SBMNH), San Nicolas (1 SBMNH)
Range: Also known from mainland (McElrath, Gimmel & Powell, 2021).
Nitops Murray, 1864
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Notes. This genus contains a single species in California (M. L. Gimmel, 2022, unpublished data).
Nitops pallipennis (Say, 1823)
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Literature Records: San Clemente (Fall, 1897: 237), Santa Catalina (Seavey, 1892: 262; Fall, 1897: 237; Grant & Connell, 1979: 100; Grant & Grant, 1979: 323), Santa Cruz (LeConte, 1876: 299; Fall, 1897: 237; Fall & Davis, 1934: 143)
Digitized Records: San Clemente (18 SBMNH), San Miguel (3 SBMNH), San Nicolas (3 SBMNH), Santa Barbara (9 SBMNH), Santa Catalina (1 LACM; 3 SBMNH), Santa Cruz (66 LACM; 3 SBMNH), Santa Rosa (29 LACM; 7 SBMNH)
Range: Also known from mainland (Parsons, 1943; Grant & Connell, 1979; Grant & Grant, 1979).
Notes. Recorded from flowers of Opuntia littoralis var. littoralis (Engelm.) Cockerell on Santa Catalina (Grant & Connell, 1979; Grant & Grant, 1979). This species was recorded by all authors cited above prior to Powell et al. (2020) as Carpophilus pallipennis.
Cryptarchinae
Notes. Two genera and seven species of Cryptarchinae have been recorded from California (Parsons, 1943; McCoshum, 2012).
Cryptarcha Shuckard, 1839
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Notes. Three species of Cryptarcha have been recorded from California (Parsons, 1943).
Cryptarcha gila Parsons, 1938
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Literature Records: Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: Santa Catalina (2 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (Parsons, 1943).
Glischrochilus Reitter, 1873
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Notes. Four species of Glischrochilus are known from California (M. L. Gimmel, 2022, unpublished data).
Glischrochilus quadrisignatus (Say, 1835)
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Literature Records: Santa Catalina (McCoshum, 2012: 348)
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (McCoshum, 2012).
Notes. This species is adventive in California (McCoshum, 2012).
Glischrochilus sanguinolentus (Olivier, 1790)
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Literature Records: Santa Catalina (McCoshum, 2012: 348)
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (McCoshum, 2012).
Notes. This species is adventive in California (McCoshum, 2012).
Meligethinae
Notes. Two genera and two species of Meligethinae have been recorded from California (Parsons, 1943; Easton, 1955). Easton (1955) revised the species in this subfamily for North America. Generic concepts and some species concepts, however, have changed since then (Audisio et al., 2009).
Brassicogethes Audisio & Cline, 2009
Nomenclatural Authority: Audisio et al. (2009)
Notes. This genus contains one species in California (Parsons, 1943; Audisio et al., 2009).
Brassicogethes aeneus (Fabricius, 1775)
Nomenclatural Authority: Audisio et al. (2009)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (Parsons, 1943; Easton, 1955).
Notes. This species is Holarctic in distribution (Audisio et al., 2009).
Nitidulinae
Notes. Eight genera and 19 species of Nitidulinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Nitidula Fabricius, 1775
Nomenclatural Authority: Parsons (1943), McElrath, Gimmel & Powell (2021)
Notes. This genus contains three species in California (Parsons, 1943). Parsons (1943) provided a key to species in North America.
Nitidula flavomaculata Rossi, 1790
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH)
Range: Also known from mainland (Parsons, 1943).
Notes. This species was introduced into North America from the Mediterranean region (Parsons, 1943).
Thalycra Erichson, 1843
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Notes. This genus contains eight species in California (Howden, 1961; M. L. Gimmel, 2022, unpublished data). Howden (1961) provided a key to New World species.
Thalycra undetermined species
Literature Records: none
Digitized Records: Santa Rosa (8 SBMNH)
CUCUJOIDEA
Cryptophagidae
Notes. Two subfamilies, 11 genera, and 60 species of Cryptophagidae are known to occur in California (M. L. Gimmel, 2022, unpublished data). Pelletier & Hébert (2019) provided a revision and identification guide to the species of Cryptophagidae in the northern US and Canada, which is helpful for making identifications in the southern US as well.
Atomariinae
Notes. Two genera and 28 species of Atomariinae, all belonging to Atomariini, are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Atomaria Stephens, 1829
Nomenclatural Authority: Pelletier & Hébert (2019)
Literature Records (genus-only): Santa Rosa (Fall, 1897: 237)
Notes. Fall (1897) did not specify subgenus when he cited the record of “Atomaria sp.”. Pelletier & Hébert (2019) provided a key to all North American species of the genus. Twenty-seven species of Atomaria have been reported from California, 11 from the subgenus Anchicera Thomson, 1863 and 16 from the subgenus Atomaria (s.str.) (M. L. Gimmel, 2022, unpublished data).
Atomaria (Anchicera) lewisi Reitter, 1877
Nomenclatural Authority: Pelletier & Hébert (2019)
Literature Records: none
Digitized Records: Santa Cruz (6 SBMNH)
Range: Also known from mainland (Pelletier & Hébert, 2019).
Notes. This species is adventive from Europe (Pelletier & Hébert, 2019).
Atomaria (Anchicera) nubipennis Casey, 1900
Nomenclatural Authority: Pelletier & Hébert (2019)
Literature Records: none
Digitized Records: San Clemente (20 SBMNH)
Range: Also known from mainland (Pelletier & Hébert, 2019).
Atomaria (Atomaria) puella (Casey, 1900)
Nomenclatural Authority: Pelletier & Hébert (2019)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Pelletier & Hébert, 2019).
Cryptophaginae: Cryptophagini
Notes. Two tribes, nine genera, and 32 species of Cryptophaginae, of which seven genera and 29 species belong to Cryptophagini, are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Cryptophagus Herbst, 1792
Nomenclatural Authority: Pelletier & Hébert (2019)
Notes. Woodroffe & Coombs (1961) revised the genus Cryptophagus for North America. Pelletier & Hébert (2019) provided an updated key to and illustrations of most North American species. In California, 22 species have been reported (M. L. Gimmel, 2022, unpublished data).
Cryptophagus tuberculosus Mäklin, 1853
Nomenclatural Authority: Woodroffe & Coombs (1961), Pelletier & Hébert (2019)
Literature Records: San Clemente (Fall, 1897: 237; Cockerell, 1940: 286), Santa Catalina (Fall, 1897: 237; Cockerell, 1940: 286)
Digitized Records: San Clemente (5 SBMNH), Santa Cruz (5 SBMNH)
Range: Also known from mainland (Woodroffe & Coombs, 1961).
Notes. Fall (1897) recorded this species as “Cryptophagus sp.”, which Cockerell (1940) indicated as Cryptophagus debilis LeConte, 1858 based on communication from H.C. Fall. Cryptophagus debilis was synonymized with C. tuberculosus by Woodroffe & Coombs (1961).
Laemophloeidae, NEW FAMILY RECORD
Notes. Eight genera and 16 species of Laemophloeidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Narthecius LeConte, 1861
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Notes. Two species of Narthecius are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Narthecius striaticeps Fall, 1907
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Fall & Cockerell, 1907).
Phalacridae
Notes. Five genera and 21 species of Phalacridae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Phalacrus Paykull, 1800
Nomenclatural Authority: Gimmel (2013)
Notes. This genus needs revision; the species in North America are not currently identifiable (Gimmel, 2013).
Phalacrus undetermined species 1
Literature Records: none
Digitized Records: San Nicolas (6 SBMNH)
Notes. This species has microsculpture on the elytra, the left mandible with a ventral tooth, and a prominent metaventral process that exceeds the mesocoxae; this possibly represents Phalacrus conjunctus Casey, 1890 (M. L. Gimmel, 2021, personal observation).
Phalacrus undetermined species 2
Literature Records: Santa Cruz (Naughton et al., 2014: 304)
Digitized Records: Santa Cruz (3 SBMNH)
Notes. This species has no microsculpture on the elytra, the left mandible with a ventral tooth, and a short metaventral process not exceeding the mesocoxae; this possibly represents Phalacrus ovalis LeConte, 1856 (M. L. Gimmel, 2021, personal observation). Santa Cruz Island vouchers from the Naughton et al. (2014) study citing “Phalacrus sp.” were examined by MLG and belong to this morphospecies.
Silvanidae, NEW FAMILY RECORD
Notes. Two subfamilies, nine genera, and 14 species of Silvanidae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Silvanoprus Reitter, 1911
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Notes. One introduced species of Silvanoprus is now known from California; the record below represents a new state record for the genus.
Silvanoprus angusticollis (Reitter, 1876)
Nomenclatural Authority: McElrath, Gimmel & Powell (2021)
Literature Records: none
Digitized Records: Santa Cruz (6 SBMNH)
Range: Also known from mainland (McElrath, Gimmel & Powell, 2021).
Notes. This represents a new state record for California. This species is adventive in North America.
CHRYSOMELOIDEA
Cerambycidae
Notes. Seven subfamilies, 143 genera, and 317 species of Cerambycidae are known to occur in California (M. L. Gimmel, 2022, unpublished data). Bezark & Monné (2013) provided a nomenclatural checklist of all New World Cerambycidae. Linsley (1962, 1963, 1964), Linsley & Chemsak (1984, 1995), and Chemsak (1996, 2005) monographed the North American fauna. The subfamily Parandrinae occurs on the nearby mainland but is not known from the Channel Islands.
Cerambycinae
Notes. Twenty-three tribes, 72 genera, and 161 species of Cerambycinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Callidiini
Notes. Six genera and 35 species of Callidiini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Callidiellum Linsley, 1940
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Two species of Callidiellum are known to occur in California, plus one species (C. rufipenne below) recorded only as an interception (M. L. Gimmel, 2022, unpublished data).
Callidiellum rufipenne (Motschulsky, 1860)
Nomenclatural Authority: Linsley (1964), Miller & Miller (1985)
Literature Records: Santa Barbara (Miller & Miller, 1985: 130)
Digitized Records: none
Range: Also known from mainland (Linsley, 1964).
Notes. Introduced to North America from eastern Asia; the Santa Barbara Island record is probably an interception and does not represent a breeding population (see Miller & Miller, 1985).
Phymatodes Mulsant, 1839
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Fifteen species of Phymatodes are known to occur in California (M. L. Gimmel, 2022, unpublished data). Swift & Ray (2010) presented a revised key to the North American species.
Phymatodes decussatus (LeConte, 1857)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: Santa Rosa (Fall, 1897: 238)
Digitized Records: Santa Cruz (2 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Linsley, 1964).
Notes. According to Linsley (1964: 50), the subspecies occurring in coastal California is P. d. decussatus. Fall (1897) reported this species as Phymatodes juglandis Leng, 1890, and listed the record with a question mark; this record was included in the species’ synonymy in Linsley (1964: 52). However, P. juglandis was recently made a junior synonym of P. decussatus by Swift & Ray (2010: 42).
Phymatodes grandis Casey, 1912
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Catalina (20 LACM; 8 SBMNH; 2 iNat), Santa Cruz (4 LACM; 7 SBMNH)
Range: Also known from mainland (Linsley, 1964).
Notes. The species Phymatodes obscurus (LeConte, 1859) was given the unnecessary replacement name of Phymatodes lecontei Linsley, 1938, but the proper name for this species is P. grandis (see Swift & Ray, 2010).
Clytini
Notes. Eight genera and 30 species of Clytini have been recorded from California (Linsley, 1964; M. L. Gimmel, 2022, unpublished data).
Xylotrechus Chevrolat, 1860
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Eight species of Xylotrechus have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Xylotrechus insignis LeConte, 1873
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: Santa Catalina (Fall, 1897: 238; Fall, 1901: 147; Hopping, 1932: 542; Cockerell, 1940: 286)
Digitized Records: Santa Catalina (1 CASC; 4 LACM)
Range: Also known from mainland (Fall, 1901; Linsley, 1964).
Notes. Fall (1897, 1901) recorded this species as Xylotrechus obliteratus LeConte, 1873, which was amended to X. insignis by communication of E.G. Linsley to Cockerell (1940) and in Hopping (1932). Fall (1897) reported that it occurred on willows.
Xylotrechus nauticus (Mannerheim, 1843)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: Santa Cruz (Linsley, 1964: 109 [map])
Digitized Records: Santa Catalina (2 CASC), Santa Cruz (7 CASC; 5 SBMNH)
Range: Also known from mainland (Linsley, 1964).
Eburiini
Notes. Two genera and three species of Eburiini have been recorded from California (Linsley, 1962, 1963).
Enaphalodes Haldeman, 1847
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Two species of Enaphalodes have been recorded from California (Linsley, 1963).
Enaphalodes hispicornis (Linnaeus, 1767)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: Santa Catalina (Fall, 1897: 238; Fall, 1901: 144; Garnett, 1918: 177; Linsley, 1963: 66 [map])
Digitized Records: Santa Catalina (6 LACM)
Range: Also known from mainland (Fall, 1901; Garnett, 1918; Linsley, 1963).
Notes. This species was reported by Fall (1897, 1901) and Garnett (1918) as Romaleum simplicicolle (Haldeman, 1847), which is now recognized as a synonym of the variable species E. hispicornis (see Linsley, 1963: 64).
Hesperophanini
Notes. Six genera and eight species of Hesperophanini have been recorded from California (Linsley, 1962).
Brothylus LeConte, 1859
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Two species of Brothylus have been recorded from California (Linsley, 1962).
Brothylus gemmulatus LeConte, 1859
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Catalina (22 LACM; 1 iNat)
Range: Also known from mainland (Linsley, 1962).
Holopleurini
Notes. One species of Holopleurini has been recorded from California (Linsley, 1962).
Holopleura LeConte, 1873
Nomenclatural Authority: Bezark & Monné (2013)
Notes. One species of Holopleura has been recorded from California (Linsley, 1962).
Holopleura marginata LeConte, 1873
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Catalina (3 LACM)
Range: Also known from mainland (Linsley, 1962).
Hyboderini
Notes. Four genera and six species of Hyboderini have been recorded from California (Linsley, 1963).
Callimus Mulsant, 1864
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Two species of Callimus have been recorded from California (Linsley, 1963, as Lampropterus Mulsant, 1863).
Callimus ruficollis (LeConte, 1873)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Catalina (3 LACM), Santa Cruz (29 LACM; 1 SBMNH)
Range: Also known from mainland (Linsley, 1963).
Notes. Linsley (1963) reported this species as Lampropterus ruficollis.
Megobrium LeConte, 1873
Nomenclatural Authority: Bezark & Monné (2013)
Notes. One species of Megobrium has been recorded from California (Linsley, 1963).
Megobrium edwardsi LeConte, 1873
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: Santa Rosa (LeConte, 1873: 193; Fall, 1897: 238; Fall, 1901: 145; Garnett, 1918: 206; Linsley, 1963: 150)
Digitized Records: Santa Catalina (2 SBMNH; 4 LACM)
Range: Also known from mainland (Fall, 1901; Garnett, 1918; Linsley, 1963).
Notes. Often misspelled M. edwardsii, this species was considered endemic at the time of its description (LeConte, 1873).
Methiini
Notes. Two genera and eight species of Methiini are known to occur in California (Linsley, 1962; M. L. Gimmel, 2022, unpublished data).
Styloxus LeConte, 1873
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Two species of Styloxus have been recorded from California (Linsley, 1962).
Styloxus fulleri (Horn, 1880)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (1 iNat)
Range: Also known from mainland (Linsley, 1962).
Notes. According to Linsley (1962: 40), the subspecies occurring in California is S. f. californicus (Fall, 1901).
Oemini
Notes. Six genera and six species of Oemini are known to occur in California (Linsley, 1962; M. L. Gimmel, 2022, unpublished data).
Paranoplium Casey, 1924
Nomenclatural Authority: Bezark & Monné (2013)
Notes. One species of Paranoplium has been recorded from California (Linsley, 1962).
Paranoplium gracile (LeConte, 1881)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: Santa Catalina (Fall, 1897: 238; Fall, 1901: 144; Garnett, 1918: 176)
Digitized Records: Santa Catalina (1 LACM)
Range: Also known from mainland (Fall, 1901; Garnett, 1918; Linsley, 1962).
Notes. This species was reported by Fall (1897, 1901) and Garnett (1918) as Oeme gracilis. According to Linsley (1962: 20), the coastal California subspecies is P. g. gracile.
Psebiini
Notes. One species of Psebiini has been recorded from California (Linsley, 1963).
Nathrius Brèthes, 1916
Nomenclatural Authority: Bezark & Monné (2013)
Notes. One species of Nathrius is known to occur in California (Linsley, 1963).
Nathrius brevipennis (Mulsant, 1839)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Linsley, 1963).
Notes. This species was introduced to North America from southern Europe (Linsley, 1963: 155; Bezark & Monné, 2013).
Phoracanthini
Notes. One genus and two species of Phoracanthini are known to occur in California (Bezark & Monné, 2013).
Phoracantha Newman, 1840
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Two species of Phoracantha are now known to occur in California (Bezark & Monné, 2013).
Phoracantha recurva Newman, 1840
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH; 1 iNat), Santa Cruz (2 SBMNH)
Range: Also known from mainland (Bezark & Monné, 2013).
Notes. This species was introduced to North America from Australia (Bezark & Monné, 2013).
Phoracantha semipunctata (Fabricius, 1775)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Catalina (2 LACM), Santa Cruz (3 SBMNH)
Range: Also known from mainland (Bezark & Monné, 2013).
Notes. This species was introduced to North America from Australia (Bezark & Monné, 2013).
Lamiinae
Notes. Nine tribes, 20 genera, and 31 species of Lamiinae have been recorded from California (Linsley & Chemsak, 1984, 1995).
Acanthocinini
Notes. Seven genera and eight species of Acanthocinini have been recorded from California (Linsley & Chemsak, 1995).
Sternidocinus Dillon, 1956
Nomenclatural Authority: Bezark & Monné (2013)
Notes. One species of Sternidocinus occurs in California (Linsley & Chemsak, 1995).
Sternidocinus barbarus (Van Dyke, 1920)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: Santa Cruz (Dillon, 1956: 167; Linsley & Chemsak, 1995: 57)
Digitized Records: Santa Cruz (11 LACM; 5 SBMNH)
Range: Also known from mainland (Dillon, 1956; Linsley & Chemsak, 1995).
Parmenini
Notes. Two genera and two species of Parmenini have been recorded from California (Linsley & Chemsak, 1984).
Ipochus LeConte, 1852
Nomenclatural Authority: Bezark & Monné (2013)
Notes. One species of Ipochus occurs in California (Linsley & Chemsak, 1984).
Ipochus fasciatus LeConte, 1852
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: Anacapa (Linsley & Chemsak, 1984: 11 [map]; Miller & Miller, 1985: 130), San Clemente (Linsley & Chemsak, 1984: 11 [map]), San Miguel (Cockerell, 1940: 287; Linsley & Chemsak, 1984: 11 [map]; Miller & Miller, 1985: 130), Santa Barbara (Miller & Miller, 1985: 130), Santa Catalina (Fall, 1897: 238; Casey, 1913b: 281; Cockerell, 1940: 287; Miller & Miller, 1985: 130), Santa Cruz (Linsley & Chemsak, 1984: 11 [map]; Miller & Miller, 1985: 130; Naughton et al., 2014: 303), Santa Rosa (Linsley & Chemsak, 1984: 11 [map]; Miller & Miller, 1985: 130)
Digitized Records: Anacapa (3 CASC; 11 LACM; 2 SBMNH), San Miguel (5 LACM; 11 SBMNH), San Nicolas (1 SBMNH), Santa Barbara (1 LACM), Santa Catalina (3 CASC; 12 LACM; 6 SBMNH; 1 SDNHM; 1 USNM; 2 iNat), Santa Cruz (1 CASC; 1 LACM; 7 SBMNH; 1 iNat), Santa Rosa (1 CASC; 6 LACM; 1 SBMNH)
Range: Also known from mainland (Linsley & Chemsak, 1984).
Notes. Fall (1897) reported this species occurring “rather plentifully… under bark and on the branches of dead Rhus laurina (or R. integrifolia).” Casey (1913b) reported the presumed endemic Santa Catalina Island population as Ipochus catalinae Casey, 1913. This species was synonymized with I. fasciatus by Linsley & Chemsak (1984: 12), a finding already suggested by Cockerell (1940: 287).
Phytoeciini
Notes. Two genera and two species of Phytoeciini have been recorded from California (Linsley & Chemsak, 1995).
Oberea Mulsant, 1839
Nomenclatural Authority: Bezark & Monné (2013)
Notes. One species of Oberea has been recorded from California (Linsley & Chemsak, 1995).
Oberea quadricallosa LeConte, 1874
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Linsley & Chemsak, 1995).
Pogonocherini
Notes. Four genera and eight species of Pognocherini have been recorded from California (Linsley & Chemsak, 1984).
Lophopogonius Linsley, 1935
Nomenclatural Authority: Bezark & Monné (2013)
Notes. One species of Lophopogonius has been recorded from California (Linsley & Chemsak, 1984).
Lophopogonius crinitus (LeConte, 1873)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH), Santa Rosa (3 SBMNH)
Range: Also known from mainland (Linsley & Chemsak, 1984).
Saperdini
Notes. One genus and three species of Saperdini have been recorded from California (Linsley & Chemsak, 1984).
Saperda Fabricius, 1775
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Three species of Saperda have been recorded from California (Linsley & Chemsak, 1995).
Saperda horni Joutel, 1902
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Linsley & Chemsak, 1995).
Lepturinae
Notes. Six tribes, 35 genera, and 89 species of Lepturinae have been recorded from California (Chemsak, 2005). Chemsak (2005) provided a guide to the genera and species of the subfamily in North America.
Desmocerini
Notes. One genus and two species of Desmocerini have been recorded from California (Chemsak, 2005).
Desmocerus Dejean, 1821
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Two species of Desmocerus have been recorded from California (Chemsak, 2005).
Desmocerus californicus Horn, 1881
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (3 LACM), Santa Rosa (1 LACM)
Range: Also known from mainland (Chemsak, 2005).
Notes. All island records of this species belong to D. c. californicus Horn, 1881. The other subspecies, D. c. dimorphus Fisher, 1921, is restricted to the Central Valley of California and is federally protected.
Lepturini
Notes. Eighteen genera and 48 species of Lepturini have been recorded from California (Chemsak, 2005).
Anastrangalia Casey, 1924
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Two species of Anastrangalia have been recorded from California (Chemsak, 2005).
Anastrangalia laetifica (LeConte, 1859)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (1 CASC; 1 LACM; 4 SBMNH; 1 iNat)
Range: Also known from mainland (Chemsak, 2005).
Strophiona Casey, 1913
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Two species of Strophiona have been recorded from California (Chemsak, 2005).
Strophiona tigrina Casey, 1913
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Catalina (2 SBMNH), Santa Cruz (1 LACM; 2 SBMNH)
Range: Also known from mainland (Chemsak, 2005).
Xestoleptura Casey, 1913
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Four species of Xestoleptura have been recorded from California (Chemsak, 2005).
Xestoleptura crassipes (LeConte, 1857)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Chemsak, 2005).
Necydalini
Notes. Two genera and seven species of Necydalini have been recorded from California (Chemsak, 2005). The tribe is sometimes recognized as a full subfamily (e.g., Bezark & Monné, 2013).
Necydalis Linnaeus, 1758
Nomenclatural Authority: Bezark & Monné (2013)
Notes. There are six California species of the Holarctic genus Necydalis (Chemsak, 2005).
Necydalis laevicollis LeConte, 1869
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Rosa (1 USNM)
Range: Also known from mainland (Chemsak, 2005).
Notes. The Santa Rosa Island record represents a significant southerly range extension for this species; the nearest recorded specimens are known from the San Francisco Bay area (Chemsak, 2005).
Rhagiini
Notes. Twelve genera and 35 species of Rhagiini have been recorded from California (Chemsak, 2005).
Brachysomida Casey, 1913
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Two species of Brachysomida have been recorded from California (Chemsak, 2005).
Brachysomida californica (LeConte, 1851)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Chemsak, 2005).
Centrodera LeConte, 1850
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Seven species of Centrodera have been recorded from California (Chemsak, 2005).
Centrodera autumnata Leech, 1963
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (2 LACM; 1 SBMNH)
Range: Also known from mainland (Chemsak, 2005).
Centrodera spurca (LeConte, 1857)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (2 iNat)
Range: Also known from mainland (Chemsak, 2005).
Stenocorus Geoffroy, 1762
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Four species of Stenocorus have been recorded from California (Chemsak, 2005).
Stenocorus (Stenocorus) vestitus (Haldeman, 1847)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (2 LACM; 11 SBMNH; 1 SDNHM)
Range: Also known from mainland (Chemsak, 2005).
Prioninae
Notes. Five tribes, six genera, and 12 species of Prioninae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Callipogonini
Notes. One genus and two species of Callipogonini have been recorded from California (Chemsak, 1996; Swift, Santos-Silva & Nearns, 2010).
Trichocnemis LeConte, 1851
Nomenclatural Authority: Bezark & Monné (2013)
Notes. This genus was reviewed by Swift, Santos-Silva & Nearns (2010). Two species occur in California (Chemsak, 1996; Swift, Santos-Silva & Nearns, 2010).
Trichocnemis spiculatus LeConte, 1851
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Linsley, 1962; Chemsak, 1996).
Notes. This species was indicated as Ergates spiculatus in Linsley (1962) and Chemsak (1996), but was returned to Trichocnemis in Swift, Santos-Silva & Nearns (2010).
Prionini
Notes. Two genera and five species of Prionini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Prionus Geoffroy, 1762
Nomenclatural Authority: Santos-Silva, Nearns & Swift (2016)
Notes. This genus was revised for the New World by Santos-Silva, Nearns & Swift (2016). Two species occur in California (Chemsak, 1996; Santos-Silva, Nearns & Swift, 2016).
Prionus (Prionus) californicus Motschulsky, 1845
Nomenclatural Authority: Santos-Silva, Nearns & Swift (2016)
Literature Records: none
Digitized Records: Santa Cruz (3 CASC; 19 LACM; 15 SBMNH; 2 UCSB; 2 iNat)
Range: Also known from mainland (Linsley, 1962; Chemsak, 1996; Santos-Silva, Nearns & Swift, 2016).
Notes. It is somewhat surprising that this large and readily recognizable species has not been reported in the literature from Santa Cruz Island before now.
Spondylidinae: Asemini
Notes. Three tribes, seven genera, and 16 species of Spondylidinae, of which four genera and nine species belong to Asemini, have been recorded from California (Chemsak, 1996).
Arhopalus Audinet-Serville, 1834
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Three species of this genus are known to occur in California (Chemsak, 1996). A key to the North American species was provided by Chemsak (1996).
Arhopalus asperatus (LeConte, 1859)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: Santa Catalina (Linsley, 1962: 71 [map])
Digitized Records: none
Range: Also known from mainland (Linsley, 1962; Chemsak, 1996).
Arhopalus productus (LeConte, 1850)
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: Santa Catalina (Linsley, 1962: 74 [map])
Digitized Records: none
Range: Also known from mainland (Linsley, 1962; Chemsak, 1996).
Asemum Eschscholtz, 1830
Nomenclatural Authority: Bezark & Monné (2013)
Notes. Three species of this genus are known to occur in California (Chemsak, 1996). A key to the North American species was provided by Chemsak (1996).
Asemum nitidum LeConte, 1873
Nomenclatural Authority: Bezark & Monné (2013)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Linsley, 1962; Chemsak, 1996).
Chrysomelidae
Notes. Nine subfamilies, 106 genera, and 436 species of Chrysomelidae have been recorded from California (Riley, Clark & Seeno, 2003; Kingsolver, 2004; M. L. Gimmel, 2022, unpublished data). The subfamilies Donaciinae and Synetinae have not been reported from the Channel Islands.
Bruchinae: Bruchini
Notes. Two tribes, 12 genera, and 43 species of Bruchinae, of which 10 genera and 35 species are in the tribe Bruchini, have been recorded from California (Kingsolver, 2004; M. L. Gimmel, 2022, unpublished data). Kingsolver (2004) reviewed the North American fauna of the subfamily.
Acanthoscelides Schilsky, 1905
Nomenclatural Authority: Kingsolver (2004)
Notes. Seventeen species of Acanthoscelides have been recorded from California (Kingsolver, 2004).
Acanthoscelides margaretae Johnson, 1970
Nomenclatural Authority: Kingsolver (2004)
Literature Records: none
Digitized Records: San Miguel (12 SBMNH), Santa Catalina (3 SBMNH), Santa Cruz (2 SBMNH), Santa Rosa (7 SBMNH)
Range: Also known from mainland (Kingsolver, 2004).
Acanthoscelides napensis Johnson, 1970
Nomenclatural Authority: Kingsolver (2004)
Literature Records: San Miguel (Miller & Davis, 1986: 550)
Digitized Records: San Clemente (1 LACM; 15 SBMNH), Santa Cruz (20 SBMNH), Santa Rosa (125 LACM; 16 SBMNH)
Range: Also known from mainland (Kingsolver, 2004).
Acanthoscelides pauperculus (LeConte, 1857)
Nomenclatural Authority: Kingsolver (2004)
Literature Records: San Miguel (Cockerell, 1940: 287), Santa Catalina (Fall, 1897: 238)
Digitized Records: none
Range: Also known from mainland (Kingsolver, 2004).
Notes. Recorded as Bruchus pauperculus by Fall (1897) and Cockerell (1940).
Acanthoscelides pullus (Fall, 1910)
Nomenclatural Authority: Kingsolver (2004)
Literature Records: Santa Catalina (Fall, 1910: 180)
Digitized Records: Anacapa (76 LACM; 1 SBMNH), San Clemente (10 LACM; 1 SBMNH), San Miguel (5 LACM; 1 SBMNH), San Nicolas (8 LACM; 1 SBMNH), Santa Cruz (2 LACM), Santa Rosa (57 LACM; 1 SBMNH)
Range: Also known from mainland (Kingsolver, 2004).
Notes. Fall (1910) recorded this species as Bruchus pullus.
Megacerus Fåhraeus, 1839
Nomenclatural Authority: Kingsolver (2004)
Notes. One species of Megacerus has been recorded from California (Kingsolver, 2004).
Megacerus (Megacerus) impiger (Horn, 1873)
Nomenclatural Authority: Kingsolver (2004)
Literature Records: Santa Cruz (Schlising, 1980: 6 [map])
Digitized Records: San Nicolas (1 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Schlising, 1980; Kingsolver, 2004).
Stator Bridwell, 1946
Nomenclatural Authority: Kingsolver (2004)
Notes. Two species of Stator have been recorded from California (Kingsolver, 2004).
Stator limbatus (Horn, 1873)
Nomenclatural Authority: Kingsolver (2004)
Literature Records: Santa Catalina (Cockerell, 1940: 287; Johnson, 1963: 861)
Digitized Records: Santa Catalina (1 iNat)
Range: Also known from mainland (Johnson, 1963; Kingsolver, 2004).
Notes. Recorded as Bruchus limbatus by Cockerell (1940).
Cassidinae: Cassidini
Notes. Three tribes, 13 genera, and 25 species of Cassidinae are known to occur in California (Riley, Clark & Seeno, 2003; M. L. Gimmel, 2022, unpublished data).
Charidotella Weise, 1896
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. One species of Charidotella has been recorded from California (Riley, Clark & Seeno, 2003).
Charidotella sexpunctata (Fabricius, 1781)
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Anacapa (68 LACM; 15 SBMNH), Santa Cruz (5 SBMNH; 1 UCSB; 6 iNat)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Notes. The only subspecies of C. sexpunctata occurring in California is C. s. bicolor (Fabricius, 1798) (Riley, Clark & Seeno, 2003).
Chrysomelinae: Chrysomelini
Notes. Two tribes, 14 genera, and 32 species of Chrysomelinae, of which 12 genera and 30 species belong to Chrysomelini, have been recorded from California (Riley, Clark & Seeno, 2003; M. L. Gimmel, 2022, unpublished data). Wilcox (1972) provided keys to all North American genera and species of Chrysomelinae known at the time.
Calligrapha Chevrolat, 1836
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Six species of Calligrapha have been reported from California (Riley, Clark & Seeno, 2003).
Calligrapha (Calligramma) sigmoidea (LeConte, 1859)
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Gastrophysa Chevrolat, 1836
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. One species of Gastrophysa has been recorded from California (Riley, Clark & Seeno, 2003).
Gastrophysa cyanea Melsheimer, 1847
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: San Nicolas (6 SBMNH), Santa Cruz (8 SBMNH), Santa Rosa (4 LACM)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Phaedon Megerle von Mühlfeld, 1823
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Five species of Phaedon have been recorded from California (Riley, Clark & Seeno, 2003).
Phaedon (Allophaedon) prasinellus (LeConte, 1861)
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Barbara (Miller & Miller, 1985: 131)
Digitized Records: Santa Barbara (1 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Plagiodera Chevrolat, 1836
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. One species of Plagiodera has been recorded from California (Riley, Clark & Seeno, 2003).
Plagiodera (Plagiomorpha) californica (Rogers, 1856)
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Cruz (Cockerell, 1940: 287)
Digitized Records: Santa Cruz (14 LACM; 24 SBMNH)
Range: Also known from mainland (Fall, 1901).
Notes. Recorded as Lina californica by Cockerell (1940).
Trachymela Weise, 1908
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. One species of Trachymela is known from California (Riley, Clark & Seeno, 2003).
Trachymela sloanei (Blackburn, 1896)
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Santa Cruz (2 UCSB; 1 iNat)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Notes. This species was introduced from Australia (Riley, Clark & Seeno, 2003).
Criocerinae: Lemiini
Notes. Two tribes, three genera, and six species of Criocerinae, of which two genera and four species belong to Lemiini, have been recorded from California (Riley, Clark & Seeno, 2003; M. L. Gimmel, 2022, unpublished data).
Lema Fabricius, 1798
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Three species of Lema have been recorded from California (Riley, Clark & Seeno, 2003).
Lema daturaphila Kogan & Goeden, 1970
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Cruz (Fall & Davis, 1934: 144)
Digitized Records: Santa Cruz (74 LACM; 5 SBMNH), Santa Rosa (25 LACM; 2 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Notes. This species was recorded as Lema trilineata var. californica Schaeffer, 1933 by Fall & Davis (1934).
Cryptocephalinae: Cryptocephalini
Notes. Three tribes, 10 genera, and 79 species of Cryptocephalinae, of which three genera and 61 species belong to Cryptocephalini, have been recorded from California (Riley, Clark & Seeno, 2003; M. L. Gimmel, 2022, unpublished data).
Cryptocephalus Geoffroy, 1762
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes: Twelve species of Cryptocephalus have been reported from California (Riley, Clark & Seeno, 2003). The species were revised for North America by White (1968).
Cryptocephalus sanguinicollis Suffrian, 1852
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Santa Catalina (1 SBMNH)
Range: The two subspecies of C. sanguinicollis together span most of western North America. The subspecies reported from the Channel Islands is C. s. nigerrimus Crotch, 1874.
Diachus LeConte, 1880
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Two species of Diachus have been recorded from California (Riley, Clark & Seeno, 2003).
Diachus auratus (Fabricius, 1801)
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: San Clemente (Fall, 1897: 238), San Miguel (Miller & Davis, 1986: 550), Santa Catalina (Fall, 1897: 238), Santa Cruz (Naughton et al., 2014: 303), Santa Rosa (Fall, 1897: 238)
Digitized Records: Anacapa (5 SBMNH), San Clemente (21 SBMNH), San Miguel (19 SBMNH), San Nicolas (9 SBMNH), Santa Catalina (15 SBMNH), Santa Cruz (16 SBMNH), Santa Rosa (4 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Notes. This species was reported from flowers of Malacothrix by Miller & Davis (1986).
Pachybrachis Chevrolat, 1836
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Digitized Records (genus-only): Santa Cruz (1 UCSB)
Notes. Forty-seven species of Pachybrachis have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Pachybrachis melanostictus Suffrian, 1852
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Santa Cruz (16 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Pachybrachis mobilis Fall, 1915
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Santa Catalina (3 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Pachybrachis pluripunctatus Fall, 1915
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Pachybrachis punctatus Bowditch, 1909
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Catalina (Fall, 1915: 343)
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Fall, 1915).
Notes. Fall (1915) reported this species as Pachybrachys punctatus. Earlier, Fall (1897: 238) listed two separate undetermined “Pachybrachys” species from Santa Catalina; this probably represents one of them.
Pachybrachis quadratus Fall, 1915
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Catalina (Fall, 1915: 406)
Digitized Records: none
Range: Also known from mainland (Fall, 1915).
Notes. Fall (1915) reported this species as Pachybrachys punctatus. Earlier, Fall (1897: 238) listed two separate undetermined “Pachybrachys” species from Santa Catalina; this probably represents one of them.
Eumolpinae
Notes. Three tribes, 10 genera, and 36 species of Eumolpinae have been recorded from California (Riley, Clark & Seeno, 2003; M. L. Gimmel, 2022, unpublished data). Straughan & Hadley (1980: 392) recorded “Eumolpinae” from Catalina Harbor, Santa Catalina Island.
Adoxini
Notes. Two genera and four species of Adoxini have been recorded from California (Riley, Clark & Seeno, 2003).
Colaspidea Laporte, 1833
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Three species of Colaspidea have been recorded from California (Riley, Clark & Seeno, 2003).
Colaspidea smaragdula (LeConte, 1857)
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: San Clemente (Fall, 1897: 238; Fall, 1901: 154; Fall, 1933: 232; Miller, 1985a: 21), Santa Catalina (Fall, 1897: 238; Linell, 1898: 481; Fall, 1901: 154; Fall, 1933: 232; Miller, 1985a: 21)
Digitized Records: San Clemente (8 SBMNH), Santa Catalina (2 SBMNH), Santa Cruz (6 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Notes. A supposed endemic, Colaspidea subvittata Fall, 1897, was described from San Clemente and Santa Catalina and reported in the works of Fall (1897, 1901, 1933), Linell (1898), and Miller (1985a). The species was later synonymized with the widespread C. smaragdula by Schultz (1970), reflected in Riley, Clark & Seeno (2003).
Eumolpini
Notes. Five genera and 22 species of Eumolpini have been recorded from California (Riley, Clark & Seeno, 2003; M. L. Gimmel, 2022, unpublished data).
Spintherophyta Dejean, 1836
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Three species of Spintherophyta have been recorded from California (Riley, Clark & Seeno, 2003; Gilbert & Clark, 2020). These were keyed by Gilbert & Clark (2020).
Spintherophyta punctum Gilbert & Clark, 2020
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Rosa (Gilbert & Clark, 2020: 558)
Digitized Records: none
Range: Endemic (Gilbert & Clark, 2020).
Notes. The holotype and paratypes of this species were collected “on leaves of willow” (Gilbert & Clark, 2020).
Galerucinae
Notes. Four tribes, 39 genera, and 199 species have been recorded from California (Riley, Clark & Seeno, 2003; M. L. Gimmel, 2022, unpublished data). Wilcox (1965) provided keys to the non-Alticini taxa for North America.
Alticini
Notes. Twenty-one genera and 114 species of Alticini have been recorded from California (Riley, Clark & Seeno, 2003; M. L. Gimmel, 2022, unpublished data).
Altica Geoffroy, 1762
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Twenty species of Altica have been recorded from California (Riley, Clark & Seeno, 2003).
Altica undetermined species
Literature Records: none
Digitized Records: Anacapa (4 SBMNH), Santa Cruz (1 SBMNH)
Aulacothorax Boheman, 1858
Nomenclatural Authority: Bezděk & Konstantinov (2017).
Notes. One species of Aulacothorax has been recorded from California (Riley, Clark & Seeno, 2003). This genus was, until recently, known as Orthaltica Crotch, 1873 (see Bezděk & Konstantinov, 2017).
Aulacothorax recticollis (LeConte, 1861)
Nomenclatural Authority: Bezděk & Konstantinov (2017)
Literature Records: none
Digitized Records: Santa Catalina (3 SBMNH), Santa Cruz (1 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Notes. This species was until recently known as Orthaltica recticollis (see Bezděk & Konstantinov, 2017).
Dibolia Latreille, 1829
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Three species of Dibolia have been recorded from California (Riley, Clark & Seeno, 2003). The genus was revised for North America by Parry (1974).
Dibolia californica Parry, 1974
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Santa Cruz (4 SBMNH)
Range: Also known from mainland (Parry, 1974).
Disonycha Chevrolat, 1836
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Nine species of Disonycha have been recorded from California (Riley, Clark & Seeno, 2003). Blake (1933) keyed out the species.
Disonycha latiovittata Hatch, 1932
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Santa Cruz (27 LACM), Santa Rosa (11 LACM)
Range: Also known from mainland (Blake, 1933).
Epitrix Foudras, 1859
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Four species of Epitrix have been recorded from California (Riley, Clark & Seeno, 2003). Seeno & Andrews (1972) provided keys to the California species.
Epitrix similaris Gentner, 1944
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Catalina (Gentner, 1944: 142; Seeno & Andrews, 1972: 59 [map])
Digitized Records: none
Range: Also known from mainland (Gentner, 1944; Seeno & Andrews, 1972).
Epitrix subcrinita (LeConte, 1857)
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: Santa Cruz (6 SBMNH)
Range: Also known from mainland (Seeno & Andrews, 1972).
Notes. MLG examined vouchers from the Naughton et al. (2014) study, which were previously identified only to genus; they belong to E. subcrinita.
Longitarsus Berthold, 1827
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Ten species of Longitarsus have been recorded from California (Riley, Clark & Seeno, 2003).
Longitarsus undetermined species 1
Literature Records: none
Digitized Records: San Clemente (1 SBMNH)
Notes. This is a pale, brachypterous species with complete elytra and long antennae, and less than 1.5 mm in total body length (M. L. Gimmel, 2021, personal observation).
Longitarsus undetermined species 2
Literature Records: none
Digitized Records: San Nicolas (2 SBMNH)
Notes. This is a pale, brachypterous species with complete elytra and short antennae, and at least 2.0 mm in total body length (M. L. Gimmel, 2021, personal observation).
Phyllotreta Chevrolat, 1836
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Digitized Records (genus-only): Santa Cruz (1 SBMNH)
Notes. Twenty-one species of Phyllotreta have been recorded from California (Riley, Clark & Seeno, 2003). The specimen from Santa Cruz Island housed in the SBMNH has been tentatively identified as Phyllotreta inconspicua Chittenden, 1927, pending examination of the type of that species (H. Douglas, 2022, personal communication).
Phyllotreta pusilla Horn, 1889
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Catalina (Fall, 1897: 238)
Digitized Records: none
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Galerucini
Notes. Ten genera and 36 species of Galerucini have been recorded from California (Riley, Clark & Seeno, 2003; Viswajyothi & Clark, 2022).
Erynephala Blake, 1936
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Two species of Erynephala have been recorded from California (Riley, Clark & Seeno, 2003).
Erynephala morosa (LeConte, 1857)
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Rosa (Fall, 1897: 238)
Digitized Records: none
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Notes. Recorded as Monoxia puncticollis (Say, 1824) by Fall (1897). Based on information in Blake (1936), this record almost certainly applies to E. morosa and not to the more easterly-occurring species currently known as Erynephala puncticollis (Say, 1824).
Monoxia LeConte, 1865
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Six species of Monoxia have been recorded from California (Riley, Clark & Seeno, 2003; Viswajyothi & Clark, 2022).
Monoxia undetermined species
Literature Records: none
Digitized Records: Anacapa (2 SBMNH)
Notes. This pair of Monoxia from Anacapa Island did not readily match any species presented in Blake (1939) (M. L. Gimmel, 2021, personal observation).
Trirhabda LeConte, 1865
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Digitized Records (genus-only): Santa Cruz (9 UCSB)
Notes. Sixteen species of Trirhabda have been recorded from California (Riley, Clark & Seeno, 2003).
Trirhabda confusa Blake, 1931
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Trirhabda sericotrachyla Blake, 1931
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: none
Digitized Records: Santa Cruz (10 SBMNH), Santa Rosa (6 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003).
Yingabruxia Viswajyothi & Clark, 2022
Nomenclatural Authority: Viswajyothi & Clark (2022)
Notes. Three species of Yingabruxia have been recorded from California (Riley, Clark & Seeno, 2003; Viswajyothi & Clark, 2022).
Yingabruxia sordida (LeConte, 1858)
Nomenclatural Authority: Viswajyothi & Clark (2022)
Literature Records: San Clemente (Miller & Miller, 1985: 130), Santa Barbara (Miller & Miller, 1985: 130)
Digitized Records: San Nicolas (1 SBMNH), Santa Barbara (3 SBMNH)
Range: Also known from mainland (Riley, Clark & Seeno, 2003). Miller & Miller (1985) reported this species as Monoxia sordida; this species was recently transferred to the genus Yingabruxia.
Luperini
Notes. Nine genera and 49 species of Luperini have been recorded from California (Riley, Clark & Seeno, 2003; M. L. Gimmel, 2022, unpublished data).
Diabrotica Chevrolat, 1836
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Two species of Diabrotica have been recorded from California (Riley, Clark & Seeno, 2003).
Diabrotica undecimpunctata Mannerheim, 1843
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Barbara (Miller & Miller, 1985: 130), Santa Catalina (Seavey, 1892: 263; Fall, 1897: 238), Santa Cruz (Smith, 1966: 109 [map]; Miller & Miller, 1985: 130)
Digitized Records: Santa Barbara (1 SBMNH), Santa Cruz (8 LACM; 3 SBMNH; 6 UCSB), Santa Rosa (7 LACM)
Range: Also known from mainland (Smith, 1966; Riley, Clark & Seeno, 2003).
Notes. This species was recorded as Diabrotica soror LeConte, 1865 by Seavey (1892) and Fall (1897).
Scelolyperus Crotch, 1874
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Notes. Twenty species of Scelolyperus have been recorded from California (Riley, Clark & Seeno, 2003). Clark (1996) revised the genus for North America.
Scelolyperus torquatus (LeConte, 1884)
Nomenclatural Authority: Riley, Clark & Seeno (2003)
Literature Records: Santa Catalina (Wilcox, 1965: 140)
Digitized Records: none
Range: Also known from mainland (Wilcox, 1965; Clark, 1996; Riley, Clark & Seeno, 2003).
CURCULIONOIDEA
Attelabidae
Notes. One subfamily (Rhynchitinae), six genera, and 18 species of Attelabidae have been recorded from California (O’Brien & Wibmer, 1982).
Deporaus Samouelle, 1819
Nomenclatural Authority: O’Brien & Wibmer (1982)
Notes. One species of Deporaus has been recorded from California (O’Brien & Wibmer, 1982).
Deporaus glastinus (LeConte, 1857)
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Cruz (Hamilton, 1969: 394)
Digitized Records: Santa Catalina (9 LACM), Santa Cruz (33 LACM; 17 SBMNH)
Range: Also known from mainland (O’Brien & Wibmer, 1982; Hamilton, 1969, 2002).
Temnocerus Thunberg, 1815
Nomenclatural Authority: Hamilton (2002)
Notes. Six species of Temnocerus have been recorded from California (O’Brien & Wibmer, 1982). This genus was until recently known as Pselaphorhynchites Schilsky, 1903.
Temnocerus aeratoides (Fall, 1901)
Nomenclatural Authority: Hamilton (1971), Hamilton (2002)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Hamilton, 1971).
Notes. Hamilton (1971) recorded this species as Pselaphorhynchites aeratoides.
Temnocerus aureus (LeConte, 1876)
Nomenclatural Authority: Hamilton (1971), Hamilton (2002)
Literature Records: San Clemente (Fall, 1897: 239; Fall, 1901: 186)
Digitized Records: Santa Cruz (21 SBMNH)
Range: Also known from mainland (Fall, 1901).
Notes. Fall (1897, 1901) recorded this species as Rhynchites aureus, and Hamilton (1971) as Pselaphorhynchites aureus.
Temnocerus insularis (Fall, 1929)
Nomenclatural Authority: Hamilton (1971), Hamilton (2002)
Literature Records: San Clemente (Fall, 1929b: 294; Hamilton, 1969: 211), Santa Catalina (Fall, 1897: 239; Fall, 1929b: 294; Hamilton, 1969: 211; Hamilton, 1971: 986)
Digitized Records: none
Range: Also known from mainland (Hamilton, 1969, 1971).
Notes. Fall (1897: 239) reported this species as “Rhynchites sp. nov.?”; Fall (1929b) recorded this species as Rhynchites insularis, and Hamilton (1969, 1971) recorded it as Pselaphorhynchites insularis. The species was thought to be endemic at the time of its description (Fall, 1929b).
Temnocerus naso (Casey, 1885)
Nomenclatural Authority: Hamilton (1971), Hamilton (2002)
Literature Records: none
Digitized Records: Santa Cruz (1 UCRC)
Range: Also known from mainland (Hamilton, 1971).
Notes. Hamilton (1971) recorded this species as Pselaphorhynchites naso. This species is distributed through much of mainland southern California (Hamilton, 1971).
Brentidae
Notes. One subfamily, 13 genera, and 35 species of Brentidae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Apioninae
Literature Records (subfamily-only): Santa Cruz (Naughton et al., 2014: 303)
Digitized Records (subfamily-only): San Clemente (3 SBMNH), San Miguel (1 SBMNH), Santa Cruz (51 SBMNH), Santa Rosa (3 SBMNH)
Notes. This subfamily is fairly diverse in California, with 13 genera and 35 species recorded (O’Brien & Wibmer, 1982; M. L. Gimmel, 2022, unpublished data). Although they were monographed for North America by Kissinger (1968), they are exceedingly challenging to identify. Because of this challenge, we have included a “Digitized Records” section above for this subfamily to indicate the amount of material in dire need of expert determination.
Coelocephalapion Wagner, 1914
Nomenclatural Authority: Anderson & Kissinger (2002)
Notes. Six species of Coelocephalapion have been recorded from California (O’Brien & Wibmer, 1982).
Coelocephalapion antennatum (Smith, 1884)
Nomenclatural Authority: Kissinger (1968)
Literature Records: Santa Catalina (Fall, 1897: 239), Santa Cruz (Kissinger, 1968: 249)
Digitized Records: none
Range: Also known from mainland (Kissinger, 1968).
Notes. Recorded as Apion antennatum by Fall (1897) and Kissinger (1968), with the latter work not including the species in a subgenus. A new classification of former Apion Herbst, 1797 was developed by Alonso-Zarazaga (1990), but not fully implemented for North American species (Anderson & Kissinger, 2002); however, this species has been informally moved to the genus Coelocephalapion in, e.g., bugguide.net.
Coelocephalapion californicum (Smith, 1884)
Nomenclatural Authority: Kissinger (1968)
Literature Records: Santa Cruz (Cockerell, 1940: 287)
Digitized Records: none
Range: Also known from mainland (Kissinger, 1968).
Notes. Reported by Cockerell (1940) as Apion californicum. Kissinger (1968) included this species in the subgenus Apion (Coelocephalapion), which was upgraded to genus by Kissinger (1992).
Coelocephalapion oedorhynchum (LeConte, 1858)
Nomenclatural Authority: Kissinger (1968)
Literature Records: Santa Catalina (Fall, 1897: 239; Fall, 1898a: 130)
Digitized Records: none
Range: Also known from mainland (Fall, 1898a; Kissinger, 1968).
Notes. Recorded as Apion oedorhynchum by Fall (1897, 1898a) and Kissinger (1968), with the latter work not including the species in a subgenus. A new classification of former Apion was developed by Alonso-Zarazaga (1990), but not fully implemented for North American species (Anderson & Kissinger, 2002); however, this species has been informally moved to the genus Coelocephalapion in, e.g., bugguide.net.
Curculionidae
Notes. Nineteen subfamilies, 221 genera, and 821 species of Curculionidae are known to occur in California (M. L. Gimmel, 2022, unpublished data). Subfamilies occurring on the nearby mainland but not known from the Channel Islands include: Bagoinae, Conoderinae, Cryptorhynchinae, Gonipterinae, Mesoptiliinae, Platypodinae.
Baridinae: Baridini
Notes. Three tribes, 15 genera, and 48 species of Baridinae, including seven genera and 24 species of Baridini, are known to occur in California (O’Brien & Wibmer, 1982; M. L. Gimmel, 2022, unpublished data).
Trichobaris LeConte, 1876
Nomenclatural Authority: Anderson (2002)
Digitized Records (genus-only): Santa Cruz (2 UCSB)
Notes. Three species of Trichobaris have been recorded from California (O’Brien & Wibmer, 1982).
Trichobaris compacta Casey, 1892
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: Santa Cruz (5 SBMNH)
Range: Also known from mainland (O’Brien & Wibmer, 1982).
Ceutorhynchinae: Ceutorhynchini
Notes. Five tribes, 14 genera, and 42 species of Ceutorhynchinae, of which six genera and 31 species belong to Ceutorhynchini, are known to occur in California (O’Brien & Wibmer, 1982; M. L. Gimmel, 2022, unpublished data).
Ceutorhynchus Germar, 1824
Nomenclatural Authority: Anderson (2002)
Notes. Twenty-four species of Ceutorhynchus are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Ceutorhynchus assimilis (Paykull, 1792)
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (O’Brien & Wibmer, 1982).
Notes. This species is introduced from Europe (O’Brien & Wibmer, 1982).
Cossoninae
Notes. Five tribes, 12 genera, and 29 species of Cossoninae are known to occur in California (O’Brien & Wibmer, 1982; M. L. Gimmel, 2022, unpublished data).
Onycholipini
Notes. Two genera and two species of Onycholipini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Pselactus Broun, 1886
Nomenclatural Authority: Anderson (2002)
Notes. A single, adventive species of Pselactus is known from North America (Anderson, 2002).
Pselactus spadix (Herbst, 1795)
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: Santa Rosa (2 SBMNH)
Range: Also known from mainland (O’Brien & Wibmer, 1982).
Notes. This beach-dwelling and driftwood-inhabiting species is introduced to North America (Anderson, 2002).
Rhyncolini
Notes. Five genera and 14 species of Rhyncolini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Elassoptes Horn, 1873
Nomenclatural Authority: Anderson (2002)
Notes. A single species of Elassoptes is known from North America (Anderson, 2002).
Elassoptes marinus Horn, 1873
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: San Clemente (2 SBMNH), San Miguel (26 SBMNH), San Nicolas (5 SBMNH), Santa Cruz (16 SBMNH), Santa Rosa (21 SBMNH)
Range: Also known from mainland (O’Brien & Wibmer, 1982).
Notes. This species is a beach inhabitant associated with driftwood (Anderson, 2002).
Rhyncolus Germar, 1817
Nomenclatural Authority: Anderson (2002)
Literature Records (genus-only): Santa Barbara (Miller & Miller, 1985: 131)
Digitized Records (genus-only): San Miguel (4 SBMNH)
Notes. Miller & Miller (1985) recorded the genus only, noting that it needs revision. Nine nominal species of Rhyncolus are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Rhyncolus cylindricollis Wollaston, 1873
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: Santa Cruz (6 SBMNH)
Range: Also known from mainland (O’Brien & Wibmer, 1982).
Curculioninae
Notes. Nine tribes, 23 genera, and 131 species of Curculioninae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Anthonomini
Notes. Six genera and 60 species of Anthonomini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Anthonomus Germar, 1817
Nomenclatural Authority: Anderson (2002)
Digitized Records (genus-only): Anacapa (2 LACM; 2 SBMNH), Santa Catalina (5 SBMNH)
Notes. Forty-five species of Anthonomus belonging to five subgenera (Anthomorphus Weise, 1883, Anthonomochaeta Dietz, 1891, Anthonomorphus Dietz, 1891, Anthonomus (s.str.), and Cnemocyllus Dietz, 1891) are known from California (M. L. Gimmel, 2022, unpublished data). The subgenus Cnemocyllus was revised by Clark & Burke (2005). All SBMNH genus-only records belong to the subgenus Cnemocyllus.
Anthonomus (Anthonomus) pauperculus LeConte, 1876
Nomenclatural Authority: Clark et al. (2019)
Literature Records: Santa Catalina (Seavey, 1892: 262; Fall, 1897: 239; Cockerell, 1940: 287; Clark et al., 2019: 796)
Digitized Records: none
Range: Also known from mainland (Clark et al., 2019).
Notes. The record of Anthonomus canus LeConte, 1876 (now a junior synonym of Anthonomus (Cnemocyllus) decipiens LeConte, 1876) by Seavey (1892) was considered to be erroneous according to Fall (1897: 235) and refers to A. pauperculus (see Cockerell, 1940: 287). Members of the Anthonomus squamosus LeConte, 1876 species-group, to which A. pauperculus belongs, were revised by Clark et al. (2019).
Anthonomus (Cnemocyllus) inermis Boheman, 1859
Nomenclatural Authority: Clark & Burke (2005)
Literature Records: San Miguel (Miller & Miller, 1985: 131), Santa Barbara (Miller & Miller, 1985: 131), Santa Rosa (Clark & Burke, 2005: 45)
Digitized Records: Santa Barbara (1 LACM)
Range: Also known from mainland (Clark & Burke, 2005).
Notes. Recorded as Anthonomus subvittatus LeConte, 1876 by Miller & Miller (1985), which is a junior synonym of A. inermis (see Clark & Burke, 2005). Miller & Miller (1985) reported it from Hemizonia clementina (Asteraceae) on Santa Barbara Island.
Anthonomus (Cnemocyllus) undescribed species
Literature Records: none
Digitized Records: Santa Barbara (2 SBMNH)
Range: ?Endemic.
Notes. These seven SBMNH specimens (on two pins) from Santa Barbara Island, one of which is a dissected male, were marked as “Anthonomus n. sp. #2, Cnemocyllus gp.” by Horace R. Burke in 2009. Based on the funicle with seven antennomeres, it belongs to the A. inermis group of Clark & Burke (2005) (M. L. Gimmel, 2021, personal observation). It is unknown whether specimens of this morphospecies exist from other localities.
Curculionini
Notes. One genus and three species of Curculionini have been recorded from California (Gibson, 1969).
Curculio Linnaeus, 1758
Nomenclatural Authority: Anderson (2002)
Literature Records (genus-only): Santa Cruz (Naughton et al., 2014: 303)
Digitized Records (genus-only): Santa Catalina (1 SBMNH), Santa Cruz (1 SBMNH)
Notes. Three species of Curculio have been recorded from California (Gibson, 1969). The Naughton et al. (2014) record refers to a single specimen of “Curculio sp.” in addition to records of C. uniformis. Seavey (1892: 262) recorded Balaninus obtusus [=Curculio obtusus (Blanchard, 1884)] from Santa Catalina Island. Gibson (1969) used an unnecessary replacement name, Curculio neocorylus Gibson, 1969 to refer to this species. Fall (1897: 235) doubted the validity of this identification; we agree that this record must be in error, since this species has not subsequently been reported from California and is otherwise only known from east of the Rocky Mountains (Gibson, 1969). This record almost certainly represents one of the two species of Curcuilo listed below. The North American species of Curculio were revised by Gibson (1969).
Curculio aurivestis Chittenden, 1927
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Catalina (Chittenden, 1927: 186, 191)
Digitized Records: Santa Catalina (1 USNM)
Range: Also known from mainland (Gibson, 1969).
Notes. Chittenden (1927) recognized two species, one from the mainland (C. aurivestis) and one insular and presumed endemic (Curculio brevinasus Chittenden, 1927), the latter representing the Santa Catalina record above. Gibson (1969) synonymized the two and recognized a single species ranging from British Columbia to southern California where it breeds in various species of Quercus.
Curculio uniformis (LeConte, 1857)
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Catalina (Fall, 1897: 239; Fall, 1901: 199; Chittenden, 1908: 22), Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: Santa Cruz (4 SBMNH)
Range: Also known from mainland (Fall, 1901; Chittenden, 1908; Gibson, 1969).
Notes. Fall (1897) recorded this species as Balaninus occidentis Casey, 1897 and noted this species had previously been confused with Balaninus uniformis, but is distinct. Fall (1901) recorded this species as B. uniformis. Chittenden (1908) synonymized B. occidentis with B. uniformis, and Gibson (1969) incorrectly used C. occidentis as the valid name.
Smicronychini
Notes. Two genera and 21 species of Smicronychini have been recorded from California (O’Brien & Anderson, 1996).
Smicronyx Schoenherr, 1843
Nomenclatural Authority: Anderson (2002)
Literature Records (genus-only): Santa Rosa (Fall, 1897: 239)
Digitized Records (genus-only): San Clemente (1 SBMNH)
Notes. Nineteen species of Smicronyx have been recorded from California (O’Brien & Anderson, 1996). Fall (1897) recorded only “Smicronyx, sp.” from Santa Rosa Island. Anderson (1962) revised the species for North America.
Smicronyx cinereus (Motschulsky, 1845)
Nomenclatural Authority: O’Brien & Anderson (1996)
Literature Records: Santa Rosa (Anderson, 1962: 208)
Digitized Records: none
Range: Also known from mainland (Anderson, 1962).
Notes. The record from Fall (1897) may refer to this species, but because there is at least one new island record for the genus (San Clemente) the genus should be reexamined from the Channel Islands.
Tychiini
Notes. Three genera and 14 species of Tychiini have been recorded from California (O’Brien & Wibmer, 1982).
Sibinia Germar, 1817
Nomenclatural Authority: Anderson (2002)
Notes. Five species of Sibinia are known to occur in California (O’Brien & Wibmer, 1982). This genus was revised for the New World by Clark (1978).
Sibinia maculata (LeConte, 1876)
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: San Miguel (Clark, 1978: 363; Miller & Miller, 1985: 131), San Nicolas (Fall, 1901: 197), Santa Barbara (Miller & Miller, 1985: 131)
Digitized Records: Santa Barbara (1 LACM)
Range: Also known from mainland (Fall, 1901; Clark, 1978).
Notes. Fall (1901) recorded this species as Paragoges maculatus LeConte. Reported from “sage brush” on Santa Barbara Island by Miller & Miller (1985).
Tychius Germar, 1817
Nomenclatural Authority: Anderson (2002)
Literature Records (genus-only): San Nicolas (Fall, 1897: 239)
Notes. Six species of Tychius have been recorded from California (O’Brien & Wibmer, 1982). The genus was revised for North America by Clark (1971). The status of the San Nicolas Island record referred to by Fall (1897) as “Tychius, n. sp.” is unknown.
Tychius lineellus LeConte, 1876
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Cruz (Clark, 1971: 18), Santa Rosa (Clark, 1971: 18)
Digitized Records: Santa Cruz (4 LACM), Santa Rosa (5 LACM; 2 SBMNH)
Range: Also known from mainland (Clark, 1971).
Cyclominae
Notes. Two genera and 17 species of Cyclominae have been recorded from California (O’Brien, 1997).
Listroderes Schoenherr, 1826
Nomenclatural Authority: Anderson (2002)
Digitized Records (genus-only): Santa Cruz (1 UCSB)
Notes. Two introduced species of Listroderes have been recorded from California (O’Brien, 1997). This genus was partially revised by Morrone (1993).
Listroderes costirostris Schoenherr, 1826
Nomenclatural Authority: Morrone (1993)
Literature Records: none
Digitized Records: San Miguel (14 SBMNH), San Nicolas (11 LACM), Santa Catalina (1 LACM), Santa Rosa (1 LACM)
Range: Also known from mainland (Morrone, 1993).
Notes. This species was accidentally introduced to North America from South America (Morrone, 1993).
Listronotus Jekel, 1865
Nomenclatural Authority: Anderson (2002)
Digitized Records (genus-only): Santa Catalina (3 SBMNH)
Notes. Fifteen species of Listronotus have been recorded from California (O’Brien, 1997). The larger species of this genus in North America were taxonomically treated by O’Brien (1981).
Listronotus sordidus (Gyllenhal, 1834)
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: San Nicolas (Cockerell, 1940: 287)
Digitized Records: none
Range: Also known from mainland (O’Brien & Wibmer, 1982).
Notes. This species was reported by Cockerell (1940) as Listronotus obliquus LeConte, 1876. This species is now considered a junior synonym of L. sordidus (see O’Brien & Wibmer, 1982: 70), but this species’ reported range does not include California (O’Brien & Wibmer, 1982). The specific identity of the San Nicolas Island record is therefore still in question.
Dryophthorinae: Rhynchophorini
Notes. Two tribes, seven genera, and 27 species of Dryophthorinae, of which all but one genus and species belong to Rhynchophorini, have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Scyphophorus Schoenherr, 1838
Nomenclatural Authority: Anderson (2002)
Notes. Two species of Scyphophorus have been recorded from California (Vaurie, 1971). The species of the genus were treated by Vaurie (1971).
Scyphophorus yuccae Horn, 1873
Nomenclatural Authority: Vaurie (1971), O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: Santa Cruz (2 SBMNH)
Range: Also known from mainland (Vaurie, 1971; O’Brien & Wibmer, 1982).
Notes. This species breeds in the stems of Hesperoyucca whipplei (Torr.) Trel. (Agavaceae), a common plant on the mainland that does not occur natively in the Channel Islands. However, a variety of yucca species have been planted there.
Sphenophorus Schoenherr, 1838
Nomenclatural Authority: Anderson (2002)
Notes. Eighteen species of Sphenophorus have been recorded from California (M. L. Gimmel, 2022, unpublished data). This genus was treated for North America by Vaurie (1951) as the genus Calendra Clairville & Schellenberg, 1798.
Sphenophorus graminis Chittenden, 1905
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Vaurie, 1951; O’Brien & Wibmer, 1982).
Sphenophorus phoeniciensis Chittenden, 1904
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Vaurie, 1951; O’Brien & Wibmer, 1982).
Sphenophorus simplex LeConte, 1860
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: San Nicolas (1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Vaurie, 1951; O’Brien & Wibmer, 1982).
Sphenophorus vomerinus LeConte, 1858
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Rosa (Fall, 1897: 239)
Digitized Records: none
Range: Also known from mainland (Vaurie, 1951; O’Brien & Wibmer, 1982).
Entiminae
Notes. Fifteen tribes, 53 genera, and 205 species of Entiminae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Geonemini
Notes. Four genera and 64 species of Geonemini are known to occur in California (O’Brien & Wibmer, 1982).
Trigonoscuta Motschulsky, 1853
Nomenclatural Authority: Anderson (2002)
Literature Records (genus-only): San Clemente (Doyen, 1974: 87), San Miguel (Van Dam & Matzke, 2016: 1527), San Nicolas (Van Dam & Matzke, 2016: 1527), Santa Catalina (Van Dam & Matzke, 2016: 1527), Santa Cruz (Van Dam & Matzke, 2016: 1527), Santa Rosa (Van Dam & Matzke, 2016: 1527)
Digitized Records (genus-only): San Clemente (2 LACM; 7 SBMNH), San Miguel (14 SBMNH), San Nicolas (32 LACM; 24 SBMNH), Santa Catalina (9 LACM), Santa Cruz (5 SBMNH), Santa Rosa (25 SBMNH)
Notes. This genus was revised in elaborate detail by Pierce (1975), who diagnosed and named minute variation in these sand-dwelling beetles, recognizing over 150 species and subspecies, most of which were limited to a single locality or even collecting event. Miller (1985a: 21), in a list of endemic species of the Channel Islands, did not bother to individually list all of the purported endemic Trigonoscuta described by Pierce (1975). A robust molecular phylogeny was published by Van Dam & Matzke (2016) which sampled specimens from five of the Channel Islands. This phylogeny seems to indicate that at least two species are present on the islands and each of those are found on multiple islands. This contradicts Pierce (1975) who enumerated different species or subspecies for nearly every beach locality collected on the islands and mainland California. It seems likely that coastal dune Trigonoscuta may have similar diversity to the tenebrionid genus Coelus which has a nearly identical distribution and habitat (see account above for this genus). It actually may be the case that the two “subgenera” identified by Pierce as inhabiting the islands align more appropriately with true species diversity than the numerous “species” described. This genus is in great need of a modern revision and likely tells a very interesting story of dune colonization between the islands and mainland California. Van Dam & Matzke (2016, see their fig. 5 and supplemental Information) found two well supported clades which one might consider species: one from the northern islands of Santa Rosa, Santa Cruz, and San Miguel sister to a specimen from the mainland, and a second from Santa Catalina and San Nicolas islands. Specimens of this genus are known from all eight Channel Islands. The records presented under each species largely reflect the work of Pierce (1975) and the specimens included within that study.
Trigonoscuta anacapensis Pierce, 1975
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Anacapa (Pierce, 1975: 48)
Digitized Records: Anacapa (18 LACM)
Range: Endemic (Pierce, 1975).
Notes. Described in the subgenus Trigonoscuta (s.str.), which Pierce (1975) understood as otherwise restricted to the mainland Pacific Coast. This species was described from West Anacapa Island and considered similar to populations described from the coast of Ventura County, California (Pierce, 1975: 49).
Trigonoscuta catalina Pierce, 1975
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Catalina (Pierce, 1975: 53)
Digitized Records: Santa Catalina (24 LACM)
Range: Endemic (Pierce, 1975).
Notes. Described in the subgenus Nesocatoecus Pierce, 1975, which Pierce (1975) understood as restricted to the Channel Islands.
Trigonoscuta clemente Pierce, 1975
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: San Clemente (Pierce, 1975: 53)
Digitized Records: San Clemente (2031 LACM), Santa Barbara (2 LACM)
Range: Endemic (Pierce, 1975).
Notes. The correct spelling of this species is T. clemente (Pierce, 1975), though in the same paper the incorrect original spelling of T. sanclemente was also used (nec. T. sanclementis Pierce, 1975; see O’Brien & Wibmer, 1982: 33). Described in the subgenus Nesocatoecus, which Pierce (1975) understood as restricted to the Channel Islands. Pierce (1975) described five subspecies largely based off of dune host plants the beetles were collected from, which seem highly unlikely to represent different taxa in an evolutionary context. The subspecies are: T. c. clemente Pierce, 1975; T. c. isola Pierce, 1975; T. c. excavata Pierce, 1975; T c. latesecula Pierce, 1975; T. c. traskiae Pierce, 1975.
Trigonoscuta curviscroba Pierce, 1975
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Barbara (Pierce, 1975: 53; Miller & Miller, 1985: 131)
Digitized Records: none
Range: Endemic (Pierce, 1975).
Notes. Described in the subgenus Nesocatoecus, which Pierce (1975) understood as restricted to the Channel Islands. Miller & Miller (1985) merely listed T. curviscroba as being described from Santa Barbara Island, and doubted the taxonomic validity of this species.
Trigonoscuta miguelensis Pierce, 1975
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: San Miguel (Pierce, 1975: 46)
Digitized Records: San Miguel (21 LACM; 10 SBMNH)
Range: Endemic (Pierce, 1975).
Notes. Described in the subgenus Nesocatoecus, which Pierce (1975) understood as restricted to the Channel Islands.
Trigonoscuta nesiotis Pierce, 1975
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Anacapa (Pierce, 1975: 48)
Digitized Records: Anacapa (35 LACM)
Range: Endemic (Pierce, 1975).
Notes. Described in the subgenus Nesocatoecus, which Pierce (1975) understood as restricted to the Channel Islands. This species was described from West Anacapa Island.
Trigonoscuta nicolana Pierce, 1975
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: San Nicolas (Pierce, 1975: 49)
Digitized Records: San Nicolas (264 LACM), Santa Barbara (2 LACM)
Range: Endemic (Pierce, 1975).
Notes. Described in the subgenus Nesocatoecus, which Pierce (1975) understood as restricted to the Channel Islands. Eight subspecies were described by Pierce (1975), which corresponded to the dune host plant the beetles were found associated with. It seems highly unlikely that these names represent different taxa in an evolutionary context. The subspecies are: T. n. nicolana Pierce, 1975; T. n. longinoda Pierce, 1975; T. n. latelobata Pierce, 1975; T. n. nonmarginata Pierce, 1975; T. n. latespiculum Pierce, 1975; T. n. lateconjuncta Pierce, 1975; T. n. sulcata Pierce, 1975; T. n. breviconjuncta Pierce, 1975.
Trigonoscuta pilosa Motschulsky, 1953
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: San Clemente (Fall, 1897: 239), Santa Rosa (Fall, 1897: 239)
Digitized Records: none
Range: Also known from mainland (Pierce, 1975).
Notes. In the genus revision by Pierce (1975), this species was interpreted to occur from Washington, Oregon, and California north of the San Andreas fault, from which were described eight subspecies. The literature records for this taxon predate Pierce’s revision and likely correspond to other taxa named therein. This species belongs to the subgenus Trigonoscuta (s.str.), which Pierce (1975) recognized as restricted to the mainland except for T. anacapensis from Anacapa island.
Trigonoscuta sanctabarbarae Pierce, 1975
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Barbara (Pierce, 1975: 52; Miller & Miller, 1985: 131).
Digitized Records: Santa Barbara (46 LACM)
Range: Endemic (Pierce, 1975).
Notes. The correct spelling of this name is T. sanctabarbarae, but in the same paper (Pierce, 1975) the incorrect original spelling T. santabarbarae was also used (see O’Brien & Wibmer, 1982: 35). Described in the subgenus Nesocatoecus, which Pierce (1975) understood as restricted to the Channel Islands. Pierce (1975) included three subspecies, all from Santa Barbara Island: T. s. sanctabarbarae Pierce, 1975; T. s. mesembryanthemi Pierce, 1975; T. s. lycii Pierce, 1975. Miller & Miller (1985) merely listed these three subspecies (as “Trigonoscuta santabarbarae”) as being described from Santa Barbara Island. They doubted the taxonomic validity of these taxa.
Trigonoscuta sanctarosae Pierce, 1975
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Rosa (Pierce, 1975: 47, 48)
Digitized Records: Santa Rosa (138 LACM)
Range: Endemic (Pierce, 1975).
Notes. Described in the subgenus Nesocatoecus, which Pierce (1975) understood as restricted to the Channel Islands. Pierce (1975: 47–48) described two subspecies, both from Santa Rosa Island: T. s. sanctarosae Pierce, 1975 and T. s. astragalensis Pierce, 1975.
Trigonoscuta stantoni Sleeper, 1975
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Cruz (Pierce, 1975: 77)
Digitized Records: Santa Cruz (35 SBMNH; 1 iNat)
Range: Endemic (Pierce, 1975).
Notes. Described in the subgenus Nesocatoecus, which Pierce (1975) understood as restricted to the Channel Islands. This species was described by the editor of the revision (Pierce, 1975) to fill in an island gap that Pierce had postulated should have a species present.
Naupactini
Notes. Four genera and seven species of Naupactini have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Naupactus Dejean, 1821
Nomenclatural Authority: Anderson (2002)
Notes. One species of Naupactus is known from California (M. L. Gimmel, 2022, unpublished data).
Naupactus cervinus Boheman, 1840
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: Santa Catalina (Cockerell, 1940: 287)
Digitized Records: Santa Catalina (1 LACM; 1 SBMNH; 1 iNat), Santa Cruz (89 LACM)
Range: Also known from mainland (O’Brien & Wibmer, 1982).
Notes. Recorded as Pantomorus fulleri (Horn) by Cockerell (1940), and often referred to as Pantomorus cervinus in the literature. This species is introduced in North America (O’Brien & Wibmer, 1982).
Otiorhynchini
Notes. Three genera and 13 species of Otiorhynchini have been recorded from California (O’Brien & Wibmer, 1982).
Otiorhynchus Germar, 1822
Nomenclatural Authority: Anderson (2002)
Notes. Five species of Otiorhynchus have been recorded from California (O’Brien & Wibmer, 1982). A key and distributional summary of the genus in North America was provided by Warner & Negley (1976).
Otiorhynchus cribricollis Gyllenhal, 1834
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: San Nicolas (1 SBMNH)
Range: Also known from mainland (Warner & Negley, 1976; O’Brien & Wibmer, 1982).
Notes. This species is adventive in North America from Europe (Warner & Negley, 1976).
Sciopithes Horn, 1876
Nomenclatural Authority: Anderson (2002)
Notes. Six species of Sciopithes have been recorded from California (O’Brien & Wibmer, 1982).
Sciopithes insularis Van Dyke, 1935
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: San Clemente (Van Dyke, 1935: 91; Miller, 1985a: 21; Miller & Miller, 1985: 131)
Digitized Records: none
Range: Endemic (Van Dyke, 1935; Miller, 1985a).
Sciopithes setosus Casey, 1888
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: San Clemente (Fall, 1897: 239; Fall, 1901: 188), Santa Barbara (Miller & Miller, 1985: 131)
Digitized Records: none
Range: Also known from mainland (Fall, 1901).
Notes. Fall (1897) recorded this species as “var.”. Miller & Miller (1985) reported this species from Coreopsis gigantea on Santa Barbara Island.
Peritelini
Notes. Sixteen genera and 37 species of Peritelini have been recorded from California (O’Brien & Wibmer, 1982).
Geodercodes Casey, 1888
Nomenclatural Authority: Anderson (2002)
Notes. One species of Geodercodes occurs in North America (Anderson, 2002).
Geodercodes latipennis Casey, 1888
Nomenclatural Authority: O’Brien & Wibmer (1982), Anderson (2002)
Literature Records: Santa Cruz (Polihronakis, Caterino & Chatzimanolis, 2010: 940; Naughton et al., 2014: 303), Santa Rosa (Polihronakis, Caterino & Chatzimanolis, 2010: 940)
Digitized Records: San Clemente (8 SBMNH), Santa Cruz (11 SBMNH), Santa Rosa (16 SBMNH)
Range: Also known from mainland (O’Brien & Wibmer, 1982; Anderson, 2002; Polihronakis, Caterino & Chatzimanolis, 2010).
Notes. Polihronakis, Caterino & Chatzimanolis (2010) showed that this species is made up of sexual and asexual populations across the Coast and Transverse ranges of California; the populations on the northern Channel Islands are asexual.
Nemocestes Van Dyke, 1936
Nomenclatural Authority: Anderson (2002)
Notes. This genus contains nine species in California (O’Brien & Wibmer, 1982, as Geoderces Horn, 1876; M. L. Gimmel, 2022, unpublished data). Eight of these species were keyed by Van Dyke (1936).
Nemocestes undetermined species
Literature Records: Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: Santa Catalina (8 SBMNH), Santa Cruz (6 SBMNH), Santa Rosa (3 SBMNH)
Peritelinus Casey, 1888
Nomenclatural Authority: Anderson (2002)
Notes. This genus contains two species in California (O’Brien & Wibmer, 1982). These were distinguished by Van Dyke (1936).
Peritelinus undetermined species
Literature Records: none
Digitized Records: Anacapa (6 SBMNH)
Stenoptochus Casey, 1888
Nomenclatural Authority: O’Brien & Wibmer (1982); Anderson (2002)
Notes. This genus contains two species in California (O’Brien & Wibmer, 1982; Anderson, 2002). These were distinguished by Van Dyke (1935).
Stenoptochus undetermined species
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Sitonini
Notes. One genus and nine species of Sitonini have been recorded from California (Bright, 1994).
Sitona Germar, 1817
Nomenclatural Authority: Bright (1994)
Notes. This genus contains nine described species in California (Bright, 1994). These were revised for North America by Bright (1994).
Sitona californius Fåhraeus, 1840
Nomenclatural Authority: Bright (1994)
Literature Records: San Miguel (Blaisdell, 1938: 33; Miller, 1985a: 21; Bright, 1994: 294 [map]), Santa Cruz (Bright, 1994: 294 [map])
Digitized Records: Anacapa (16 LACM), San Miguel (10 LACM; 1 SBMNH), Santa Catalina (2 SBMNH), Santa Cruz (5 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Bright, 1994).
Notes. This species was described and recorded by Blaisdell (1938) as the purported endemic Sitona cockerelli Blaisdell, 1938, listed as such by Miller (1985a: 21). Bright (1989: 77), however, synonymized this species with S. californicus, later recognized with the spelling S. californius.
Erirhininae
Notes. Six genera and 10 species of Erirhininae have been recorded from California (O’Brien & Anderson, 1996; M. L. Gimmel, 2022, unpublished data).
Notiodes Schoenherr, 1838
Nomenclatural Authority: Anderson (2002)
Notes. Five species of Notiodes have been recorded from California (O’Brien & Anderson, 1996).
Notiodes aeratus (LeConte, 1876)
Nomenclatural Authority: O’Brien & Anderson (1996)
Literature Records: none
Digitized Records: San Clemente (1 SBMNH)
Range: Also known from mainland (O’Brien & Anderson, 1996).
Hyperinae
Notes. Two genera and five species of Hyperinae have been recorded from California (O’Brien & Wibmer, 1982; M. L. Gimmel, 2022, unpublished data).
Hypera Germar, 1817
Nomenclatural Authority: Anderson (2002)
Notes. Four introduced species of Hypera have been recorded from California (O’Brien & Wibmer, 1982).
Hypera postica (Gyllenhal, 1813)
Nomenclatural Authority: O’Brien & Wibmer (1982)
Literature Records: none
Digitized Records: San Clemente (2 SBMNH), San Nicolas (1 SBMNH), Santa Cruz (16 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (O’Brien & Wibmer, 1982).
Notes. This species is introduced in North America (O’Brien & Wibmer, 1982).
Lixinae
Notes. Three tribes, seven genera, and 30 species of Lixinae are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Cleonini
Notes. Two genera and 17 species of Cleonini have been recorded from California (Anderson, 1988). The species were revised for the New World by Anderson (1988).
Apleurus Chevrolat, 1873
Nomenclatural Authority: Anderson (2002)
Notes. Six species of Apleurus are known to occur in California (Anderson, 1988).
Apleurus (Apleurus) jacobinus (Casey, 1891)
Nomenclatural Authority: Anderson (1988)
Literature Records: San Miguel (Anderson, 1988: 646 [map])
Digitized Records: San Miguel (1 SBMNH)
Range: Also known from mainland (Anderson, 1988).
Scaphomorphus Motschulsky, 1860
Nomenclatural Authority: Anderson (2002)
Notes. This genus contains 12 species in California (Anderson, 1988).
Scaphomorphus americanus (Csiki, 1934)
Nomenclatural Authority: Anderson (1988), Alonso-Zarazaga & Lyal (1999), Anderson (2002)
Literature Records: San Clemente (Fall, 1897: 239; Anderson, 1988: 649 [map])
Digitized Records: none
Range: Also recorded from mainland (Anderson, 1988).
Notes. This species was recorded by Fall (1897) as Cleonus basalis Fall, 1897, which is a junior homonym that was replaced by Csiki (1934) with Cleonus (Cleonidius) americanus Csiki, 1934. O’Brien & Wibmer (1982) corrected the name to Cleonis americanus (Csiki, 1934); later, Anderson (1988) moved the species to Cleonidius Casey, 1891. Cleonidius was more recently synonymized under the resurrected Scaphomorphus in Alonso-Zarazaga & Lyal (1999: 192). The species was considered endemic to San Clemente Island at the time of its original description (Fall, 1897).
Rhinocyllini
Notes. Two genera and three species of Rhinocyllini are known to occur in California (M. L. Gimmel, 2022, unpublished data).
Rhinocyllus Germar, 1817
Nomenclatural Authority: Anderson (2002)
Notes. This genus contains a single species in California (O’Brien & Wibmer, 1982).
Rhinocyllus conicus (Froelich, 1792)
Nomenclatural Authority: Anderson (2002)
Literature Records: Santa Cruz (Hawkes, Andres & Dunn, 1972: 14; Goeden & Ricker, 1974: 9; Goeden & Ricker, 1977: 289), Santa Rosa (Miller, 1996: 197)
Digitized Records: Santa Cruz (1 iNat)
Range: Also known from mainland (O’Brien & Wibmer, 1982; Anderson, 2002).
Notes. This species was introduced into North America for biological control of Carduus nutans L. (Asteraceae) (Goeden & Ricker, 1974, 1977; Anderson, 2002). Although Hawkes, Andres & Dunn (1972) and Goeden & Ricker (1974, 1977) reported that the 1971 Santa Cruz Island introductions were unsuccessful, the 2017 iNaturalist record above provides evidence that the species may be established there. Miller (1996) reported on an attempted 1979 introduction to Santa Rosa Island, the results of which are unknown.
Molytinae
Notes. Eight tribes, 13 genera, and 24 species of Molytinae have been recorded from California (M. L. Gimmel, 2022, unpublished data).
Conotrachelini
Notes. Two genera and three species of Conotrachelini have been recorded from California (O’Brien & Wibmer, 1982).
Micromastus LeConte, 1876
Nomenclatural Authority: Anderson (2002)
Notes. This genus contains a single species in California (O’Brien & Wibmer, 1982; Anderson, 2002).
Micromastus gracilis (Boheman, 1859)
Nomenclatural Authority: Anderson (2002)
Literature Records: none
Digitized Records: Santa Catalina (12 SBMNH)
Range: Also known from mainland (O’Brien & Wibmer, 1982).
Emphyastini
Notes. Two genera and two species of Emphyastini have been recorded from California (O’Brien, 1997).
Emphyastes Mannerheim, 1852
Nomenclatural Authority: Anderson (2002)
Notes. This genus contains a single species in California (O’Brien & Wibmer, 1982; Anderson, 2002). It was treated in the subfamily Cyclominae by Anderson (2002).
Emphyastes fucicola Mannerheim, 1852
Nomenclatural Authority: Anderson (2002)
Literature Records: none
Digitized Records: San Clemente (7 SBMNH), San Nicolas (14 LACM; 5 SBMNH), Santa Cruz (2 SBMNH), Santa Rosa (1 iNat)
Range: Also known from mainland (O’Brien & Wibmer, 1982; Anderson, 2002).
Notes. Adults and larvae are associated with buried, decaying seaweed on sandy beaches (Anderson, 2002).
Thalasselephas Egorov & Korotyaev, 1976
Nomenclatural Authority: Anderson (2002)
Notes. This genus contains a single species in California (Anderson, 2002). The genus was renamed from Phycocoetes LeConte, 1876 as Neophycocoetes O’Brien & Wibmer, 1982 in O’Brien & Wibmer (1982).
Thalasselephas testaceus (LeConte, 1876)
Nomenclatural Authority: Anderson (2002)
Literature Records: none
Digitized Records: San Clemente (4 SBMNH), San Nicolas (11 SBMNH), Santa Rosa (2 SBMNH)
Range: Also known from mainland (O’Brien & Wibmer, 1982; Anderson, 2002).
Notes. Adults of this species are found under seaweed on sandy beaches (Anderson, 2002).
Raymondionyminae
Notes. Three genera and five species of Raymondionyminae have been recorded from California (O’Brien & Wibmer, 1982).
Gilbertiola Osella, 1982
Nomenclatural Authority: Anderson (2002)
Notes. This genus contains two described species from California (Anderson, 2002).
Gilbertiola undetermined species
Literature Records: Santa Catalina (Caterino & Chandler, 2010: 191)
Digitized Records: Santa Catalina (1 SBMNH)
Notes. This species, represented by a single SBMNH specimen collected on Santa Catalina Island, was reported only as the genus Gilbertiola by Caterino & Chandler (2010).
Scolytinae
Notes. Two tribes, 49 genera, and 196 species of Scolytinae are known to occur in California (Atkinson, 2021; M. L. Gimmel, 2022, unpublished data). Bright & Stark (1973) reviewed the California fauna of Scolytinae, and Wood (1982) reviewed the entire North American fauna known at the time. Atkinson (2021) is an excellent online reference for staying up-to-date with North American scolytine nomenclature.
Hylesinini
Notes. Sixteen genera and 69 species of Hylesinini are known to occur in California (Atkinson, 2021; M. L. Gimmel, 2022, unpublished data).
Carphobius Blackman, 1943
Nomenclatural Authority: Wood (1982)
Notes. The genus Carphobius has not been previously reported from California (Atkinson, 2021), but is known from two species occurring from Arizona to Guatemala (Wood, 1982).
Carphobius undescribed species
Literature Records: none
Digitized Records: San Clemente (1 SBMNH)
Range: Endemic (A. Cognato & S. Smith, 2022, unpublished data)
Notes. The San Clemente Island specimen cited above belongs to a new species that is currently under description (A. Cognato & S. Smith, 2022, personal communication).
Carphoborus Eichhoff, 1864
Nomenclatural Authority: Wood (1982)
Notes. Nine species of this genus are reported from California (Atkinson, 2021).
Carphoborus declivis Wood, 1954
Nomenclatural Authority: Wood (1982)
Literature Records: none
Digitized Records: Santa Rosa (1 SBMNH)
Range: Also known from mainland (Wood, 1982).
Dendroctonus Erichson, 1836
Nomenclatural Authority: Wood (1982)
Notes. Six species of this genus are reported to occur in California (Atkinson, 2021).
Dendroctonus valens LeConte, 1859
Nomenclatural Authority: Atkinson (2021)
Literature Records: none
Digitized Records: Santa Cruz (5 SBMNH; 1 UCSB)
Range: Also known from mainland (Wood, 1982).
Scolytini
Notes. Thirty-three genera and 127 species of Scolytini are known to occur in California (Atkinson, 2021; M. L. Gimmel, 2022, unpublished data).
Coccotrypes Eichhoff, 1878
Nomenclatural Authority: Wood (1982)
Notes. Three adventive species of Coccotrypes have been recorded from California (Atkinson, 2021).
Coccotrypes dactyliperda (Fabricius, 1801)
Nomenclatural Authority: Atkinson (2021)
Literature Records: Santa Catalina (Holzman et al., 2009: 1078)
Digitized Records: none
Range: Also known from mainland (Wood, 1982).
Notes. This species was introduced to North America (Wood, 1982).
Dendrocranulus Schedl, 1937
Nomenclatural Authority: Wood (1982)
Notes. One species of Dendrocranulus has been recorded from California (Atkinson, 2021).
Dendrocranulus cucurbitae (LeConte, 1879)
Nomenclatural Authority: Atkinson (2021)
Literature Records: Santa Cruz (Naughton et al., 2014: 303)
Digitized Records: Anacapa (2 SBMNH), San Clemente (3 SBMNH), Santa Catalina (5 SBMNH), Santa Cruz (7 SBMNH), Santa Rosa (10 SBMNH)
Range: Also known from mainland (Wood, 1982).
Notes. This species was reported by Naughton et al. (2014) as Dendrocranulus californicus (Hopkins, 1915), now considered a junior synonym of D. cucurbitae (Wood, 1982; Atkinson, 2021).
Gnathotrichus Eichhoff, 1869
Nomenclatural Authority: Wood (1982)
Notes. This genus is represented in California by five species (Atkinson, 2021).
Gnathotrichus pilosus (LeConte, 1868)
Nomenclatural Authority: Atkinson (2021)
Literature Records: none
Digitized Records: Santa Cruz (4 SBMNH)
Range: Also known from mainland (Wood, 1982).
Hypothenemus Westwood, 1836
Nomenclatural Authority: Wood (1982)
Notes. Only three species of this large genus have been recorded from California (Atkinson, 2021).
Hypothenemus eruditus Westwood, 1836
Nomenclatural Authority: Atkinson (2021)
Literature Records: none
Digitized Records: San Clemente (1 SBMNH), Santa Catalina (1 SBMNH), Santa Rosa (11 SBMNH)
Range: Also known from mainland (Wood, 1982).
Ips DeGeer, 1775
Nomenclatural Authority: Wood (1982)
Notes. Lanier (1970: 1418) hypothesized that Ips plastographus maritimus Lanier, 1970 probably occurred on pines “on the islands off the coast of southern California and Mexico”, but this species has not yet been recorded from the Channel Islands.
Ips paraconfusus Lanier, 1970
Nomenclatural Authority: Atkinson (2021)
Literature Records: Santa Cruz (Bright & Stark, 1973: 92 [map])
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Bright & Stark, 1973; Wood, 1982).
Monarthrum Kirsch, 1866
Nomenclatural Authority: Wood (1982)
Notes. Three species of this genus are known to occur in California (Atkinson, 2021).
Monarthrum scutellare (LeConte, 1857)
Nomenclatural Authority: Atkinson (2021)
Literature Records: none
Digitized Records: Santa Cruz (6 SBMNH)
Range: Also known from mainland (Wood, 1982).
Pityophthorus Eichhoff, 1864
Nomenclatural Authority: Wood (1982)
Notes. Thirty-three species of Pityophthorus are known to occur in California (Atkinson, 2021).
Pityophthorus carmeli Swaine, 1918
Nomenclatural Authority: Atkinson (2021)
Literature Records: none
Digitized Records: Santa Cruz (10 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Wood, 1982).
Procryphalus Hopkins, 1915
Nomenclatural Authority: Wood (1982)
Notes. One species of this genus occurs in California (Atkinson, 2021).
Procryphalus utahensis Hopkins, 1915
Nomenclatural Authority: Atkinson (2021)
Literature Records: none
Digitized Records: San Miguel (1 SBMNH)
Range: Also known from mainland (Wood, 1982).
Pseudips Cognato, 2000
Nomenclatural Authority: Cognato (2000)
Notes. This genus contains two species occurring in California (Atkinson, 2021).
Pseudips mexicanus (Hopkins, 1905)
Nomenclatural Authority: Atkinson (2021)
Literature Records: Santa Cruz (Bright & Stark, 1973: 86)
Digitized Records: none
Range: Also known from mainland (Bright & Stark, 1973).
Notes. Bright & Stark (1973) and Wood (1982) reported this species as Ips mexicanus; however, Cognato (2000) placed this species in the new genus Pseudips.
Pseudopityophthorus Swaine, 1918
Nomenclatural Authority: Wood (1982)
Literature Records (genus-only): Santa Barbara (Miller & Miller, 1985: 132)
Notes. Three species of this genus are known to occur in California (Atkinson, 2021). Miller & Miller (1985) reported an unidentified species of this genus from Santa Barbara Island that was collected from Eriophyllum (Asteraceae).
Pseudopityophthorus agrifoliae Blackman, 1931
Nomenclatural Authority: Atkinson (2021)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH)
Range: Also known from mainland (Wood, 1982).
Pseudopityophthorus pubipennis (LeConte, 1857)
Nomenclatural Authority: Atkinson (2021)
Literature Records: none
Digitized Records: Santa Cruz (1 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Wood, 1982).
Stenoclyptus Blackman, 1943
Nomenclatural Authority: Wood (1982), Atkinson (2021)
Notes. One species of this genus is known from California (Atkinson, 2021). The name of this genus was misspelled “Stenocleptus” in Wood (1982).
Stenoclyptus sulcatus (Bruck, 1936)
Nomenclatural Authority: Atkinson (2021)
Literature Records: none
Digitized Records: Santa Rosa (11 SBMNH)
Range: Also known from mainland (Wood, 1982).
Xyleborinus Reitter, 1913
Nomenclatural Authority: Wood (1982)
Notes. One species of this genus is known from California (Atkinson, 2021).
Xyleborinus saxesenii (Ratzeburg, 1837)
Nomenclatural Authority: Atkinson (2021)
Literature Records: none
Digitized Records: Santa Cruz (9 SBMNH), Santa Rosa (1 SBMNH)
Range: Also known from mainland (Wood, 1982; Rabaglia, Dole & Cognato, 2006).
Notes. This species was probably introduced to North America from Asia (Rabaglia, Dole & Cognato, 2006).